743 lines
92 KiB
XML
743 lines
92 KiB
XML
<document id="994664B13CC20067A7B8C150CE247747" ID-DOI="10.37520/aemnp.2023.005" ID-ISSN="1804-6487" ID-Zenodo-Dep="8005982" ID-ZooBank="5774BE11-4CDE-4703-8CB6-16FC7FAFCEED" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685971728898" checkinUser="felipe" docAuthor="Sato, Yuya & Maruyama, Munetoshi" docDate="2023" docId="6F3B0917FFF1FFFECA9384F1FE59F780" docLanguage="en" docName="ActaEntMusNatPra.63.1.111-124.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 63 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2023.005" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Lathrobium nomurai Nakane 1955" docType="treatment" docVersion="2" lastPageNumber="117" masterDocId="9302716FFFF0FFF8C934850CFFDAFF95" masterDocTitle="Taxonomy of the Lathrobium nomurai group (Coleoptera: Staphylinidae) from Northern Kyushu, Japan, with descriptions of two new species" masterLastPageNumber="124" masterPageNumber="111" pageNumber="112" updateTime="1685972318776" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="270F9C3EB8A76A1EC3E86BFE99F53D40">Taxonomy of the Lathrobium nomurai group (Coleoptera: Staphylinidae) from Northern Kyushu, Japan, with descriptions of two new species</mods:title>
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<mods:namePart id="83784B04F6FCE11FDD9518291C46FA56">Sato, Yuya</mods:namePart>
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<mods:affiliation id="0844CA3FC5018583B6AD90E1E8A6DFE4">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, JP- 819 - 0395, Japan; e-mail: inperiadm @ gmail. com & The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:namePart id="E25916E2997FFBA4B09E85BFC4D5ACDD">Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation id="0C021B638846103BB3FA20A65EA51386">The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:title id="8A66950A7B2C7E16094B7AE05B44B96C">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="FB4EF1D44E8A86D57A5359CF78506D29">2023</mods:date>
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<mods:number id="34CA277BEE2F17581E1103AB211A1835">2023-05-06</mods:number>
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<subSubSection id="AF88EB8AFFF1FFF9CA9384F1FAE9FDBA" pageId="1" pageNumber="112" type="nomenclature">
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<paragraph id="E72DB801FFF1FFF9CA9384F1FAEBFD80" blockId="1.[935,1329,509,533]" box="[935,1329,509,533]" pageId="1" pageNumber="112">
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<heading id="BC650F6DFFF1FFF9CA9384F1FAEBFD80" box="[935,1329,509,533]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="112" reason="3">
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<taxonomicName id="2092C382FFF1FFF9CA9384F1FAEBFD80" ID-CoL="3SJHT" authority="Nakane, 1955" authorityName="Nakane" authorityYear="1955" box="[935,1329,509,533]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="112" phylum="Arthropoda" rank="species" species="nomurai">
|
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<emphasis id="D5E66413FFF1FFF9CA9384F1FAEBFD80" bold="true" box="[935,1329,509,533]" pageId="1" pageNumber="112">
|
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<emphasis id="D5E66413FFF1FFF9CA9384F1FB4AFD80" bold="true" box="[935,1168,509,533]" italics="true" pageId="1" pageNumber="112">Lathrobium nomurai</emphasis>
|
||
<bibRefCitation id="8303C5F0FFF1FFF9CDA384F2FAEBFD80" author="NAKANE T." box="[1175,1329,510,533]" pageId="1" pageNumber="112" pagination="24 - 42" refId="ref8544" refString="NAKANE T. 1955: New or little-known Coleoptera from Japan and its adjacent region, XII. Scientific Reports of the Saikyo University, Natural Science and Living Science 2 (1): 24 - 42." type="journal article" year="1955">Nakane, 1955</bibRefCitation>
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</emphasis>
|
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</taxonomicName>
|
||
</heading>
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</paragraph>
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||
<paragraph id="E72DB801FFF1FFF9CA918710FAE9FDBA" blockId="1.[933,1331,540,586]" box="[933,1331,540,559]" pageId="1" pageNumber="112">[Japanese name: Ô-kobane-naga-hanekakushi]</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF1FFF9CD01873BFB79FDDF" box="[1077,1187,567,586]" pageId="1" pageNumber="112" type="description">
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<paragraph id="E72DB801FFF1FFF9CD01873BFB79FDDF" blockId="1.[933,1331,540,586]" box="[1077,1187,567,586]" pageId="1" pageNumber="112">
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(
|
||
<figureCitation id="7FA9A484FFF1FFF9CD0F873BFBABFDDF" box="[1083,1137,567,586]" captionStart="Fig" captionStartId="2.[151,182,1520,1539]" captionTargetBox="[336,1212,198,1467]" captionTargetId="figure-13@2.[299,1288,186,1477]" captionTargetPageId="2" captionText="Fig. 1. Lathrobium nomurai Nakane, 1955. A – habitus in dorsal view; B – labels of holotype; C – male sternite VII; D – male sternite VIII; E – female sternite VIII. Scale bars = 2.0 mm for A, 0.5 mm for C–E." figureDoi="http://doi.org/10.5281/zenodo.8005984" httpUri="https://zenodo.org/record/8005984/files/figure.png" pageId="1" pageNumber="112">Figs 1</figureCitation>
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,
|
||
<figureCitation id="7FA9A484FFF1FFF9CD48873BFB52FDDF" box="[1148,1160,567,586]" captionStart="Fig" captionStartId="3.[150,181,1021,1040]" captionTargetBox="[319,1306,190,973]" captionTargetId="figure-478@3.[214,1371,175,986]" captionTargetPageId="3" captionText="Fig. 2. Male aedeagus of Lathrobium nomurai Nakane, 1955: A – ventral view; B – lateral view; C – dorsal view. Scale bar = 0.5 mm. Abbreviations: AP – apical part of ventral process; BF – basal foramen; DP – dorsal plate; VP – ventral process. Black arrow indicates the V-shaped impression of basal margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.8005986" httpUri="https://zenodo.org/record/8005986/files/figure.png" pageId="1" pageNumber="112">2</figureCitation>
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,
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||
<figureCitation id="7FA9A484FFF1FFF9CDA5873BFB46FDDF" box="[1169,1180,567,586]" captionStart="Fig" captionStartId="11.[151,182,1153,1172]" captionTargetBox="[254,1353,201,1126]" captionTargetId="figure-402@11.[207,1379,186,1137]" captionTargetPageId="11" captionText="Fig. 9. Map showing the distribution of the Lathrobium nomurai group in Kyushu, Japan. Black rhombus – L. nomurai Nakane, 1955; black triangle – L. fujimotoi Y. Watanabe, 2001; black circle – L. hikosanense Y. Watanabe, 1998; black square – L. yufuense sp. nov.; black pentagon – L. denchu sp. nov.; A–G – localities of L. hikosanense: A – Mt. Daisen; B – Mt. Hiko-san (type locality for L. hikosanense); C – Hiraodai; D – Mt. Buccho-zan; E – Mt. Tachibana-yama; F – Mt. Iwara-yama; G – Kinsenji Temple." figureDoi="http://doi.org/10.5281/zenodo.8006000" httpUri="https://zenodo.org/record/8006000/files/figure.png" pageId="1" pageNumber="112">9</figureCitation>
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)
|
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</paragraph>
|
||
</subSubSection>
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<subSubSection id="AF88EB8AFFF1FFF9CA08876DFBB7FC7C" pageId="1" pageNumber="112" type="materials_examined">
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<paragraph id="E72DB801FFF1FFF9CA08876DFB22FD6C" blockId="1.[828,1437,609,1001]" pageId="1" pageNumber="112">
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<emphasis id="D5E66413FFF1FFF9CA08876DFBC6FDE1" bold="true" box="[828,1052,609,628]" pageId="1" pageNumber="112">Type material examined.</emphasis>
|
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<materialsCitation id="57FAB25CFFF1FFF9CD15876DFB22FD6C" collectingDate="1954-10-03" collectionCode="SEHU, OHARA, INARI, KANBE, T" collectorName="S. Nomura" country="Japan" location="Japan" municipality="Kyushu" pageId="1" pageNumber="112" specimenCount="3" stateProvince="Hokkaido" typeStatus="holotype">
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<typeStatus id="382906A3FFF1FFF9CD15876DFBA1FDE1" box="[1057,1147,609,628]" pageId="1" pageNumber="112">HOLOTYPE</typeStatus>
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: 3,
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:
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<collectingMunicipality id="0749227BFFF1FFF9CDEF876DFAFAFDE1" box="[1243,1312,609,628]" pageId="1" pageNumber="112">Kyushu</collectingMunicipality>
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, “0000000424 / Sys. Ent /
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<collectingRegion id="255676E3FFF1FFF9CAA98777FC2BFD1B" box="[925,1009,635,654]" country="Japan" name="Hokkaido" pageId="1" pageNumber="112">Hokkaido</collectingRegion>
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Univ. / Japan [
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<collectionCode id="818320C4FFF1FFF9CD5B8777FB71FD1B" box="[1135,1195,635,654]" pageId="1" pageNumber="112">SEHU</collectionCode>
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] //
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<typeStatus id="382906A3FFF1FFF9CDF18777FAECFD1B" box="[1221,1334,635,654]" pageId="1" pageNumber="112">HOLOTYPE</typeStatus>
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/ Appended labeled by /
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<collectionCode id="818320C4FFF1FFF9CA94879AFC31FD3C" box="[928,1003,662,681]" pageId="1" pageNumber="112">ÔHARA</collectionCode>
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/
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<collectionCode id="818320C4FFF1FFF9CACD879AFBEEFD3C" box="[1017,1076,662,681]" pageId="1" pageNumber="112">INARI</collectionCode>
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/
|
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<collectionCode id="818320C4FFF1FFF9CD76879AFB51FD3C" box="[1090,1163,662,681]" pageId="1" pageNumber="112">KANBE</collectionCode>
|
||
/ SUZUKI / and / HIRONAGA / 2006 // NAKANE Coll. /
|
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<collectionCode id="818320C4FFF1FFF9CD2487BDFB92FD51" box="[1040,1096,689,708]" pageId="1" pageNumber="112">SEHU</collectionCode>
|
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/
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||
<collectingCountry id="9F85F891FFF1FFF9CD6187BDFB4FFD51" box="[1109,1173,689,708]" name="Japan" pageId="1" pageNumber="112">JAPAN</collectingCountry>
|
||
/ 1999 //
|
||
<taxonomicName id="2092C382FFF1FFF9CDD687BDFA9CFD51" authorityName="Gravenhorst" authorityYear="1802" box="[1250,1350,689,708]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="112" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D5E66413FFF1FFF9CDD687BDFA9CFD51" box="[1250,1350,689,708]" italics="true" pageId="1" pageNumber="112">Lathrobium</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="2092C382FFF1FFF9CC6087BDFA46FD51" authorityName="Nakane" authorityYear="1955" box="[1364,1436,689,708]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="112" phylum="Arthropoda" rank="species" species="nomurai">
|
||
<emphasis id="D5E66413FFF1FFF9CC6087BDFA46FD51" box="[1364,1436,689,708]" italics="true" pageId="1" pageNumber="112">nomurai</emphasis>
|
||
</taxonomicName>
|
||
/ Det.
|
||
<collectionCode id="818320C4FFF1FFF9CA5B87C7FCA7FD4B" box="[879,893,715,734]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="112">T</collectionCode>
|
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.
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||
<bibRefCitation id="8303C5F0FFF1FFF9CAB187C7FC2CFD4B" author="NAKANE T." box="[901,1014,715,734]" pageId="1" pageNumber="112" pagination="24 - 42" refId="ref8544" refString="NAKANE T. 1955: New or little-known Coleoptera from Japan and its adjacent region, XII. Scientific Reports of the Saikyo University, Natural Science and Living Science 2 (1): 24 - 42." type="journal article" year="1955">Nakane 1955</bibRefCitation>
|
||
// Tsuruoka-Dô / Saeki /
|
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<collectingRegion id="255676E3FFF1FFF9CDFE87C7FB35FD4B" box="[1226,1263,715,734]" country="Japan" name="Oita" pageId="1" pageNumber="112">Oita</collectingRegion>
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||
Prf. /
|
||
<date id="932C9EC1FFF1FFF9CC1687C7FAA6FD4B" box="[1314,1404,715,734]" pageId="1" pageNumber="112" value="1954-10-03">
|
||
<collectingDate id="83686729FFF1FFF9CC1687C7FAA6FD4B" box="[1314,1404,715,734]" pageId="1" pageNumber="112" value="1954-10-03">3. X. 1954</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="4A67DDD7FFF1FFF9CCBF87C7FC58FD6C" pageId="1" pageNumber="112">S. Nomura</collectorName>
|
||
// 45–9 //
|
||
<typeStatus id="382906A3FFF1FFF9CAED87EAFB96FD6C" box="[985,1100,742,761]" pageId="1" pageNumber="112">HOLOTYPE</typeStatus>
|
||
” (
|
||
<collectionCode id="818320C4FFF1FFF9CD6B87EAFB43FD6C" box="[1119,1177,742,761]" pageId="1" pageNumber="112">SEHU</collectionCode>
|
||
) (
|
||
<figureCitation id="7FA9A484FFF1FFF9CD9E87EAFB37FD6C" box="[1194,1261,742,761]" captionStart="Fig" captionStartId="2.[151,182,1520,1539]" captionTargetBox="[336,1212,198,1467]" captionTargetId="figure-13@2.[299,1288,186,1477]" captionTargetPageId="2" captionText="Fig. 1. Lathrobium nomurai Nakane, 1955. A – habitus in dorsal view; B – labels of holotype; C – male sternite VII; D – male sternite VIII; E – female sternite VIII. Scale bars = 2.0 mm for A, 0.5 mm for C–E." figureDoi="http://doi.org/10.5281/zenodo.8005984" httpUri="https://zenodo.org/record/8005984/files/figure.png" pageId="1" pageNumber="112">Fig. 1B</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF1FFF9CA08860DFBB7FC7C" blockId="1.[828,1437,609,1001]" pageId="1" pageNumber="112">
|
||
<emphasis id="D5E66413FFF1FFF9CA08860DFB88FC81" bold="true" box="[828,1106,769,788]" pageId="1" pageNumber="112">Additional material examined.</emphasis>
|
||
1 3, “[
|
||
<collectingCountry id="9F85F891FFF1FFF9CDBB860DFB03FC81" box="[1167,1241,769,788]" name="Japan" pageId="1" pageNumber="112">JAPAN</collectingCountry>
|
||
]: Oita-ken / Saeki-shi / Kariu /
|
||
<quantity id="206A15E4FFF1FFF9CA4F8617FC7FFCBB" box="[891,933,795,814]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="112" unit="m" value="90.0">90 m</quantity>
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in alt. / [
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<geoCoordinate id="82A6DEC6FFF1FFF9CAD88617FBF1FCBB" box="[1004,1067,795,814]" degrees="33.031" direction="north" orientation="latitude" pageId="1" pageNumber="112" precision="55" value="33.031">33.031</geoCoordinate>
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,
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] /
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<date id="932C9EC1FFF1FFF9CDB88617FB35FCBB" box="[1164,1263,795,814]" pageId="1" pageNumber="112" value="2019-10-27">27. X. 2019</date>
|
||
/ Tsubasa NOZAKI, leg.” (
|
||
<collectionCode id="818320C4FFF1FFF9CA44863AFC7CFCDC" box="[880,934,822,841]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="1" pageNumber="112" type="Herbarium">KUM</collectionCode>
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); 1 3, “[
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||
<collectingCountry id="9F85F891FFF1FFF9CADF863AFBEFFCDC" box="[1003,1077,822,841]" name="Japan" pageId="1" pageNumber="112">JAPAN</collectingCountry>
|
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]: Oita-ken / Saeki-shi / Kariu /
|
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<quantity id="206A15E4FFF1FFF9CC08863AFABDFCDC" box="[1340,1383,822,841]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="112" unit="m" value="90.0">90 m</quantity>
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in alt. / [
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<geoCoordinate id="82A6DEC6FFF1FFF9CA7F865DFC50FCF1" box="[843,906,849,868]" degrees="33.031" direction="north" orientation="latitude" pageId="1" pageNumber="112" precision="55" value="33.031">33.031</geoCoordinate>
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,
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<geoCoordinate id="82A6DEC6FFF1FFF9CAA6865DFC00FCF1" box="[914,986,849,868]" degrees="131.895" direction="east" orientation="longitude" pageId="1" pageNumber="112" precision="55" value="131.895">131.895</geoCoordinate>
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] /
|
||
<date id="932C9EC1FFF1FFF9CAD9865DFB97FCF1" box="[1005,1101,849,868]" pageId="1" pageNumber="112" value="2019-11-09">9. XI. 2019</date>
|
||
/ Tsubasa NOZAKI, leg.” (
|
||
<collectionCode id="818320C4FFF1FFF9CC07865DFAB3FCF1" box="[1331,1385,849,868]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="1" pageNumber="112" type="Herbarium">KUM</collectionCode>
|
||
); 1 3,
|
||
<specimenCount id="F1947388FFF1FFF9CA088667FC86FCEB" box="[828,860,875,894]" pageId="1" pageNumber="112" type="female">1 ♀</specimenCount>
|
||
, “
|
||
<collectingRegion id="255676E3FFF1FFF9CA5B8667FC4EFCEB" box="[879,916,875,894]" country="Japan" name="Oita" pageId="1" pageNumber="112">Oita</collectingRegion>
|
||
–ken / Tsukumi-shi / Kamiaoe /
|
||
<date id="932C9EC1FFF1FFF9CD9E8667FADFFCEB" box="[1194,1285,875,894]" pageId="1" pageNumber="112" value="2007-10-09">9. X. 2007</date>
|
||
/
|
||
<collectionCode id="818320C4FFF1FFF9CC228667FAFEFCEB" box="[1302,1316,875,894]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="112">T</collectionCode>
|
||
. Miyake leg.” (
|
||
<collectionCode id="818320C4FFF1FFF9CA76868AFCA8FC0C" box="[834,882,902,921]" pageId="1" pageNumber="112">TUA</collectionCode>
|
||
);
|
||
<specimenCount id="F1947388FFF1FFF9CAB4868AFC41FC0C" box="[896,923,902,921]" pageId="1" pageNumber="112" type="female">1♀</specimenCount>
|
||
, “(Tsuruoka Mine) / Saeki /
|
||
<collectingRegion id="255676E3FFF1FFF9CDB8868AFB3AFC0C" box="[1164,1248,902,921]" country="Japan" name="Oita" pageId="1" pageNumber="112">Oita Pref.</collectingRegion>
|
||
/
|
||
<collectingCountry id="9F85F891FFF1FFF9CDDA868AFAC5FC0C" box="[1262,1311,902,921]" name="Japan" pageId="1" pageNumber="112">Japan</collectingCountry>
|
||
/
|
||
<date id="932C9EC1FFF1FFF9CC1A868AFA48FC0C" box="[1326,1426,902,921]" pageId="1" pageNumber="112" value="1955-03-23">23-III-1955</date>
|
||
/ Coll. S-I. Uéno” (
|
||
<collectionCode id="818320C4FFF1FFF9CAE086ADFBDEFC21" box="[980,1028,929,948]" pageId="1" pageNumber="112">TUA</collectionCode>
|
||
);
|
||
<specimenCount id="F1947388FFF1FFF9CD2786ADFBF4FC21" box="[1043,1070,929,948]" pageId="1" pageNumber="112" type="female">1♀</specimenCount>
|
||
, “(
|
||
<collectionCode id="818320C4FFF1FFF9CD7386ADFB50FC21" box="[1095,1162,929,948]" pageId="1" pageNumber="112">SAEKI</collectionCode>
|
||
) /
|
||
<collectingRegion id="255676E3FFF1FFF9CD9586ADFB1CFC21" box="[1185,1222,929,948]" country="Japan" name="Oita" pageId="1" pageNumber="112">Oita</collectingRegion>
|
||
/ Kyushu / XI–20.1946 / Coll. K. Kutosa” (
|
||
<collectionCode id="818320C4FFF1FFF9CAE086B7FBDEFC5B" box="[980,1028,955,974]" pageId="1" pageNumber="112">TUA</collectionCode>
|
||
);
|
||
<specimenCount id="F1947388FFF1FFF9CD2686B7FBF7FC5B" box="[1042,1069,955,974]" pageId="1" pageNumber="112" type="female">1♀</specimenCount>
|
||
, “
|
||
<collectingCountry id="9F85F891FFF1FFF9CD0A86B7FB5EFC5B" box="[1086,1156,955,974]" name="Japan" pageId="1" pageNumber="112">JAPAN</collectingCountry>
|
||
:
|
||
<collectingRegion id="255676E3FFF1FFF9CDBF86B7FB05FC5B" box="[1163,1247,955,974]" country="Japan" name="Oita" pageId="1" pageNumber="112">Ôita Pref.</collectingRegion>
|
||
/ Saeki-shi / Kadomae /
|
||
<date id="932C9EC1FFF1FFF9CA7386DAFC1BFC7C" box="[839,961,982,1001]" pageId="1" pageNumber="112" value="2020-03-12">12. Mar. 2020</date>
|
||
. / Tainaka” (
|
||
<collectionCode id="818320C4FFF1FFF9CD1886DAFBB8FC7C" box="[1068,1122,982,1001]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="1" pageNumber="112" type="Herbarium">KUM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF1FFFBCA08810DFE47F8E8" lastPageId="3" lastPageNumber="114" pageId="1" pageNumber="112" type="description">
|
||
<paragraph id="E72DB801FFF1FFF9CA08810DFB8BFB6D" blockId="1.[828,1437,1025,2072]" pageId="1" pageNumber="112">
|
||
<emphasis id="D5E66413FFF1FFF9CA08810DFC05FB8D" bold="true" box="[828,991,1025,1048]" pageId="1" pageNumber="112">Redescription.</emphasis>
|
||
BL = 11.0–
|
||
<quantity id="206A15E4FFF1FFF9CD63810DFB6FFB8D" box="[1111,1205,1025,1048]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="1" pageNumber="112" unit="mm" value="15.4">15.4 mm</quantity>
|
||
, FBL =
|
||
<quantity id="206A15E4FFF1FFF9CC31810DFA5BFB8D" box="[1285,1409,1025,1048]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="5.8" metricValueMin="5.5" pageId="1" pageNumber="112" unit="mm" value="5.65" valueMax="5.8" valueMin="5.5">5.5–5.8 mm</quantity>
|
||
(n = 8). Coloration: Body (
|
||
<figureCitation id="7FA9A484FFF1FFF9CD08812DFB54FBAD" box="[1084,1166,1057,1080]" captionStart="Fig" captionStartId="2.[151,182,1520,1539]" captionTargetBox="[336,1212,198,1467]" captionTargetId="figure-13@2.[299,1288,186,1477]" captionTargetPageId="2" captionText="Fig. 1. Lathrobium nomurai Nakane, 1955. A – habitus in dorsal view; B – labels of holotype; C – male sternite VII; D – male sternite VIII; E – female sternite VIII. Scale bars = 2.0 mm for A, 0.5 mm for C–E." figureDoi="http://doi.org/10.5281/zenodo.8005984" httpUri="https://zenodo.org/record/8005984/files/figure.png" pageId="1" pageNumber="112">Fig. 1A</figureCitation>
|
||
) reddish brown, moderately shining; maxillary palpi light red; legs reddish brown but basal part of tibia black or darken; mandible, scutellum, bases of antennomeres I–X black or darken; elytra reddish brown but elytral suture black; abdomen reddish brown, posterior margin of tergite VII and VIII light brownish red; tergite IX and X light red.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF1FFF9CA6E800DFBDCF8AD" blockId="1.[828,1437,1025,2072]" pageId="1" pageNumber="112">
|
||
Head suborbicular, HL =
|
||
<quantity id="206A15E4FFF1FFF9CD59800DFAD1FA8D" box="[1133,1291,1281,1304]" metricMagnitude="-3" metricUnit="m" metricValue="1.725" metricValueMax="1.8" metricValueMin="1.65" pageId="1" pageNumber="112" unit="mm" value="1.725" valueMax="1.8" valueMin="1.65">1.65–1.80 mm</quantity>
|
||
, HW =
|
||
<quantity id="206A15E4FFF1FFF9CC54800DFC47FAAD" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.95" metricValueMin="1.65" pageId="1" pageNumber="112" unit="mm" value="1.7999999999999998" valueMax="1.95" valueMin="1.65">1.65– 1.95 mm</quantity>
|
||
, HW/HL = 1.03, widest at the basal 1/5 of head; frontal area flattened; surface coarsely punctate; punctures sparser on frons and vertex, becoming denser toward posterior and lateral margins, with dark brown setae. Eyes very small, slightly protruding; longitudinal diameter of eye about 1/5 as long as temple in dorsal view. Antennae elongate, AL=
|
||
<quantity id="206A15E4FFF1FFF9CAE380EDFBA8FA6D" box="[983,1138,1505,1528]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.75" metricValueMin="3.85" pageId="1" pageNumber="112" unit="mm" value="4.3" valueMax="4.75" valueMin="3.85">3.85–4.75 mm</quantity>
|
||
, AL/HL = 2.56, with whitish yellow setae; antennomere I robust, widest at apical 1/3; antennomere II slightly elongated, shorter than III; antennomeres III–X slightly narrow at base, widening towards apex, successively shorter and narrower toward apical antennomeres but IX and X almost same size; antennomere XI fusiform; relative length of antennomeres as follows: 1.83: 1.08: 1.23: 1.21: 1.20: 1.13: 1.13: 1.10: 1.07: 1.00: 1.20; relative maximum width of antennomeres as follows: 1.74:1.57: 1.43: 1.43: 1.31: 1.31: 1.31: 1.29: 1.00: 1.00: 1.03.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF1FFFACA6E824DFDFEF90C" blockId="1.[828,1437,1025,2072]" lastBlockId="2.[151,760,1634,2073]" lastPageId="2" lastPageNumber="113" pageId="1" pageNumber="112">
|
||
Pronotum oblong-oval, with anterior margin wider than posterior one, PL = 2.00–
|
||
<quantity id="206A15E4FFF1FFF9CD7F826DFB76F8ED" box="[1099,1196,1889,1912]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="112" unit="mm" value="2.2">2.20 mm</quantity>
|
||
, PW =
|
||
<quantity id="206A15E4FFF1FFF9CDCF826DFA42F8ED" box="[1275,1432,1889,1912]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="1.74" metricValueMin="1.51" pageId="1" pageNumber="112" unit="mm" value="1.625" valueMax="1.74" valueMin="1.51">1.51–1.74 mm</quantity>
|
||
, PL/PW = 1.29, PL/HL = 1.24, PW/HW = 0.92, widest at apical 1/3; lateral sides of pronotum gently dilated to widest point, thence gradually convergent toward posterior margin; all corners obtusely rounded; disc sparsely covered with coarse and weak punctations except medial longitudinal line; punctures slightly sparser and less than on head, with brownish yellow setae.
|
||
</paragraph>
|
||
<caption id="B3EDE889FFF2FFFAC9A380FCFD5DF98B" ID-DOI="http://doi.org/10.5281/zenodo.8005984" ID-Zenodo-Dep="8005984" httpUri="https://zenodo.org/record/8005984/files/figure.png" pageId="2" pageNumber="113" startId="2.[151,182,1520,1539]" targetBox="[336,1212,198,1467]" targetPageId="2" targetType="figure">
|
||
<paragraph id="E72DB801FFF2FFFAC9A380FCFD5DF98B" blockId="2.[151,1436,1520,1566]" pageId="2" pageNumber="113">
|
||
Fig. 1.
|
||
<taxonomicName id="2092C382FFF2FFFAC9E780FCFDD9F996" authority="Nakane, 1955" authorityName="Nakane" authorityYear="1955" box="[211,515,1520,1539]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="nomurai">
|
||
<emphasis id="D5E66413FFF2FFFAC9E780FCFE5FF996" box="[211,389,1520,1539]" italics="true" pageId="2" pageNumber="113">Lathrobium nomurai</emphasis>
|
||
<bibRefCitation id="8303C5F0FFF2FFFAC8BE80FCFDD9F996" author="NAKANE T." box="[394,515,1520,1539]" pageId="2" pageNumber="113" pagination="24 - 42" refId="ref8544" refString="NAKANE T. 1955: New or little-known Coleoptera from Japan and its adjacent region, XII. Scientific Reports of the Saikyo University, Natural Science and Living Science 2 (1): 24 - 42." type="journal article" year="1955">Nakane, 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
. A – habitus in dorsal view; B – labels of holotype; C – male sternite VII; D – male sternite VIII; E – female sternite VIII. Scale bars = 2.0 mm for A, 0.5 mm for C–E.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E72DB801FFF2FFFAC98183AEFEB7F82C" blockId="2.[151,760,1634,2073]" pageId="2" pageNumber="113">
|
||
Scutellum with sparse punctures. Elytra subtrapezoidal, widened posteriorly, EL =
|
||
<quantity id="206A15E4FFF2FFFAC89E83CEFD9EF94C" box="[426,580,1730,1753]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" metricValueMax="1.3" metricValueMin="1.08" pageId="2" pageNumber="113" unit="mm" value="1.19" valueMax="1.3" valueMin="1.08">1.08–1.30 mm</quantity>
|
||
, EW =
|
||
<quantity id="206A15E4FFF2FFFACBBA83CEFF19F96C" metricMagnitude="-3" metricUnit="m" metricValue="1.805" metricValueMax="1.91" metricValueMin="1.7" pageId="2" pageNumber="113" unit="mm" value="1.805" valueMax="1.91" valueMin="1.7">1.70–1.91 mm</quantity>
|
||
, EW/EL = 1.54, EL/PL = 0.56, EW/PW = 1.11; lateral margins almost straight; posterior corners rounded; disc covered with sparse and coarse punctations; punctations denser than on pronotum, with brownish yellow setae. Hind wing completely reduced. Legs with shortened tibiae and tarsi which are covered with golden setae; tarsomeres I–IV of protarsus dilated.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF2FFFAC98182CEFA47F90C" blockId="2.[151,760,1634,2073]" lastBlockId="2.[828,1437,1634,2073]" pageId="2" pageNumber="113">Abdomen elongate, gradually widened from segment III–V, thence contracted to the posterior end; each tergite covered with micropunctations and brownish yellow setae, distinctly long setae on each side near anterior margin; tergite X as long as posterolateral processes of tergite IX.</paragraph>
|
||
<paragraph id="E72DB801FFF2FFFACA6E83ADFB7AF8AC" blockId="2.[828,1437,1634,2073]" pageId="2" pageNumber="113">
|
||
<emphasis id="D5E66413FFF2FFFACA6E83ADFC43F92C" bold="true" box="[858,921,1697,1721]" italics="true" pageId="2" pageNumber="113">Male.</emphasis>
|
||
Sternite VII (
|
||
<figureCitation id="7FA9A484FFF2FFFACD0783AEFB5CF92C" box="[1075,1158,1698,1721]" captionStart="Fig" captionStartId="2.[151,182,1520,1539]" captionTargetBox="[336,1212,198,1467]" captionTargetId="figure-13@2.[299,1288,186,1477]" captionTargetPageId="2" captionText="Fig. 1. Lathrobium nomurai Nakane, 1955. A – habitus in dorsal view; B – labels of holotype; C – male sternite VII; D – male sternite VIII; E – female sternite VIII. Scale bars = 2.0 mm for A, 0.5 mm for C–E." figureDoi="http://doi.org/10.5281/zenodo.8005984" httpUri="https://zenodo.org/record/8005984/files/figure.png" pageId="2" pageNumber="113">Fig. 1C</figureCitation>
|
||
) transverse, without posteromedian impression, posterior margin almost straight; sternite VIII (
|
||
<figureCitation id="7FA9A484FFF2FFFACAE183EEFBF1F96C" box="[981,1067,1762,1785]" captionStart="Fig" captionStartId="2.[151,182,1520,1539]" captionTargetBox="[336,1212,198,1467]" captionTargetId="figure-13@2.[299,1288,186,1477]" captionTargetPageId="2" captionText="Fig. 1. Lathrobium nomurai Nakane, 1955. A – habitus in dorsal view; B – labels of holotype; C – male sternite VII; D – male sternite VIII; E – female sternite VIII. Scale bars = 2.0 mm for A, 0.5 mm for C–E." figureDoi="http://doi.org/10.5281/zenodo.8005984" httpUri="https://zenodo.org/record/8005984/files/figure.png" pageId="2" pageNumber="113">Fig. 1D</figureCitation>
|
||
) with posteromedian impression, with sparse setae; posterior margin of sternite VIII slightly asymmetric; sternite IX elongate.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF2FFFBCA6E824EFE77FA48" blockId="2.[828,1437,1634,2073]" lastBlockId="3.[151,761,1190,2077]" lastPageId="3" lastPageNumber="114" pageId="2" pageNumber="113">
|
||
Aedeagus (
|
||
<figureCitation id="7FA9A484FFF2FFFACAE2824EFB8BF8CC" box="[982,1105,1858,1881]" captionStart="Fig" captionStartId="3.[150,181,1021,1040]" captionTargetBox="[319,1306,190,973]" captionTargetId="figure-478@3.[214,1371,175,986]" captionTargetPageId="3" captionText="Fig. 2. Male aedeagus of Lathrobium nomurai Nakane, 1955: A – ventral view; B – lateral view; C – dorsal view. Scale bar = 0.5 mm. Abbreviations: AP – apical part of ventral process; BF – basal foramen; DP – dorsal plate; VP – ventral process. Black arrow indicates the V-shaped impression of basal margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.8005986" httpUri="https://zenodo.org/record/8005986/files/figure.png" pageId="2" pageNumber="113">Figs 2A–C</figureCitation>
|
||
)
|
||
<quantity id="206A15E4FFF2FFFACD55824EFB1EF8CC" box="[1121,1220,1858,1881]" metricMagnitude="-3" metricUnit="m" metricValue="2.63" pageId="2" pageNumber="113" unit="mm" value="2.63">2.63 mm</quantity>
|
||
long, elongate; basal part of aedeagus with V-shaped impression; ventral process well sclerotized, slightly asymmetrical in ventral view, gradually narrowed toward apical 1/5, strongly narrowed toward apex, curved ventrally from 1/2 to apex; apical part of ventral process slightly curved toward left side in ventral view; apex of ventral process with small projection; basal and middle parts of ventral process slightly thicker than apex in lateral view; part above basal foramen transverse, with two projections at lateral side; dorsal plate sclerotized, widest at apical 1/4, gradually widened to the widest point at apical 1/4, thus contracted toward apex, with rounded apex; two tubes attached with apex of ventral process in between ventral process and dorsal plate; internal sac with sclerotized structures; ringed part connected with large flattened sac by sclerotized long-narrowed part.
|
||
</paragraph>
|
||
<caption id="B3EDE889FFF3FFFBC9A286F1FE9EFBD3" ID-DOI="http://doi.org/10.5281/zenodo.8005986" ID-Zenodo-Dep="8005986" httpUri="https://zenodo.org/record/8005986/files/figure.png" pageId="3" pageNumber="114" startId="3.[150,181,1021,1040]" targetBox="[319,1306,190,973]" targetPageId="3" targetType="figure">
|
||
<paragraph id="E72DB801FFF3FFFBC9A286F1FE9EFBD3" blockId="3.[150,1437,1021,1094]" pageId="3" pageNumber="114">
|
||
Fig. 2. Male aedeagus of
|
||
<taxonomicName id="2092C382FFF3FFFBC84586F1FD79FB85" authority="Nakane, 1955" authorityName="Nakane" authorityYear="1955" box="[369,675,1021,1040]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="nomurai">
|
||
<emphasis id="D5E66413FFF3FFFBC84586F1FDF8FB85" box="[369,546,1021,1040]" italics="true" pageId="3" pageNumber="114">Lathrobium nomurai</emphasis>
|
||
<bibRefCitation id="8303C5F0FFF3FFFBCB1D86F1FD79FB85" author="NAKANE T." box="[553,675,1021,1040]" pageId="3" pageNumber="114" pagination="24 - 42" refId="ref8544" refString="NAKANE T. 1955: New or little-known Coleoptera from Japan and its adjacent region, XII. Scientific Reports of the Saikyo University, Natural Science and Living Science 2 (1): 24 - 42." type="journal article" year="1955">Nakane, 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
: A – ventral view; B – lateral view; C – dorsal view. Scale bar = 0.5 mm. Abbreviations: AP – apical part of ventral process; BF – basal foramen; DP – dorsal plate; VP – ventral process. Black arrow indicates the V-shaped impression of basal margin of aedeagus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E72DB801FFF3FFFBC98180E9FE47F8E8" blockId="3.[151,761,1190,2077]" pageId="3" pageNumber="114">
|
||
<emphasis id="D5E66413FFF3FFFBC98180E9FEC9FA68" bold="true" box="[181,275,1509,1533]" italics="true" pageId="3" pageNumber="114">Female.</emphasis>
|
||
Sternite VIII (
|
||
<figureCitation id="7FA9A484FFF3FFFBC8F180EAFDC6FA68" box="[453,540,1510,1533]" captionStart="Fig" captionStartId="2.[151,182,1520,1539]" captionTargetBox="[336,1212,198,1467]" captionTargetId="figure-13@2.[299,1288,186,1477]" captionTargetPageId="2" captionText="Fig. 1. Lathrobium nomurai Nakane, 1955. A – habitus in dorsal view; B – labels of holotype; C – male sternite VII; D – male sternite VIII; E – female sternite VIII. Scale bars = 2.0 mm for A, 0.5 mm for C–E." figureDoi="http://doi.org/10.5281/zenodo.8005984" httpUri="https://zenodo.org/record/8005984/files/figure.png" pageId="3" pageNumber="114">Fig. 1E</figureCitation>
|
||
) elongate, densely fringed with numerous setae, post-median margin straight.
|
||
<emphasis id="D5E66413FFF3FFFBC9A3832AFE4AF9A8" bold="true" box="[151,400,1574,1597]" pageId="3" pageNumber="114">Differential diagnosis.</emphasis>
|
||
This species is similar to
|
||
<taxonomicName id="2092C382FFF3FFFBCB93832AFEEEF9C8" authorityName="Y. Watanabe" authorityYear="2001" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="fujimotoi">
|
||
<emphasis id="D5E66413FFF3FFFBCB93832AFEEEF9C8" italics="true" pageId="3" pageNumber="114">Lathrobium fujimotoi</emphasis>
|
||
</taxonomicName>
|
||
from Mt. Kurodake,
|
||
<collectingRegion id="255676E3FFF3FFFBCB29834AFD5CF9C8" box="[541,646,1606,1629]" country="Japan" name="Oita" pageId="3" pageNumber="114">Oita Pref.</collectingRegion>
|
||
in general appearance and aedeagal shape. However, this species can be easily distinguished from it by the combination of the following characteristics: 1) lateral side of ventral process of aedeagus straight; 2) internal sac without large sclerotized plate; 3) male sternite VIII slightly asymmetric and smaller than
|
||
<taxonomicName id="2092C382FFF3FFFBC87A820AFE13F888" authorityName="Y. Watanabe" authorityYear="2001" box="[334,457,1798,1821]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="fujimotoi">
|
||
<emphasis id="D5E66413FFF3FFFBC87A820AFE13F888" box="[334,457,1798,1821]" italics="true" pageId="3" pageNumber="114">L. fujimotoi</emphasis>
|
||
</taxonomicName>
|
||
(in
|
||
<taxonomicName id="2092C382FFF3FFFBC8C0820AFDAAF888" authorityName="Y. Watanabe" authorityYear="2001" box="[500,624,1798,1821]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="fujimotoi">
|
||
<emphasis id="D5E66413FFF3FFFBC8C0820AFDAAF888" box="[500,624,1798,1821]" italics="true" pageId="3" pageNumber="114">L. fujimotoi</emphasis>
|
||
</taxonomicName>
|
||
, each lateral side of ventral process is bulged at basal 1/3, the internal sac has a large sclerotized oval part and sternite VIII is symmetrical and larger).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF3FFFBC9A3828AFD3FF868" pageId="3" pageNumber="114" type="biology_ecology">
|
||
<paragraph id="E72DB801FFF3FFFBC9A3828AFD3FF868" blockId="3.[151,761,1190,2077]" pageId="3" pageNumber="114">
|
||
<emphasis id="D5E66413FFF3FFFBC9A3828AFF29F808" bold="true" box="[151,243,1926,1949]" pageId="3" pageNumber="114">Biology.</emphasis>
|
||
Some specimens of this species collected from Kadomae by authors were found in the upper hypogean zone along mountain stream, at an altitude of about
|
||
<specimenCount id="F1947388FFF3FFFBCB8882CAFD22F848" box="[700,760,1990,2013]" pageId="3" pageNumber="114" type="generic">
|
||
<quantity id="206A15E4FFF3FFFBCB8882CAFD2EF848" box="[700,756,1990,2013]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="114" unit="m" value="70.0">70 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
We dug laterally
|
||
<quantity id="206A15E4FFF3FFFBC87982EAFE62F868" box="[333,440,2022,2045]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="3" pageNumber="114" unit="cm" value="35.0" valueMax="50.0" valueMin="20.0">20–50 cm</quantity>
|
||
up the slope of this locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF3FFFBC9A38D0AFD6AF788" box="[151,688,2054,2077]" pageId="3" pageNumber="114" type="distribution">
|
||
<paragraph id="E72DB801FFF3FFFBC9A38D0AFD6AF788" blockId="3.[151,761,1190,2077]" box="[151,688,2054,2077]" pageId="3" pageNumber="114">
|
||
<emphasis id="D5E66413FFF3FFFBC9A38D0AFEF0F788" bold="true" box="[151,298,2054,2077]" pageId="3" pageNumber="114">Distribution.</emphasis>
|
||
<collectingCountry id="9F85F891FFF3FFFBC8048D0AFEB7F788" box="[304,365,2054,2077]" name="Japan" pageId="3" pageNumber="114">Japan</collectingCountry>
|
||
(
|
||
<collectingRegion id="255676E3FFF3FFFBC84E8D0AFE3DF788" box="[378,487,2054,2077]" country="Japan" name="Oita" pageId="3" pageNumber="114">Oita Pref.</collectingRegion>
|
||
, north of Kyushu).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF3FFFBCA0881A9FB42F9C9" pageId="3" pageNumber="114" type="discussion">
|
||
<paragraph id="E72DB801FFF3FFFBCA0881A9FBBBFA49" blockId="3.[828,1437,1189,1500]" pageId="3" pageNumber="114">
|
||
<emphasis id="D5E66413FFF3FFFBCA0881A9FC70FB29" bold="true" box="[828,938,1189,1212]" pageId="3" pageNumber="114">Remarks.</emphasis>
|
||
The aedeagus of the
|
||
<typeStatus id="382906A3FFF3FFFBCDA781A9FB2AFB29" box="[1171,1264,1189,1212]" pageId="3" pageNumber="114">holotype</typeStatus>
|
||
is not available for examination. The additional specimens examined here have the characteristics of the antennae and male sternite VIII corresponding to the
|
||
<typeStatus id="382906A3FFF3FFFBCD7D8009FB72FA89" box="[1097,1192,1285,1308]" pageId="3" pageNumber="114">holotype</typeStatus>
|
||
, and we hence consider them as conspecific with the
|
||
<typeStatus id="382906A3FFF3FFFBCD548029FB65FAA9" box="[1120,1215,1317,1340]" pageId="3" pageNumber="114">holotype</typeStatus>
|
||
. Their collection sites are near the type locality (Tsuruoka-Dô Cave, Saiki-City of
|
||
<collectingRegion id="255676E3FFF3FFFBCA6F8069FBDBFAE9" box="[859,1025,1381,1404]" country="Japan" name="Oita" pageId="3" pageNumber="114">Oita Prefecture</collectingRegion>
|
||
,
|
||
<collectingCountry id="9F85F891FFF3FFFBCD3A8069FB96FAE9" box="[1038,1100,1381,1404]" name="Japan" pageId="3" pageNumber="114">Japan</collectingCountry>
|
||
) and the identical morphology of their aedeagus indicates that they all belong to the same species. Our illustrations of the aedeagus are based on these newly collected specimens.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF3FFFBCAB38303FA86F9D7" blockId="3.[893,1372,1551,1628]" pageId="3" pageNumber="114">
|
||
<taxonomicName id="2092C382FFF3FFFBCAB38303FA8BF9B2" authority="Y. Watanabe, 2001" authorityName="Y. Watanabe" authorityYear="2001" box="[903,1361,1551,1575]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="fujimotoi">
|
||
<emphasis id="D5E66413FFF3FFFBCAB38303FA8BF9B2" bold="true" box="[903,1361,1551,1575]" pageId="3" pageNumber="114">
|
||
<emphasis id="D5E66413FFF3FFFBCAB38303FBADF9B2" bold="true" box="[903,1143,1551,1575]" italics="true" pageId="3" pageNumber="114">Lathrobium fujimotoi</emphasis>
|
||
Y.
|
||
<bibRefCitation id="8303C5F0FFF3FFFBCDAF831CFA8BF9B2" author="WATANABE Y." box="[1179,1361,1552,1575]" pageId="3" pageNumber="114" pagination="37 - 43" refId="ref8893" refString="WATANABE Y. 2001: Two new species of the group of Lathrobium nomurai (Coleoptera, Staphylinidae) from West Japan. Journal of the Speleological Society of Japan 26: 37 - 43." type="journal article" year="2001">Watanabe, 2001</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
[Japanese name: Fujimoto-ô-kobane-naga-hanekakushi]
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF3FFFBCD748345FB42F9C9" blockId="3.[893,1372,1551,1628]" box="[1088,1176,1609,1628]" pageId="3" pageNumber="114">
|
||
(
|
||
<figureCitation id="7FA9A484FFF3FFFBCD728345FBA7F9C9" box="[1094,1149,1609,1628]" captionStart="Fig" captionStartId="4.[151,182,1178,1197]" captionTargetBox="[271,1358,192,1134]" captionTargetId="figure-385@4.[193,1396,179,1152]" captionTargetPageId="4" captionText="Fig. 3. Male aedeagus of Lathrobium fujimotoi Y.Watanabe, 2001:A – ventral view; B – lateral view; C – dorsal view. Scale bar = 0.5 mm.Abbreviation: SP – large sclerotized plate of internal sac." figureDoi="http://doi.org/10.5281/zenodo.8005988" httpUri="https://zenodo.org/record/8005988/files/figure.png" pageId="3" pageNumber="114">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="7FA9A484FFF3FFFBCDB28345FB4BF9C9" box="[1158,1169,1609,1628]" captionStart="Fig" captionStartId="11.[151,182,1153,1172]" captionTargetBox="[254,1353,201,1126]" captionTargetId="figure-402@11.[207,1379,186,1137]" captionTargetPageId="11" captionText="Fig. 9. Map showing the distribution of the Lathrobium nomurai group in Kyushu, Japan. Black rhombus – L. nomurai Nakane, 1955; black triangle – L. fujimotoi Y. Watanabe, 2001; black circle – L. hikosanense Y. Watanabe, 1998; black square – L. yufuense sp. nov.; black pentagon – L. denchu sp. nov.; A–G – localities of L. hikosanense: A – Mt. Daisen; B – Mt. Hiko-san (type locality for L. hikosanense); C – Hiraodai; D – Mt. Buccho-zan; E – Mt. Tachibana-yama; F – Mt. Iwara-yama; G – Kinsenji Temple." figureDoi="http://doi.org/10.5281/zenodo.8006000" httpUri="https://zenodo.org/record/8006000/files/figure.png" pageId="3" pageNumber="114">9</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF3FFFCCA08837FFBFCF8AF" lastPageId="4" lastPageNumber="115" pageId="3" pageNumber="114" type="materials_examined">
|
||
<paragraph id="E72DB801FFF3FFFBCA08837FFC1BF943" blockId="3.[828,1437,1651,1803]" pageId="3" pageNumber="114">
|
||
<emphasis id="D5E66413FFF3FFFBCA08837FFBC5F913" bold="true" box="[828,1055,1651,1670]" pageId="3" pageNumber="114">Type material examined.</emphasis>
|
||
<materialsCitation id="57FAB25CFFF3FFFBCD11837FFC1BF943" collectingDate="1997-09-21" collectionCode="TUA" collectorName="Coll. H. Fujimoto & Y. Watanabe" country="Japan" location="Kyushu" pageId="3" pageNumber="114" specimenCount="2" stateProvince="Oita" typeStatus="holotype">
|
||
<typeStatus id="382906A3FFF3FFFBCD11837FFBA5F913" box="[1061,1151,1651,1670]" pageId="3" pageNumber="114">HOLOTYPE</typeStatus>
|
||
: 3,
|
||
<collectingCountry id="9F85F891FFF3FFFBCD92837FFB0DF913" box="[1190,1239,1651,1670]" name="Japan" pageId="3" pageNumber="114">Japan</collectingCountry>
|
||
<location id="E24DEEDAFFF3FFFBCDE9837FFAF8F913" LSID="urn:lsid:plazi:treatment:6F3B0917FFF1FFFECA9384F1FE59F780:E24DEEDAFFF3FFFBCDE9837FFAF8F913" box="[1245,1314,1651,1670]" country="Japan" name="Kyushu" pageId="3" pageNumber="114" stateProvince="Oita">Kyushu</location>
|
||
, “Mt. KURODAKE / Shônai-cho /
|
||
<collectingRegion id="255676E3FFF3FFFBCACC8382FBC7F934" box="[1016,1053,1678,1697]" country="Japan" name="Oita" pageId="3" pageNumber="114">Oita</collectingRegion>
|
||
/ Kyushu /
|
||
<collectingCountry id="9F85F891FFF3FFFBCD4B8382FB6AF934" box="[1151,1200,1678,1697]" name="Japan" pageId="3" pageNumber="114">Japan</collectingCountry>
|
||
/
|
||
<date id="932C9EC1FFF3FFFBCDF48382FA93F934" box="[1216,1353,1678,1697]" pageId="3" pageNumber="114" value="1997-09-21">
|
||
<collectingDate id="83686729FFF3FFFBCDF48382FA93F934" box="[1216,1353,1678,1697]" pageId="3" pageNumber="114" value="1997-09-21">Sept. 21st. 1997</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="4A67DDD7FFF3FFFBCC6C8382FC50F92E" pageId="3" pageNumber="114">Coll. H. Fujimoto</collectorName>
|
||
// [
|
||
<typeStatus id="382906A3FFF3FFFBCA9083A4FBC3F92E" box="[932,1049,1704,1723]" pageId="3" pageNumber="114">HOLOTYPE</typeStatus>
|
||
] /
|
||
<taxonomicName id="2092C382FFF3FFFBCD1A83A4FAF9F92E" authorityName="Y. Watanabe" authorityYear="2001" box="[1070,1315,1704,1723]" class="Insecta" family="Staphylinidae" genus="Lathrobium" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="fujimotoi">
|
||
<emphasis id="D5E66413FFF3FFFBCD1A83A4FB48F92E" box="[1070,1170,1704,1723]" italics="true" pageId="3" pageNumber="114">Lathrobium</emphasis>
|
||
(s. str.)
|
||
<emphasis id="D5E66413FFF3FFFBCDE283A4FAF9F92E" box="[1238,1315,1704,1723]" italics="true" pageId="3" pageNumber="114">fujimotoi</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<collectorName id="4A67DDD7FFF3FFFBCC0683A4FA47F92E" box="[1330,1437,1704,1723]" pageId="3" pageNumber="114">Y. Watanabe</collectorName>
|
||
/ 2001” (
|
||
<collectionCode id="818320C4FFF3FFFBCAB383CFFC6DF943" box="[903,951,1731,1750]" pageId="3" pageNumber="114">TUA</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF3FFFBCA0883D2FA5FF89E" blockId="3.[828,1437,1651,1803]" pageId="3" pageNumber="114">
|
||
<emphasis id="D5E66413FFF3FFFBCA0883D2FB85F964" bold="true" box="[828,1119,1758,1777]" pageId="3" pageNumber="114">Additional material examined.</emphasis>
|
||
1 3, “
|
||
<collectingCountry id="9F85F891FFF3FFFBCD9483D2FB3DF964" box="[1184,1255,1758,1777]" name="Japan" pageId="3" pageNumber="114">JAPAN</collectingCountry>
|
||
: Oita-ken / Asono / Shonai-cho / Yufu-shi / Mt. Kurodake /
|
||
<date id="932C9EC1FFF3FFFBCDBB83F4FB27F89E" box="[1167,1277,1784,1803]" pageId="3" pageNumber="114" value="2021-11-16">16. XI. 2021</date>
|
||
. / Y. Takagaki”.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF3FFFBCA088228FBCAF8EE" blockId="3.[828,1437,1828,2075]" pageId="3" pageNumber="114">
|
||
<emphasis id="D5E66413FFF3FFFBCA088228FBA3F8AE" bold="true" box="[828,1145,1828,1851]" pageId="3" pageNumber="114">Supplementary description.</emphasis>
|
||
BL =
|
||
<quantity id="206A15E4FFF3FFFBCDF58228FABAF8AE" box="[1217,1376,1828,1851]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" metricValueMax="1.44" metricValueMin="1.34" pageId="3" pageNumber="114" unit="mm" value="13.9" valueMax="14.4" valueMin="13.4">13.4–14.4 mm</quantity>
|
||
, FBL =
|
||
<quantity id="206A15E4FFF3FFFBCA678248FC0EF8CE" box="[851,980,1860,1883]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.6" metricValueMin="5.7" pageId="3" pageNumber="114" unit="mm" value="6.15" valueMax="6.6" valueMin="5.7">5.7–6.6 mm</quantity>
|
||
(n = 2). Coloration: Body reddish brown, moderately shining.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF3FFFBCA6E8288FB52F84E" blockId="3.[828,1437,1828,2075]" pageId="3" pageNumber="114">
|
||
Head suborbicular, HL =
|
||
<quantity id="206A15E4FFF3FFFBCD598288FAD1F80E" box="[1133,1291,1924,1947]" metricMagnitude="-3" metricUnit="m" metricValue="1.825" metricValueMax="1.9" metricValueMin="1.75" pageId="3" pageNumber="114" unit="mm" value="1.825" valueMax="1.9" valueMin="1.75">1.75–1.90 mm</quantity>
|
||
, HW = 1.75– 1.85, HW/HL = 0.99, widest at the basal 1/4 of head. Eyes very small, slightly protruding.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF3FFFCCA6E82E8FEDCFA0F" blockId="3.[828,1437,1828,2075]" lastBlockId="4.[151,761,1379,2074]" lastPageId="4" lastPageNumber="115" pageId="3" pageNumber="114">
|
||
Pronotum oblong-oval, with anterior margin wider than posterior one,
|
||
<collectionCode id="818320C4FFF3FFFBCAE18D08FC29F78E" box="[981,1011,2052,2075]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="3" pageNumber="114" type="Museum">PL</collectionCode>
|
||
= 2.00–
|
||
<quantity id="206A15E4FFF3FFFBCD788D08FB76F78E" box="[1100,1196,2052,2075]" metricMagnitude="-3" metricUnit="m" metricValue="2.23" pageId="3" pageNumber="114" unit="mm" value="2.23">2.23 mm</quantity>
|
||
,
|
||
<collectionCode id="818320C4FFF3FFFBCD838D08FB05F78E" box="[1207,1247,2052,2075]" country="Thailand" name="Paleontological Collections" pageId="3" pageNumber="114">PW</collectionCode>
|
||
=
|
||
<quantity id="206A15E4FFF3FFFBCDCF8D08FA42F78E" box="[1275,1432,2052,2075]" metricMagnitude="-3" metricUnit="m" metricValue="1.675" metricValueMax="1.75" metricValueMin="1.6" pageId="3" pageNumber="114" unit="mm" value="1.675" valueMax="1.75" valueMin="1.6">1.60–1.75 mm</quantity>
|
||
,
|
||
<collectionCode id="818320C4FFF4FFFCC9A3806FFF6CFAEF" box="[151,182,1379,1402]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="4" pageNumber="115" type="Museum">PL</collectionCode>
|
||
/
|
||
<collectionCode id="818320C4FFF4FFFCC98A806FFF3CFAEF" box="[190,230,1379,1402]" country="Thailand" name="Paleontological Collections" pageId="4" pageNumber="115">PW</collectionCode>
|
||
= 1.26,
|
||
<collectionCode id="818320C4FFF4FFFCC80B806FFE84FAEF" box="[319,350,1379,1402]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="4" pageNumber="115" type="Museum">PL</collectionCode>
|
||
/HL = 1.16,
|
||
<collectionCode id="818320C4FFF4FFFCC8D6806FFDD0FAEF" box="[482,522,1379,1402]" country="Thailand" name="Paleontological Collections" pageId="4" pageNumber="115">PW</collectionCode>
|
||
/HW = 0.93, widest at apical 1/5.
|
||
</paragraph>
|
||
<caption id="B3EDE889FFF4FFFCC9A38196FDD9FB5D" ID-DOI="http://doi.org/10.5281/zenodo.8005988" ID-Zenodo-Dep="8005988" httpUri="https://zenodo.org/record/8005988/files/figure.png" pageId="4" pageNumber="115" startId="4.[151,182,1178,1197]" targetBox="[271,1358,192,1134]" targetPageId="4" targetType="figure">
|
||
<paragraph id="E72DB801FFF4FFFCC9A38196FDD9FB5D" blockId="4.[151,1438,1178,1224]" pageId="4" pageNumber="115">
|
||
Fig. 3. Male aedeagus of
|
||
<taxonomicName id="2092C382FFF4FFFCC85E8196FD1FFB38" authority="Y. Watanabe, 2001" authorityName="Y. Watanabe" authorityYear="2001" box="[362,709,1178,1197]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="fujimotoi">
|
||
<emphasis id="D5E66413FFF4FFFCC85E8196FDFAFB38" box="[362,544,1178,1197]" italics="true" pageId="4" pageNumber="115">Lathrobium fujimotoi</emphasis>
|
||
Y.
|
||
<bibRefCitation id="8303C5F0FFF4FFFCCB0E8196FD1FFB38" author="WATANABE Y." box="[570,709,1178,1197]" pageId="4" pageNumber="115" pagination="37 - 43" refId="ref8893" refString="WATANABE Y. 2001: Two new species of the group of Lathrobium nomurai (Coleoptera, Staphylinidae) from West Japan. Journal of the Speleological Society of Japan 26: 37 - 43." type="journal article" year="2001">Watanabe, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
:A – ventral view; B – lateral view; C – dorsal view. Scale bar = 0.5 mm.Abbreviation: SP – large sclerotized plate of internal sac.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E72DB801FFF4FFFCC98180AFFD23F98F" blockId="4.[151,761,1379,2074]" pageId="4" pageNumber="115">
|
||
Elytra subtrapezoidal, widened posteriorly, EL =
|
||
<quantity id="206A15E4FFF4FFFCC9A380CFFEEDFA4F" box="[151,311,1475,1498]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" metricValueMax="1.2" metricValueMin="1.16" pageId="4" pageNumber="115" unit="mm" value="1.18" valueMax="1.2" valueMin="1.16">1.16–1.20 mm</quantity>
|
||
, EW =
|
||
<quantity id="206A15E4FFF4FFFCC8BB80CFFE29FA4F" box="[399,499,1475,1498]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="4" pageNumber="115" unit="mm" value="1.78">1.78 mm</quantity>
|
||
, EW/EL = 1.51, EL/
|
||
<collectionCode id="818320C4FFF4FFFCCBED80CFFD22FA4F" box="[729,760,1475,1498]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="4" pageNumber="115" type="Museum">PL</collectionCode>
|
||
= 0.56, EW/
|
||
<collectionCode id="818320C4FFF4FFFCC82A80EFFE9CFA6F" box="[286,326,1507,1530]" country="Thailand" name="Paleontological Collections" pageId="4" pageNumber="115">PW</collectionCode>
|
||
= 1.06; lateral margins almost straight; posterior corners rounded. Hind wing completely reduced.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF4FFFCC981832EFEA5F90F" blockId="4.[151,761,1379,2074]" pageId="4" pageNumber="115">
|
||
<emphasis id="D5E66413FFF4FFFCC981832EFF2EF9AF" bold="true" box="[181,244,1570,1594]" italics="true" pageId="4" pageNumber="115">Male.</emphasis>
|
||
Abdominal sternite VII transverse, with weak posteromedian impression; several setae at posteromedian area grown toward interior; sternite VIII with weak posteromedian impression.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF4FFFCC98183AFFBFCF8AF" blockId="4.[151,761,1379,2074]" lastBlockId="4.[828,1437,1379,2074]" pageId="4" pageNumber="115">
|
||
Aedeagus (
|
||
<figureCitation id="7FA9A484FFF4FFFCC81983AFFE7EF92F" box="[301,420,1699,1722]" captionStart="Fig" captionStartId="4.[151,182,1178,1197]" captionTargetBox="[271,1358,192,1134]" captionTargetId="figure-385@4.[193,1396,179,1152]" captionTargetPageId="4" captionText="Fig. 3. Male aedeagus of Lathrobium fujimotoi Y.Watanabe, 2001:A – ventral view; B – lateral view; C – dorsal view. Scale bar = 0.5 mm.Abbreviation: SP – large sclerotized plate of internal sac." figureDoi="http://doi.org/10.5281/zenodo.8005988" httpUri="https://zenodo.org/record/8005988/files/figure.png" pageId="4" pageNumber="115">Figs 3A–C</figureCitation>
|
||
)
|
||
<quantity id="206A15E4FFF4FFFCC88083AFFDC9F92F" box="[436,531,1699,1722]" metricMagnitude="-3" metricUnit="m" metricValue="3.28" pageId="4" pageNumber="115" unit="mm" value="3.28">3.28 mm</quantity>
|
||
long, elongate; basal part of aedeagus with V-shaped impression; ventral process well sclerotized, slightly asymmetrical in ventral view, gradually narrowed toward apical 1/4, strongly narrowed toward apex, curved ventrally from 1/2 to apex; apical part of ventral process straight; apex of ventral process with small projection; basal and middle parts of ventral process slightly thicker than apex in lateral view; each lateral side of ventral process weakly bulged at basal 1/3 area; part above basal foramen almost pentagonal, with two projections at lateral side; dorsal plate sclerotized, widest at apical 1/4, gradually widened to the widest point at apical 1/4, thus strongly contracted toward apex, with subacute apex; two tubes attached with apex of ventral process in between ventral process and dorsal plate; internal sac large, with sclerotized and ringed part, including large sclerotized plate in dorsal side; sclerotized plate in internal sac with some spine lines dorsally; apical area of internal sac folded in complex way.
|
||
<emphasis id="D5E66413FFF4FFFCCA08832FFBE3F9AF" bold="true" box="[828,1081,1571,1594]" pageId="4" pageNumber="115">Differential diagnosis.</emphasis>
|
||
This species is similar to other species of the
|
||
<taxonomicName id="2092C382FFF4FFFCCAFB834FFB99F9CF" authorityName="Nakane" authorityYear="1955" box="[975,1091,1603,1626]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="nomurai">
|
||
<emphasis id="D5E66413FFF4FFFCCAFB834FFB99F9CF" box="[975,1091,1603,1626]" italics="true" pageId="4" pageNumber="115">L. nomurai</emphasis>
|
||
</taxonomicName>
|
||
group in general appearance and aedeagus. However, this species can be easily distinguished from them by the combination of the following characteristics: 1) basal part of ventral process with V-shaped impression; 2) internal sac of aedeagus with large, sclerotized plate in dorsal side; 3) each lateral side of ventral process weakly bulged at basal 1/3 area; 4) apex of ventral process with small projection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF4FFFCCA08824FFA47F80F" pageId="4" pageNumber="115" type="biology_ecology">
|
||
<paragraph id="E72DB801FFF4FFFCCA08824FFA47F80F" blockId="4.[828,1437,1379,2074]" pageId="4" pageNumber="115">
|
||
<emphasis id="D5E66413FFF4FFFCCA08824FFC4DF8CF" bold="true" box="[828,919,1859,1882]" pageId="4" pageNumber="115">Biology.</emphasis>
|
||
<typeStatus id="382906A3FFF4FFFCCAAB824FFBD8F8CF" box="[927,1026,1859,1882]" pageId="4" pageNumber="115">Holotype</typeStatus>
|
||
and additional specimen were collected from the upper hypogean zone and under stones in a deciduous broadleaf forest, at an altitude of about
|
||
<specimenCount id="F1947388FFF4FFFCCC66828FFA47F80F" box="[1362,1437,1923,1946]" pageId="4" pageNumber="115" type="generic">
|
||
<quantity id="206A15E4FFF4FFFCCC66828FFA43F80F" box="[1362,1433,1923,1946]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="115" unit="m" value="850.0">850 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF4FFFCCA0882AFFA8FF82F" box="[828,1365,1955,1978]" pageId="4" pageNumber="115" type="distribution">
|
||
<paragraph id="E72DB801FFF4FFFCCA0882AFFA8FF82F" blockId="4.[828,1437,1379,2074]" box="[828,1365,1955,1978]" pageId="4" pageNumber="115">
|
||
<emphasis id="D5E66413FFF4FFFCCA0882AFFC14F82F" bold="true" box="[828,974,1955,1978]" pageId="4" pageNumber="115">Distribution.</emphasis>
|
||
<collectingCountry id="9F85F891FFF4FFFCCAE082AFFBCBF82F" box="[980,1041,1955,1978]" name="Japan" pageId="4" pageNumber="115">Japan</collectingCountry>
|
||
(
|
||
<collectingRegion id="255676E3FFF4FFFCCD2B82AFFB56F82F" box="[1055,1164,1955,1978]" country="Japan" name="Oita" pageId="4" pageNumber="115">Oita Pref.</collectingRegion>
|
||
, north of Kyushu).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF4FFFDCA0882CFFE29FE9C" lastPageId="5" lastPageNumber="116" pageId="4" pageNumber="115" type="discussion">
|
||
<paragraph id="E72DB801FFF4FFFCCA0882CFFB77F78F" blockId="4.[828,1437,1379,2074]" pageId="4" pageNumber="115">
|
||
<emphasis id="D5E66413FFF4FFFCCA0882CFFC70F84F" bold="true" box="[828,938,1987,2010]" pageId="4" pageNumber="115">Remarks.</emphasis>
|
||
This species has been reported only from Mt. Kurodake,
|
||
<collectingRegion id="255676E3FFF4FFFCCA8782EFFBC5F86F" box="[947,1055,2019,2042]" country="Japan" name="Oita" pageId="4" pageNumber="115">Oita Pref.</collectingRegion>
|
||
,
|
||
<collectingCountry id="9F85F891FFF4FFFCCD1F82EFFBB1F86F" box="[1067,1131,2019,2042]" name="Japan" pageId="4" pageNumber="115">Japan</collectingCountry>
|
||
. See
|
||
<bibRefCitation id="8303C5F0FFF4FFFCCD9082EFFAB8F86F" author="WATANABE Y." box="[1188,1378,2019,2042]" pageId="4" pageNumber="115" pagination="37 - 43" refId="ref8893" refString="WATANABE Y. 2001: Two new species of the group of Lathrobium nomurai (Coleoptera, Staphylinidae) from West Japan. Journal of the Speleological Society of Japan 26: 37 - 43." type="journal article" year="2001">WATANABE (2001)</bibRefCitation>
|
||
for a detailed description of the species.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF5FFFDC9E385B0FD69FF7A" blockId="5.[215,694,188,265]" pageId="5" pageNumber="116">
|
||
<taxonomicName id="2092C382FFF5FFFDC9E385B0FD6CFF41" authority="Y. Watanabe, 1998" authorityName="Y. Watanabe" authorityYear="1998" box="[215,694,188,212]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="116" phylum="Arthropoda" rank="species" species="hikosanese">
|
||
<emphasis id="D5E66413FFF5FFFDC9E385B0FD6CFF41" bold="true" box="[215,694,188,212]" pageId="5" pageNumber="116">
|
||
<emphasis id="D5E66413FFF5FFFDC9E385B0FE01FF41" bold="true" box="[215,475,188,212]" italics="true" pageId="5" pageNumber="116">Lathrobium hikosanese</emphasis>
|
||
Y.
|
||
<bibRefCitation id="8303C5F0FFF5FFFDCB3485B1FD6CFF41" author="WATANABE Y." box="[512,694,189,212]" pageId="5" pageNumber="116" pagination="85 - 98" refId="ref8856" refString="WATANABE Y. 1998: Five new species of the Lathrobium (s. str.) nomurai group (Coleoptera, Staphylinidae) from Japan. Elytra 26 (1): 85 - 98." type="journal article" year="1998">Watanabe, 1998</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
[Japanese name: Hikosan-ô-kobane-naga-hanekakushi]
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF5FFFDC8AF85FAFE29FE9C" blockId="5.[215,694,188,265]" box="[411,499,246,265]" pageId="5" pageNumber="116">
|
||
(
|
||
<figureCitation id="7FA9A484FFF5FFFDC89485FAFE02FE9C" box="[416,472,246,265]" captionStart="Fig" captionStartId="5.[150,181,2010,2029]" captionTargetBox="[289,1292,705,1970]" captionTargetId="figure-15@5.[261,1325,667,1990]" captionTargetPageId="5" captionText="Fig. 4. Male aedeagus of Lathrobium hikosanense Y. Watanabe, 1998 in each locality.A–C – Mt. Hiko-san (type locality): A – ventral view; B – lateral view; C – dorsal view. D, J – Mt. Daisen; E, K – Hiraodai; F, L – Mt. Buccho-zan; G, M – Mt. Tachibana-yama; H, N – Mt. Iwara-yama; I, O – Kinsenji Temple. Scale bars = 0.5 mm. Each locality referred to Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.8005990" httpUri="https://zenodo.org/record/8005990/files/figure.png" pageId="5" pageNumber="116">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="7FA9A484FFF5FFFDC8D585FAFE36FE9C" box="[481,492,246,265]" captionStart="Fig" captionStartId="11.[151,182,1153,1172]" captionTargetBox="[254,1353,201,1126]" captionTargetId="figure-402@11.[207,1379,186,1137]" captionTargetPageId="11" captionText="Fig. 9. Map showing the distribution of the Lathrobium nomurai group in Kyushu, Japan. Black rhombus – L. nomurai Nakane, 1955; black triangle – L. fujimotoi Y. Watanabe, 2001; black circle – L. hikosanense Y. Watanabe, 1998; black square – L. yufuense sp. nov.; black pentagon – L. denchu sp. nov.; A–G – localities of L. hikosanense: A – Mt. Daisen; B – Mt. Hiko-san (type locality for L. hikosanense); C – Hiraodai; D – Mt. Buccho-zan; E – Mt. Tachibana-yama; F – Mt. Iwara-yama; G – Kinsenji Temple." figureDoi="http://doi.org/10.5281/zenodo.8006000" httpUri="https://zenodo.org/record/8006000/files/figure.png" pageId="5" pageNumber="116">9</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF5FFFEC9A2842CFD59FCE0" lastPageId="6" lastPageNumber="117" pageId="5" pageNumber="116" type="materials_examined">
|
||
<paragraph id="E72DB801FFF5FFFDC9A2842CFE02FD1B" blockId="5.[150,760,288,654]" pageId="5" pageNumber="116">
|
||
<emphasis id="D5E66413FFF5FFFDC9A2842CFE9EFEA6" bold="true" box="[150,324,288,307]" pageId="5" pageNumber="116">Material examined.</emphasis>
|
||
1 3, “Mt.Hiko / Soeda-Town / FukuokaPref./ [
|
||
<collectionCode id="818320C4FFF5FFFDCBFF842CFD28FEA6" box="[715,754,288,307]" pageId="5" pageNumber="116">JPN</collectionCode>
|
||
] /
|
||
<date id="932C9EC1FFF5FFFDC9968437FECEFEDB" box="[162,276,315,334]" pageId="5" pageNumber="116" value="2018-10-21">2018. 10. 21.</date>
|
||
/ H. Tanaka leg.” (
|
||
<collectionCode id="818320C4FFF5FFFDC88C8437FE34FEDB" box="[440,494,315,334]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="116" type="Herbarium">KUM</collectionCode>
|
||
); 1 3,
|
||
<specimenCount id="F1947388FFF5FFFDCB1D8437FD90FEDB" box="[553,586,315,334]" pageId="5" pageNumber="116" type="female">1 ♀</specimenCount>
|
||
, “[
|
||
<collectingCountry id="9F85F891FFF5FFFDCB568437FD76FEDB" box="[610,684,315,334]" name="Japan" pageId="5" pageNumber="116">JAPAN</collectingCountry>
|
||
]: Fukuoka-ken. / Soeda-machi. / Hikosan. /
|
||
<date id="932C9EC1FFF5FFFDC8E78459FDE0FEFD" box="[467,570,341,360]" pageId="5" pageNumber="116" value="2019-06-08">
|
||
8.
|
||
<collectionCode id="818320C4FFF5FFFDC8DE8459FDD9FEFD" box="[490,515,341,360]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="116">VI</collectionCode>
|
||
. 2019
|
||
</date>
|
||
. / Tsubasa NOZAKI” (
|
||
<collectionCode id="818320C4FFF5FFFDC9A9847CFF09FE16" box="[157,211,368,387]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="116" type="Herbarium">KUM</collectionCode>
|
||
); 1 3, “
|
||
<collectingCountry id="9F85F891FFF5FFFDC823847CFE80FE16" box="[279,346,368,387]" name="Japan" pageId="5" pageNumber="116">JAPAN</collectingCountry>
|
||
/ Fukuoka-ken / Umi-machi / Mt. Buccho /
|
||
<date id="932C9EC1FFF5FFFDCBE8847CFF05FE0B" pageId="5" pageNumber="116" value="2015-10-25">25. X. 2015</date>
|
||
. / Naomichi Tsuji leg.” (
|
||
<collectionCode id="818320C4FFF5FFFDC8828487FE36FE0B" box="[438,492,395,414]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="116" type="Herbarium">KUM</collectionCode>
|
||
); 1 3, “
|
||
<collectingCountry id="9F85F891FFF5FFFDCB048487FDA9FE0B" box="[560,627,395,414]" name="Japan" pageId="5" pageNumber="116">JAPAN</collectingCountry>
|
||
/ Fukuoka-ken / Itoshima-shi / Mt. Iwara-yama (alt.
|
||
<quantity id="206A15E4FFF5FFFDC8E084A9FDD6FE2D" box="[468,524,421,440]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="116" unit="m" value="400.0">400 m</quantity>
|
||
) /
|
||
<date id="932C9EC1FFF5FFFDCB1784A9FD55FE2D" box="[547,655,421,440]" pageId="5" pageNumber="116" value="2014-04-27">27. IV. 2014</date>
|
||
. / Naomichi Tsuji leg.” (
|
||
<collectionCode id="818320C4FFF5FFFDC9C884CCFEE8FE46" box="[252,306,448,467]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="116" type="Herbarium">KUM</collectionCode>
|
||
); 1 3, “
|
||
<collectingCountry id="9F85F891FFF5FFFDC84284CCFE66FE46" box="[374,444,448,467]" name="Japan" pageId="5" pageNumber="116">JAPAN</collectingCountry>
|
||
: Fukuoka-ken / Mizunashi /
|
||
<date id="932C9EC1FFF5FFFDCB8084CCFF19FE7B" pageId="5" pageNumber="116" value="2019-07-15">15. VII. 2019</date>
|
||
. / Tanaka” (
|
||
<collectionCode id="818320C4FFF5FFFDC81F84D7FEBBFE7B" box="[299,353,475,494]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="116" type="Herbarium">KUM</collectionCode>
|
||
); 13, “Mt. Tachibana / Higashi-ku /
|
||
<collectingRegion id="255676E3FFF5FFFDCB9484D7FD31FE7B" box="[672,747,475,494]" country="Japan" name="Fukuoka" pageId="5" pageNumber="116">Fukuoka</collectingRegion>
|
||
/
|
||
<collectingRegion id="255676E3FFF5FFFDC9A284F9FEC8FD9D" box="[150,274,501,520]" country="Japan" name="Fukuoka" pageId="5" pageNumber="116">Fukuoka Pref.</collectingRegion>
|
||
/
|
||
<collectingCountry id="9F85F891FFF5FFFDC81784F9FEB9FD9D" box="[291,355,501,520]" name="Japan" pageId="5" pageNumber="116">JAPAN</collectingCountry>
|
||
/
|
||
<date id="932C9EC1FFF5FFFDC84284F9FE0EFD9D" box="[374,468,501,520]" pageId="5" pageNumber="116" value="2022-10-06">6. X. 2022</date>
|
||
. / Naomichi
|
||
<collectionCode id="818320C4FFF5FFFDCB7084F9FDA6FD9D" box="[580,636,501,520]" pageId="5" pageNumber="116">TSUJI</collectionCode>
|
||
leg.” (
|
||
<collectionCode id="818320C4FFF5FFFDCB8384F9FD37FD9D" box="[695,749,501,520]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="116" type="Herbarium">KUM</collectionCode>
|
||
); 1 3, “Mt. Daisen / Kujyu
|
||
<collectingRegion id="255676E3FFF5FFFDC841871CFE40FDB6" box="[373,410,528,547]" country="Japan" name="Oita" pageId="5" pageNumber="116">Oita</collectingRegion>
|
||
/
|
||
<date id="932C9EC1FFF5FFFDC89F871CFDD0FDB6" box="[427,522,528,547]" pageId="5" pageNumber="116" value="1983-06-05">5. Vi. 1983</date>
|
||
/ S. Nomura” (
|
||
<collectionCode id="818320C4FFF5FFFDCBB8871CFD18FDB6" box="[652,706,528,547]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="116" type="Herbarium">KUM</collectionCode>
|
||
); 1 3, “[
|
||
<collectingCountry id="9F85F891FFF5FFFDC9978727FF30FDAB" box="[163,234,555,574]" name="Japan" pageId="5" pageNumber="116">JAPAN</collectingCountry>
|
||
] / Fukuoka-ken / Kitakyûshû-shi / Kokura-Minami-ku / Hiraodai /
|
||
<geoCoordinate id="82A6DEC6FFF5FFFDC9FA8749FEFBFDCD" box="[206,289,581,600]" degrees="33.769" direction="north" orientation="latitude" pageId="5" pageNumber="116" precision="55" value="33.769">33.769°N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="82A6DEC6FFF5FFFDC8008749FE55FDCD" box="[308,399,581,600]" degrees="130.908" direction="east" orientation="longitude" pageId="5" pageNumber="116" precision="55" value="130.908">130.908°E</geoCoordinate>
|
||
/
|
||
<date id="932C9EC1FFF5FFFDC8968749FDD3FDCD" box="[418,521,581,600]" pageId="5" pageNumber="116" value="2021-05-22">
|
||
22.
|
||
<collectionCode id="818320C4FFF5FFFDC8F78749FE08FDCD" box="[451,466,581,600]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="116" type="Museum">V</collectionCode>
|
||
. 2021
|
||
</date>
|
||
. / S.
|
||
<collectionCode id="818320C4FFF5FFFDCB038749FDA1FDCD" box="[567,635,581,600]" pageId="5" pageNumber="116">INOUE</collectionCode>
|
||
” (
|
||
<collectionCode id="818320C4FFF5FFFDCBBB8749FD1FFDCD" box="[655,709,581,600]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="116" type="Herbarium">KUM</collectionCode>
|
||
); 1 3 “[
|
||
<collectingCountry id="9F85F891FFF5FFFDC997876CFF37FDE6" box="[163,237,608,627]" name="Japan" pageId="5" pageNumber="116">JAPAN</collectingCountry>
|
||
]: Nagasaki-ken / Isahaya-shi / Kinsenji / [
|
||
<geoCoordinate id="82A6DEC6FFF5FFFDCB7A876CFD42FDE6" box="[590,664,608,627]" degrees="32.9739" direction="north" orientation="latitude" pageId="5" pageNumber="116" precision="5" value="32.9739">32.9739</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="82A6DEC6FFF5FFFDCB94876CFD28FDE6" box="[672,754,608,627]" degrees="130.0879" direction="east" orientation="longitude" pageId="5" pageNumber="116" precision="5" value="130.0879">130.0879</geoCoordinate>
|
||
] /
|
||
<date id="932C9EC1FFF5FFFDC9968777FF24FD1B" box="[162,254,635,654]" pageId="5" pageNumber="116" value="2022-10-01">1. X. 2022</date>
|
||
. / TANAKA H.” (
|
||
<collectionCode id="818320C4FFF5FFFDC8A38777FE17FD1B" box="[407,461,635,654]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="116" type="Herbarium">KUM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF5FFFDCA0885B1FB12FE81" blockId="5.[828,1437,189,660]" pageId="5" pageNumber="116">
|
||
<emphasis id="D5E66413FFF5FFFDCA0885B1FBA3FF41" bold="true" box="[828,1145,189,212]" pageId="5" pageNumber="116">Supplementary description.</emphasis>
|
||
BL =
|
||
<quantity id="206A15E4FFF5FFFDCDF685B1FABAFF41" box="[1218,1376,189,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.275" metricValueMax="1.39" metricValueMin="1.16" pageId="5" pageNumber="116" unit="mm" value="12.75" valueMax="13.9" valueMin="11.6">11.6–13.9 mm</quantity>
|
||
, FBL =
|
||
<quantity id="206A15E4FFF5FFFDCA6685D1FC09FF61" box="[850,979,221,244]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.6" metricValueMin="4.9" pageId="5" pageNumber="116" unit="mm" value="5.25" valueMax="5.6" valueMin="4.9">4.9–5.6 mm</quantity>
|
||
(n = 10). Coloration: Body reddish brown to reddish black, moderately shining.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF5FFFDCA6E8411FB52FEE1" blockId="5.[828,1437,189,660]" pageId="5" pageNumber="116">
|
||
Head suborbicular, HL =
|
||
<quantity id="206A15E4FFF5FFFDCD598411FAD1FEA1" box="[1133,1291,285,308]" metricMagnitude="-3" metricUnit="m" metricValue="1.555" metricValueMax="1.68" metricValueMin="1.43" pageId="5" pageNumber="116" unit="mm" value="1.555" valueMax="1.68" valueMin="1.43">1.43–1.68 mm</quantity>
|
||
, HW = 1.40– 1.65, HW/HL = 0.96, widest at the basal 1/3 of head. Eyes very small, slightly protruding.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF5FFFDCA6E8471FC71FE61" blockId="5.[828,1437,189,660]" pageId="5" pageNumber="116">
|
||
Pronotum oblong-oval, with anterior margin wider than posterior one,
|
||
<collectionCode id="818320C4FFF5FFFDCAE18491FC29FE21" box="[981,1011,413,436]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="116" type="Museum">PL</collectionCode>
|
||
=
|
||
<quantity id="206A15E4FFF5FFFDCD3B8491FB76FE21" box="[1039,1196,413,436]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.95" metricValueMin="1.75" pageId="5" pageNumber="116" unit="mm" value="1.85" valueMax="1.95" valueMin="1.75">1.75–1.95 mm</quantity>
|
||
,
|
||
<collectionCode id="818320C4FFF5FFFDCD838491FB05FE21" box="[1207,1247,413,436]" country="Thailand" name="Paleontological Collections" pageId="5" pageNumber="116">PW</collectionCode>
|
||
=
|
||
<quantity id="206A15E4FFF5FFFDCDCF8491FA42FE21" box="[1275,1432,413,436]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="5" pageNumber="116" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.30–1.50 mm</quantity>
|
||
,
|
||
<collectionCode id="818320C4FFF5FFFDCA0884B1FC81FE41" box="[828,859,445,468]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="116" type="Museum">PL</collectionCode>
|
||
/
|
||
<collectionCode id="818320C4FFF5FFFDCA5684B1FC50FE41" box="[866,906,445,468]" country="Thailand" name="Paleontological Collections" pageId="5" pageNumber="116">PW</collectionCode>
|
||
= 1.35,
|
||
<collectionCode id="818320C4FFF5FFFDCAD084B1FBD9FE41" box="[996,1027,445,468]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="116" type="Museum">PL</collectionCode>
|
||
/HL = 1.20,
|
||
<collectionCode id="818320C4FFF5FFFDCDB384B1FB75FE41" box="[1159,1199,445,468]" country="Thailand" name="Paleontological Collections" pageId="5" pageNumber="116">PW</collectionCode>
|
||
/HW = 0.92, widest at apical 1/4.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF5FFFDCA6E84F1FC13FD01" blockId="5.[828,1437,189,660]" pageId="5" pageNumber="116">
|
||
Elytra subtrapezoidal, widened posteriorly, EL = 1.00–
|
||
<quantity id="206A15E4FFF5FFFDCA4E8711FC3BFDA1" box="[890,993,541,564]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="116" unit="mm" value="1.25">1.25 mm</quantity>
|
||
, EW =
|
||
<quantity id="206A15E4FFF5FFFDCD088711FB3BFDA1" box="[1084,1249,541,564]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.65" metricValueMin="1.4" pageId="5" pageNumber="116" unit="mm" value="1.525" valueMax="1.65" valueMin="1.4">1.40–1.65 mm</quantity>
|
||
, EW/EL = 1.38, EL/
|
||
<collectionCode id="818320C4FFF5FFFDCA508731FC59FDC1" box="[868,899,573,596]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="116" type="Museum">PL</collectionCode>
|
||
= 0.59, EW/
|
||
<collectionCode id="818320C4FFF5FFFDCD2C8731FB9AFDC1" box="[1048,1088,573,596]" country="Thailand" name="Paleontological Collections" pageId="5" pageNumber="116">PW</collectionCode>
|
||
= 1.10; lateral margins almost straight; posterior corners rounded. Hind wing completely reduced.
|
||
</paragraph>
|
||
<caption id="B3EDE889FFF5FFFDC9A282D6FD7CF7B7" ID-DOI="http://doi.org/10.5281/zenodo.8005990" ID-Zenodo-Dep="8005990" httpUri="https://zenodo.org/record/8005990/files/figure.png" pageId="5" pageNumber="116" startId="5.[150,181,2010,2029]" targetBox="[289,1292,705,1970]" targetPageId="5" targetType="figure">
|
||
<paragraph id="E72DB801FFF5FFFDC9A282D6FD7CF7B7" blockId="5.[150,1435,2010,2082]" pageId="5" pageNumber="116">
|
||
Fig. 4. Male aedeagus of
|
||
<taxonomicName id="2092C382FFF5FFFDC85A82D6FD3EF878" authority="Y. Watanabe, 1998" authorityName="Y. Watanabe" authorityYear="1998" box="[366,740,2010,2029]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="116" phylum="Arthropoda" rank="species" species="hikosanense">
|
||
<emphasis id="D5E66413FFF5FFFDC85A82D6FDE5F878" box="[366,575,2010,2029]" italics="true" pageId="5" pageNumber="116">Lathrobium hikosanense</emphasis>
|
||
Y.
|
||
<bibRefCitation id="8303C5F0FFF5FFFDCB6F82D6FD3EF878" author="WATANABE Y." box="[603,740,2010,2029]" pageId="5" pageNumber="116" pagination="85 - 98" refId="ref8856" refString="WATANABE Y. 1998: Five new species of the Lathrobium (s. str.) nomurai group (Coleoptera, Staphylinidae) from Japan. Elytra 26 (1): 85 - 98." type="journal article" year="1998">Watanabe, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
in each locality.A–C – Mt. Hiko-san (type locality): A – ventral view; B – lateral view; C – dorsal view. D, J – Mt. Daisen; E, K – Hiraodai; F, L – Mt. Buccho-zan; G, M – Mt. Tachibana-yama; H, N – Mt. Iwara-yama; I, O – Kinsenji Temple. Scale bars = 0.5 mm. Each locality referred to Fig. 9.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E72DB801FFF6FFFEC98185B1FEFAFEC0" blockId="6.[151,761,189,2069]" pageId="6" pageNumber="117">
|
||
<emphasis id="D5E66413FFF6FFFEC98185B1FF2EFF40" bold="true" box="[181,244,189,213]" italics="true" pageId="6" pageNumber="117">Male.</emphasis>
|
||
Sternite VII transverse, with weak posteromedian impression; sternite VIII with posteromedian impression; each side of posteromedian impression protruded weakly; lateral area of sternite VIII with slightly longer setae than median area.
|
||
</paragraph>
|
||
<paragraph id="E72DB801FFF6FFFEC9818452FD59FCE0" blockId="6.[151,761,189,2069]" pageId="6" pageNumber="117">
|
||
Aedeagus (
|
||
<figureCitation id="7FA9A484FFF6FFFEC8188452FE79FEE0" box="[300,419,350,373]" captionStart="Fig" captionStartId="5.[150,181,2010,2029]" captionTargetBox="[289,1292,705,1970]" captionTargetId="figure-15@5.[261,1325,667,1990]" captionTargetPageId="5" captionText="Fig. 4. Male aedeagus of Lathrobium hikosanense Y. Watanabe, 1998 in each locality.A–C – Mt. Hiko-san (type locality): A – ventral view; B – lateral view; C – dorsal view. D, J – Mt. Daisen; E, K – Hiraodai; F, L – Mt. Buccho-zan; G, M – Mt. Tachibana-yama; H, N – Mt. Iwara-yama; I, O – Kinsenji Temple. Scale bars = 0.5 mm. Each locality referred to Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.8005990" httpUri="https://zenodo.org/record/8005990/files/figure.png" pageId="6" pageNumber="117">Figs 4A–C</figureCitation>
|
||
)
|
||
<quantity id="206A15E4FFF6FFFEC8878452FDC8FEE0" box="[435,530,350,373]" metricMagnitude="-3" metricUnit="m" metricValue="2.27" pageId="6" pageNumber="117" unit="mm" value="2.27">2.27 mm</quantity>
|
||
long, oblong; ventral process well sclerotized, asymmetric in ventral view, gradually narrowed toward 1/2, thus gradually narrowed toward apex, curved ventrally from 1/2 to apex; apical part of ventral process elongate, curved toward left side in ventral view; apex of ventral process with small projection; ventral process from basal to about 1/2 of area in ventral view two times thicker than that of apical part; basal and middle parts of ventral process ball-shaped, thicker than apical part in lateral view; part above basal foramen triangular, with two projections at lateral side; surrounding of basal foramen sclerotized; dorsal plate sclerotized, triangular with base at anterior margin, contracted toward apex, with acute apex; internal sac large, with sclerotized and ringed part; basal area of internal sac including two sclerotized long–narrow parts; each sclerotized part formed by gathering some thin plates; middle and apical area of internal sac flattened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF6FFFEC9A38672FE4CFAA0" pageId="6" pageNumber="117" type="diagnosis">
|
||
<paragraph id="E72DB801FFF6FFFEC9A38672FE4CFAA0" blockId="6.[151,761,189,2069]" pageId="6" pageNumber="117">
|
||
<emphasis id="D5E66413FFF6FFFEC9A38672FE7EFC00" bold="true" box="[151,420,894,917]" pageId="6" pageNumber="117">Differential diagnosis.</emphasis>
|
||
This species is similar to
|
||
<taxonomicName id="2092C382FFF6FFFECBD58672FDD3FC20" authority="Y. Watanabe, 1998" authorityName="Y. Watanabe" authorityYear="1998" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="daisensanum">
|
||
<emphasis id="D5E66413FFF6FFFECBD58672FEFDFC20" italics="true" pageId="6" pageNumber="117">L. daisensanum</emphasis>
|
||
Y.
|
||
<bibRefCitation id="8303C5F0FFF6FFFEC8668692FDD3FC20" author="WATANABE Y." box="[338,521,926,949]" pageId="6" pageNumber="117" pagination="85 - 98" refId="ref8856" refString="WATANABE Y. 1998: Five new species of the Lathrobium (s. str.) nomurai group (Coleoptera, Staphylinidae) from Japan. Elytra 26 (1): 85 - 98." type="journal article" year="1998">Watanabe, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
, a member of the
|
||
<taxonomicName id="2092C382FFF6FFFECBD68692FF2BFC40" authorityName="Nakane" authorityYear="1955" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="nomurai">
|
||
<emphasis id="D5E66413FFF6FFFECBD68692FF2BFC40" italics="true" pageId="6" pageNumber="117">L. nomurai</emphasis>
|
||
</taxonomicName>
|
||
group described from Mt. Daisen-zan,
|
||
<collectingRegion id="255676E3FFF6FFFECB9486B2FF14FC60" country="Japan" name="Kagawa" pageId="6" pageNumber="117">Kagawa Pref.</collectingRegion>
|
||
, Shikoku, in general appearance and aedeagal shape. However, this species can be easily distinguished from it by the combination of the following characteristics: 1) dorsal plate of aedeagus triangular with base at anterior margin; 2) apical part of ventral process curved toward left side in ventral view; 3) middle area of ventral process of aedeagus gradually narrowed toward apex (while in
|
||
<taxonomicName id="2092C382FFF6FFFECBD68192FEF8FB40" authorityName="Y. Watanabe" authorityYear="1998" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="daisensanum">
|
||
<emphasis id="D5E66413FFF6FFFECBD68192FEF8FB40" italics="true" pageId="6" pageNumber="117">L. daisensanum</emphasis>
|
||
</taxonomicName>
|
||
, apex of dorsal plate is truncate, apical part of ventral process of aedeagus is straight in ventral view, and middle area of ventral process of aedeagus is strongly narrowed toward apex).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF6FFFEC9A38032FD22F980" pageId="6" pageNumber="117" type="description">
|
||
<paragraph id="E72DB801FFF6FFFEC9A38032FD22F980" blockId="6.[151,761,189,2069]" pageId="6" pageNumber="117">
|
||
<emphasis id="D5E66413FFF6FFFEC9A38032FEDCFAC0" bold="true" box="[151,262,1342,1365]" pageId="6" pageNumber="117">Variation.</emphasis>
|
||
Size and shape of aedeagus tend to vary by locality (
|
||
<figureCitation id="7FA9A484FFF6FFFEC9FC8052FED2FAE0" box="[200,264,1374,1397]" captionStart="Fig" captionStartId="5.[150,181,2010,2029]" captionTargetBox="[289,1292,705,1970]" captionTargetId="figure-15@5.[261,1325,667,1990]" captionTargetPageId="5" captionText="Fig. 4. Male aedeagus of Lathrobium hikosanense Y. Watanabe, 1998 in each locality.A–C – Mt. Hiko-san (type locality): A – ventral view; B – lateral view; C – dorsal view. D, J – Mt. Daisen; E, K – Hiraodai; F, L – Mt. Buccho-zan; G, M – Mt. Tachibana-yama; H, N – Mt. Iwara-yama; I, O – Kinsenji Temple. Scale bars = 0.5 mm. Each locality referred to Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.8005990" httpUri="https://zenodo.org/record/8005990/files/figure.png" pageId="6" pageNumber="117">Fig. 4</figureCitation>
|
||
). It can be observed that individuals in the west tend to have larger aedeagus than those in the east.Although tendency cannot be observed, thickness and degree of curvature on apical part of ventral process also vary in each region; the smallest in Hiraodai,
|
||
<collectingRegion id="255676E3FFF6FFFEC8EB80D2FDABFA60" box="[479,625,1502,1525]" country="Japan" name="Fukuoka" pageId="6" pageNumber="117">Fukuoka Pref.</collectingRegion>
|
||
(
|
||
<figureCitation id="7FA9A484FFF6FFFECB4880D2FD36FA60" box="[636,748,1502,1525]" captionStart="Fig" captionStartId="5.[150,181,2010,2029]" captionTargetBox="[289,1292,705,1970]" captionTargetId="figure-15@5.[261,1325,667,1990]" captionTargetPageId="5" captionText="Fig. 4. Male aedeagus of Lathrobium hikosanense Y. Watanabe, 1998 in each locality.A–C – Mt. Hiko-san (type locality): A – ventral view; B – lateral view; C – dorsal view. D, J – Mt. Daisen; E, K – Hiraodai; F, L – Mt. Buccho-zan; G, M – Mt. Tachibana-yama; H, N – Mt. Iwara-yama; I, O – Kinsenji Temple. Scale bars = 0.5 mm. Each locality referred to Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.8005990" httpUri="https://zenodo.org/record/8005990/files/figure.png" pageId="6" pageNumber="117">Figs 4E, K</figureCitation>
|
||
), the largest in Mt. Iwara-yama,
|
||
<collectingRegion id="255676E3FFF6FFFEC8E180F2FDB3F980" box="[469,617,1534,1557]" country="Japan" name="Fukuoka" pageId="6" pageNumber="117">Fukuoka Pref.</collectingRegion>
|
||
(
|
||
<figureCitation id="7FA9A484FFF6FFFECB4180F2FD31F980" box="[629,747,1534,1557]" captionStart="Fig" captionStartId="5.[150,181,2010,2029]" captionTargetBox="[289,1292,705,1970]" captionTargetId="figure-15@5.[261,1325,667,1990]" captionTargetPageId="5" captionText="Fig. 4. Male aedeagus of Lathrobium hikosanense Y. Watanabe, 1998 in each locality.A–C – Mt. Hiko-san (type locality): A – ventral view; B – lateral view; C – dorsal view. D, J – Mt. Daisen; E, K – Hiraodai; F, L – Mt. Buccho-zan; G, M – Mt. Tachibana-yama; H, N – Mt. Iwara-yama; I, O – Kinsenji Temple. Scale bars = 0.5 mm. Each locality referred to Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.8005990" httpUri="https://zenodo.org/record/8005990/files/figure.png" pageId="6" pageNumber="117">Figs 4H, N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF6FFFEC9A38312FEA8F900" pageId="6" pageNumber="117" type="biology_ecology">
|
||
<paragraph id="E72DB801FFF6FFFEC9A38312FEA8F900" blockId="6.[151,761,189,2069]" pageId="6" pageNumber="117">
|
||
<emphasis id="D5E66413FFF6FFFEC9A38312FF2EF9A0" bold="true" box="[151,244,1566,1589]" pageId="6" pageNumber="117">Biology.</emphasis>
|
||
Examined specimens were collected from the upper hypogean zone or under stones. Compared to other species of
|
||
<taxonomicName id="2092C382FFF6FFFEC8308352FEA3F9E0" authorityName="Nakane" authorityYear="1955" box="[260,377,1630,1653]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="nomurai">
|
||
<emphasis id="D5E66413FFF6FFFEC8308352FEA3F9E0" box="[260,377,1630,1653]" italics="true" pageId="6" pageNumber="117">L. nomurai</emphasis>
|
||
</taxonomicName>
|
||
group, this species tends to be found in a shallow ground.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF6FFFEC9A38392FDE1F920" box="[151,571,1694,1717]" pageId="6" pageNumber="117" type="distribution">
|
||
<paragraph id="E72DB801FFF6FFFEC9A38392FDE1F920" blockId="6.[151,761,189,2069]" box="[151,571,1694,1717]" pageId="6" pageNumber="117">
|
||
<emphasis id="D5E66413FFF6FFFEC9A38392FEF0F920" bold="true" box="[151,298,1694,1717]" pageId="6" pageNumber="117">Distribution.</emphasis>
|
||
<collectingCountry id="9F85F891FFF6FFFEC8048392FEB7F920" box="[304,365,1694,1717]" name="Japan" pageId="6" pageNumber="117">Japan</collectingCountry>
|
||
(north of Kyushu).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF88EB8AFFF6FFFEC9A383B2FE59F780" pageId="6" pageNumber="117" type="discussion">
|
||
<paragraph id="E72DB801FFF6FFFEC9A383B2FE59F780" blockId="6.[151,761,189,2069]" pageId="6" pageNumber="117">
|
||
<emphasis id="D5E66413FFF6FFFEC9A383B2FEDCF940" bold="true" box="[151,262,1726,1749]" pageId="6" pageNumber="117">Remarks.</emphasis>
|
||
This species has been described based on the specimens collected from only Mt. Hiko-san,
|
||
<collectingRegion id="255676E3FFF6FFFECBAE83D2FF14F880" country="Japan" name="Fukuoka" pageId="6" pageNumber="117">Fukuoka Pref.</collectingRegion>
|
||
, Kyushu (
|
||
<bibRefCitation id="8303C5F0FFF6FFFEC80383F2FE32F880" author="WATANABE Y." box="[311,488,1790,1813]" pageId="6" pageNumber="117" pagination="85 - 98" refId="ref8856" refString="WATANABE Y. 1998: Five new species of the Lathrobium (s. str.) nomurai group (Coleoptera, Staphylinidae) from Japan. Elytra 26 (1): 85 - 98." type="journal article" year="1998">WATANABE 1998</bibRefCitation>
|
||
). This study revealed that this species occurs from
|
||
<collectingRegion id="255676E3FFF6FFFEC8AA8212FDDCF8A0" box="[414,518,1822,1845]" country="Japan" name="Oita" pageId="6" pageNumber="117">Oita Pref.</collectingRegion>
|
||
to
|
||
<collectingRegion id="255676E3FFF6FFFECB138212FD1DF8A0" box="[551,711,1822,1845]" country="Japan" name="Nagasaki" pageId="6" pageNumber="117">Nagasaki Pref.</collectingRegion>
|
||
, and its distribution is wider than other species of the
|
||
<taxonomicName id="2092C382FFF6FFFECBB18232FD22F8C0" authorityName="Nakane" authorityYear="1955" box="[645,760,1854,1877]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="nomurai">
|
||
<emphasis id="D5E66413FFF6FFFECBB18232FD22F8C0" box="[645,760,1854,1877]" italics="true" pageId="6" pageNumber="117">L. nomurai</emphasis>
|
||
</taxonomicName>
|
||
group in Kyushu (
|
||
<figureCitation id="7FA9A484FFF6FFFEC8628252FE4CF8E0" box="[342,406,1886,1909]" captionStart="Fig" captionStartId="11.[151,182,1153,1172]" captionTargetBox="[254,1353,201,1126]" captionTargetId="figure-402@11.[207,1379,186,1137]" captionTargetPageId="11" captionText="Fig. 9. Map showing the distribution of the Lathrobium nomurai group in Kyushu, Japan. Black rhombus – L. nomurai Nakane, 1955; black triangle – L. fujimotoi Y. Watanabe, 2001; black circle – L. hikosanense Y. Watanabe, 1998; black square – L. yufuense sp. nov.; black pentagon – L. denchu sp. nov.; A–G – localities of L. hikosanense: A – Mt. Daisen; B – Mt. Hiko-san (type locality for L. hikosanense); C – Hiraodai; D – Mt. Buccho-zan; E – Mt. Tachibana-yama; F – Mt. Iwara-yama; G – Kinsenji Temple." figureDoi="http://doi.org/10.5281/zenodo.8006000" httpUri="https://zenodo.org/record/8006000/files/figure.png" pageId="6" pageNumber="117">Fig. 9</figureCitation>
|
||
). In addition, the aedeagus of this species shows a geographic variation in each region (
|
||
<figureCitation id="7FA9A484FFF6FFFECBF98272FF7DF820" captionStart="Fig" captionStartId="5.[150,181,2010,2029]" captionTargetBox="[289,1292,705,1970]" captionTargetId="figure-15@5.[261,1325,667,1990]" captionTargetPageId="5" captionText="Fig. 4. Male aedeagus of Lathrobium hikosanense Y. Watanabe, 1998 in each locality.A–C – Mt. Hiko-san (type locality): A – ventral view; B – lateral view; C – dorsal view. D, J – Mt. Daisen; E, K – Hiraodai; F, L – Mt. Buccho-zan; G, M – Mt. Tachibana-yama; H, N – Mt. Iwara-yama; I, O – Kinsenji Temple. Scale bars = 0.5 mm. Each locality referred to Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.8005990" httpUri="https://zenodo.org/record/8005990/files/figure.png" pageId="6" pageNumber="117">Fig. 4</figureCitation>
|
||
). We consider this variation as intraspecific because it is continuous, and the structures of internal sac are about the same in all examined specimens. See
|
||
<bibRefCitation id="8303C5F0FFF6FFFECB2E82D2FD0EF860" author="WATANABE Y." box="[538,724,2014,2037]" pageId="6" pageNumber="117" pagination="85 - 98" refId="ref8856" refString="WATANABE Y. 1998: Five new species of the Lathrobium (s. str.) nomurai group (Coleoptera, Staphylinidae) from Japan. Elytra 26 (1): 85 - 98." type="journal article" year="1998">WATANABE (1998)</bibRefCitation>
|
||
for a detailed description.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |