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<document id="62C888CE6B7A312823729D9AA1AAB4C3" ID-DOI="10.21248/contrib.entomol.65.1.173-196" ID-ISSN="0005-805X" ID-Zenodo-Dep="4753843" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620838100588" checkinUser="carolina" docAuthor="Irmler, Ulrich" docDate="2015" docId="6F2ED1232376FF8E9A8DFE7CA576C828" docLanguage="en" docName="ContribEntomol.65.1.173-196.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 65 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1876" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Tumboecus verhaaghi Irmler 2015, spec. nov." docType="treatment" docUuid="BE7E8724-7B2F-4AE9-AAAC-84EACAB2D89B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="180" masterDocId="9317A95B2371FF899A30FFEFA154CA09" masterDocTitle="New Neotropical genera and species of the tribe Osoriini (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="196" masterPageNumber="173" pageNumber="180" updateTime="1698916821617" updateUser="plazi">
<mods:mods id="93E62DC65BBF17B451200340F73D918D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AA6E962E2BBB95892271BB54C9CDD29C">
<mods:title id="374ADC169DDA5D111CDA97D89F6BBEF9">New Neotropical genera and species of the tribe Osoriini (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
</mods:titleInfo>
<mods:name id="697FAC9CBF834E2B7DDD0B888B5463BB" type="personal">
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</mods:role>
<mods:namePart id="32B21F79D2BCEF26BDE104AEE6324EC5">Irmler, Ulrich</mods:namePart>
</mods:name>
<mods:typeOfResource id="103A3DEDA7BEC9797EA9228DCE88B751">text</mods:typeOfResource>
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<mods:title id="90DF5169F899629CC2B8D4E62265E8BA">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="FB2F3338374E97506F2FC075B6428782">
<mods:date id="ABE00461F4F553256D4E3491BE6097C3">2015</mods:date>
<mods:detail id="D86A6DBC7C963EFDD0E43F7756C0956D" type="pubDate">
<mods:number id="2068F764B6759D0B8C98194A7D9BADA9">2015-06-30</mods:number>
</mods:detail>
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<mods:number id="A0A9E22EEC090EE31A75F95B72598C78">65</mods:number>
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<mods:number id="19DE44799D3ACDC4E3ACCF434B969D1C">1</mods:number>
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</mods:location>
<mods:classification id="F18910EF50E544C4F4621B5CEEB69F7A">journal article</mods:classification>
<mods:identifier id="D9E72A6BD77825DD7F143C33415AB17D" type="DOI">10.21248/contrib.entomol.65.1.173-196</mods:identifier>
<mods:identifier id="C59349FE55BB72DC134F3F96129148D8" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="64BF21C9D8D8E6DCEA7C1F1863BD42F5" type="Zenodo-Dep">4753843</mods:identifier>
<mods:identifier id="180F83A95249BAE934F9C5A53222C5AF" type="ZooBank">34012399-A256-429D-A5C0-58667E25EEEB</mods:identifier>
</mods:mods>
<treatment id="6F2ED1232376FF8E9A8DFE7CA576C828" ID-DOI="http://doi.org/10.5281/zenodo.4754120" ID-Zenodo-Dep="4754120" LSID="urn:lsid:zoobank.org:act:BE7E8724-7B2F-4AE9-AAAC-84EACAB2D89B" httpUri="http://treatment.plazi.org/id/6F2ED1232376FF8E9A8DFE7CA576C828" lastPageNumber="180" pageId="7" pageNumber="180">
<subSubSection id="AF9D33BE2376FF8E9A8DFE7CA35ACBA7" box="[189,526,403,430]" pageId="7" pageNumber="180" type="nomenclature">
<paragraph id="E73860352376FF8E9A8DFE7CA35ACBA7" blockId="7.[189,526,403,463]" box="[189,526,403,430]" pageId="7" pageNumber="180">
<heading id="BC70D7592376FF8E9A8DFE7CA35ACBA7" box="[189,526,403,430]" centered="true" fontSize="12" level="2" pageId="7" pageNumber="180" reason="2">
<taxonomicName id="20871BB62376FF8E9A8DFE7CA0CACBA7" ID-CoL="59LKT" authority="Irmler, 2015" authorityName="Irmler" authorityYear="2015" box="[189,414,403,430]" class="Insecta" family="Staphylinidae" genus="Tumboecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="180" phylum="Arthropoda" rank="species" species="verhaaghi" status="sp. nov.">
<emphasis id="D5F3BC272376FF8E9A8DFE7CA0CACBA7" box="[189,414,403,430]" italics="true" pageId="7" pageNumber="180">Tumboecus verhaaghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CEC0015C2376FF8E9B94FE7CA35ACBA7" box="[420,526,403,430]" pageId="7" pageNumber="180" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AF9D33BE2376FF8E9A8DFE56A01DCBC6" box="[189,329,441,463]" pageId="7" pageNumber="180" type="description">
<paragraph id="E73860352376FF8E9A8DFE56A01DCBC6" blockId="7.[189,526,403,463]" box="[189,329,441,463]" pageId="7" pageNumber="180">
(
<figureCitation id="7FBC7CB02376FF8E9AF1FE56A049CBC6" box="[193,285,441,463]" captionStart="Fig" captionStartId="17.[151,184,1874,1896]" captionTargetBox="[202,1376,427,1748]" captionTargetId="figure-41@17.[151,1398,400,1862]" captionTargetPageId="17" captionText="Fig. 5–9: Neorhapdopsis brasiliensis (5), N. pecki (6), N. lutzi (7), N. panamae (8), Tumboecus verhaaghi (9), antenna (A), protibia in posterior aspect (B), aedeagus (C); scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4753851" httpUri="https://zenodo.org/record/4753851/files/figure.png" pageId="7" pageNumber="180">Figs 9A–C</figureCitation>
,
<figureCitation id="7FBC7CB02376FF8E9B14FE56A010CBC6" box="[292,324,441,463]" captionStart="Fig" captionStartId="19.[151,184,2022,2044]" captionTargetBox="[184,1374,265,1981]" captionTargetId="figure-42@19.[151,1398,253,2011]" captionTargetPageId="19" captionText="Fig. 15: Head, pronotum, and elytra of Ouloglene barberi (A), Oryssma schwarzi (B), O. politum (C), O. exalatum (D), Tumboecus verhaaghi (E), and T. struyvei (F); scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4753859" httpUri="https://zenodo.org/record/4753859/files/figure.png" pageId="7" pageNumber="180">15E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="AF9D33BE2376FF8E9AA7FE06A02DC928" pageId="7" pageNumber="180" type="materials_examined">
<paragraph id="E73860352376FF8E9AA7FE06A03AC848" blockId="7.[151,757,489,801]" pageId="7" pageNumber="180">
<emphasis id="D5F3BC272376FF8E9AA7FE06A079C808" bold="true" box="[151,301,489,513]" pageId="7" pageNumber="180">Type material</emphasis>
:
<materialsCitation id="57EF6A682376FF8E9B09FE06A03AC848" collectingDate="1984-06-18" collectionCode="UIC" collectorName="M. Verhaagh" country="Peru" latitude="-9.616667" location="Panguana" longLatPrecision="1299" longitude="-74.933334" pageId="7" pageNumber="180" specimenCount="1" stateProvince="Huanuco" typeStatus="holotype">
<typeStatus id="383CDE972376FF8E9B09FE06A0C8C808" box="[313,412,489,513]" pageId="7" pageNumber="180" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="9F9020A52376FF8E9BD1FE05A341C808" box="[481,533,490,513]" name="Peru" pageId="7" pageNumber="180">Peru</collectingCountry>
,
<collectingRegion id="2543AED72376FF8E982EFE05A3D6C808" box="[542,642,490,513]" country="Peru" name="Huanuco" pageId="7" pageNumber="180">Huanuco</collectingRegion>
,
<location id="E25836EE2376FF8E98BBFE05A3A0C80B" LSID="urn:lsid:plazi:treatment:6F2ED1232376FF8E9A8DFE7CA576C828:E25836EE2376FF8E98BBFE05A3A0C80B" box="[651,756,490,514]" country="Peru" latitude="-9.616667" longLatPrecision="1299" longitude="-74.933334" name="Panguana" pageId="7" pageNumber="180" stateProvince="Huanuco">Panguana</location>
(
<geoCoordinate id="82B306F22376FF8E9AAEFDE6A1ADC828" box="[158,249,521,545]" degrees="74" direction="west" minutes="56" orientation="longitude" pageId="7" pageNumber="180" precision="925" value="-74.933334">74°56'W</geoCoordinate>
,
<geoCoordinate id="82B306F22376FF8E9B38FDE6A018C828" box="[264,332,521,545]" degrees="9" direction="south" minutes="37" orientation="latitude" pageId="7" pageNumber="180" precision="925" value="-9.616667">9°37'S</geoCoordinate>
),
<collectingMethod id="3EC618222376FF8E9B53FDE6A08DC828" box="[355,473,521,545]" pageId="7" pageNumber="180">pitfall trap</collectingMethod>
, male, 3.–
<date id="933946F52376FF8E9877FDE6A395C828" box="[583,705,521,545]" pageId="7" pageNumber="180" value="1984-06-18">
<collectingDate id="837DBF1D2376FF8E9877FDE6A395C828" box="[583,705,521,545]" pageId="7" pageNumber="180" value="1984-06-18">18.06.1984</collectingDate>
</date>
, leg.
<collectorName id="4A7205E32376FF8E9AA7FDC5A075C848" box="[151,289,553,577]" pageId="7" pageNumber="180">M. Verhaagh</collectorName>
(
<collectionCode id="8196F8F02376FF8E9B1FFDC6A036C848" box="[303,354,553,577]" pageId="7" pageNumber="180">UIC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="E73860352376FF8E9AA7FDA5A02DC928" blockId="7.[151,757,489,801]" pageId="7" pageNumber="180">
<materialsCitation id="57EF6A682376FF8E9AA7FDA5A0C6C8A8" collectingDate="1984-05-20" collectionCode="UIC" collectorName="M. Verhaagh" country="Peru" latitude="-9.616667" location="Panguana" longLatPrecision="1299" longitude="-74.933334" pageId="7" pageNumber="180" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Huanuco" typeStatus="paratype">
<typeStatus id="383CDE972376FF8E9AA7FDA5A055C86B" box="[151,257,586,610]" pageId="7" pageNumber="180" type="paratype">Paratypes</typeStatus>
: female with same data as holotype;
<collectingCountry id="9F9020A52376FF8E988CFDA5A3A4C868" box="[700,752,586,609]" name="Peru" pageId="7" pageNumber="180">Peru</collectingCountry>
,
<collectingRegion id="2543AED72376FF8E9AA7FD85A1AFC888" box="[151,251,618,641]" country="Peru" name="Huanuco" pageId="7" pageNumber="180">Huanuco</collectingRegion>
,
<location id="E25836EE2376FF8E9B38FD85A027C88B" LSID="urn:lsid:plazi:treatment:6F2ED1232376FF8E9A8DFE7CA576C828:E25836EE2376FF8E9B38FD85A027C88B" box="[264,371,618,642]" country="Peru" name="Panguana" pageId="7" pageNumber="180" stateProvince="Huanuco">Panguana</location>
, rainforest,
<specimenCount id="F181ABBC2376FF8E9BC6FD86A36BC888" box="[502,575,617,641]" pageId="7" pageNumber="180" type="male">1 male</specimenCount>
,
<specimenCount id="F181ABBC2376FF8E987CFD86A3FCC888" box="[588,680,617,641]" pageId="7" pageNumber="180" type="female">1 female</specimenCount>
,
<date id="933946F52376FF8E9885FD86A198C8A8" pageId="7" pageNumber="180" value="1984-05-20">
<collectingDate id="837DBF1D2376FF8E9885FD86A198C8A8" pageId="7" pageNumber="180" value="1984-05-20">20.05. 1984</collectingDate>
</date>
, leg.
<collectorName id="4A7205E32376FF8E9B37FD65A0C6C8A8" box="[263,402,649,673]" pageId="7" pageNumber="180">M. Verhaagh</collectorName>
</materialsCitation>
; rainforest,
<specimenCount id="F181ABBC2376FF8E9824FD66A32EC8A8" box="[532,634,649,673]" pageId="7" pageNumber="180" type="female">4 females</specimenCount>
,
<date id="933946F52376FF8E98B6FD66A3A4C8A8" box="[646,752,649,673]" pageId="7" pageNumber="180" value="1975-02">Feb. 1975</date>
,
<date id="933946F52376FF8E9AA7FD46A058C8C8" box="[151,268,681,705]" pageId="7" pageNumber="180" value="1975-06-05">06.05.1975</date>
, 08.–09.1975, 10.–11.1975 leg. W. Hanagarth; grassland, female,
<date id="933946F52376FF8E9B5DFD26A0B6C8E8" box="[365,482,713,737]" pageId="7" pageNumber="180" value="1975-03-09">03.09.1975</date>
, leg. W. Hanagarth;
<materialsCitation id="57EF6A682376FF8E98E3FD25A021C928" collectingDate="1975-03-20" collectionCode="UIC" collectorName="W. Hanagarth" country="Peru" pageId="7" pageNumber="180" specimenCount="1" stateProvince="Huanuco" typeStatus="paratype">
cut rainforest planted with manioc, female,
<date id="933946F52376FF8E987DFD06A396C908" box="[589,706,745,769]" pageId="7" pageNumber="180" value="1975-03-20">
<collectingDate id="837DBF1D2376FF8E987DFD06A396C908" box="[589,706,745,769]" pageId="7" pageNumber="180" value="1975-03-20">20.03.1975</collectingDate>
</date>
, leg.
<collectorName id="4A7205E32376FF8E9AA7FCE5A07FC928" box="[151,299,777,801]" pageId="7" pageNumber="180">W. Hanagarth</collectorName>
(
<collectionCode id="8196F8F02376FF8E9B09FCE6A038C928" box="[313,364,777,801]" pageId="7" pageNumber="180">UIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AF9D33BE2376FF8E9AA7FCA6A051C9A8" pageId="7" pageNumber="180" type="etymology">
<paragraph id="E73860352376FF8E9AA7FCA6A051C9A8" blockId="7.[151,756,841,929]" pageId="7" pageNumber="180">
<emphasis id="D5F3BC272376FF8E9AA7FCA6A05AC968" bold="true" box="[151,270,841,865]" pageId="7" pageNumber="180">Etymology</emphasis>
: The specific name is given in honor of the collector of the species to whom the species is heartily dedicated.
</paragraph>
</subSubSection>
<subSubSection id="AF9D33BE2376FF8E9AA7FC26A50CCB48" pageId="7" pageNumber="180" type="description">
<paragraph id="E73860352376FF8E9AA7FC26A0FBCE08" blockId="7.[151,757,969,2081]" pageId="7" pageNumber="180">
<emphasis id="D5F3BC272376FF8E9AA7FC26A04DC9E8" bold="true" box="[151,281,969,993]" pageId="7" pageNumber="180">Description</emphasis>
: Length:
<quantity id="207FCDD02376FF8E9BB6FC26A08EC9E8" box="[390,474,969,993]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="180" unit="mm" value="1.5">1.5 mm</quantity>
.
<emphasis id="D5F3BC272376FF8E9BD8FC26A331C9E8" box="[488,613,969,993]" italics="true" pageId="7" pageNumber="180">Colouration</emphasis>
: brown; legs and antennae light brown.
</paragraph>
<paragraph id="E73860352376FF8E9AA7FBE6A00CCEC8" blockId="7.[151,757,969,2081]" pageId="7" pageNumber="180">
Head:
<quantity id="207FCDD02376FF8E9AD5FBE6A011CE28" box="[229,325,1033,1057]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="7" pageNumber="180" unit="mm" value="0.18">0.18 mm</quantity>
long,
<quantity id="207FCDD02376FF8E9BA6FBE6A0A2CE28" box="[406,502,1033,1057]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="180" unit="mm" value="0.31">0.31 mm</quantity>
wide; eyes large, not prominent; longer than temples; sides of forehead strongly convergent; anterior edge of clypeus even; setiferous punctation dense and coarse; neck without punctation; weak microsculpture netlike; surface moderately shiny.
</paragraph>
<paragraph id="E73860352376FF8E9AA7FB26A1A8CF88" blockId="7.[151,757,969,2081]" pageId="7" pageNumber="180">Antennae nearly as long as head and pronotum combined; first antennomere thick; as thick as fourth antennomere; second narrower and oval; third antennomere conical; following antennomeres increasing in width; fourth to ninth wider than long; tenth antennomere approximately quadrate.</paragraph>
<paragraph id="E73860352376FF8E9AA7FA65A327CCA8" blockId="7.[151,757,969,2081]" pageId="7" pageNumber="180">
Pronotum:
<quantity id="207FCDD02376FF8E9B24FA66A026CFA8" box="[276,370,1417,1441]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="180" unit="mm" value="0.31">0.31 mm</quantity>
long, 0.36 wide; widest at anterior angles; moderately narrowed to posterior angles in anterior half; more strongly narrowed in posterior half; in front of posterior angles more strongly narrowed; lateral margin fine; in dorsal aspect covered in anterior half; setiferous punctation coarse and large; interstices between punctures irregularly wide; on average, half as wide as diameter of punctures; in lateral posterior half, interstices wider; without microsculpture; surface shiny.
</paragraph>
<paragraph id="E73860352376FF8E9AA7F946A07CCD68" blockId="7.[151,757,969,2081]" pageId="7" pageNumber="180">
Elytra:
<quantity id="207FCDD02376FF8E9AD4F946A017CCC8" box="[228,323,1705,1729]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="7" pageNumber="180" unit="mm" value="0.39">0.39 mm</quantity>
long,
<quantity id="207FCDD02376FF8E9BB6F946A0B0CCC8" box="[390,484,1705,1729]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="7" pageNumber="180" unit="mm" value="0.38">0.38 mm</quantity>
wide; shoulders approximately rectangular; sides subparallel; slightly widened posteriorly; setiferous punctation large and dense; on average, interstices as wide as or slightly shorter than diameter of punctures; with remains of microsculpture; surface shiny.
</paragraph>
<paragraph id="E73860352376FF8E9AA7F886A0BFCDA8" blockId="7.[151,757,969,2081]" pageId="7" pageNumber="180">Abdomen with similar setiferous punctation and similar microsculpture as on fore-body.</paragraph>
<paragraph id="E73860352376FF8E9AA7F846A0B0C228" blockId="7.[151,757,969,2081]" pageId="7" pageNumber="180">Protibia only slightly enlarged; with four spines on outer edge inserting directly on edge; in posterior aspect, comb at inner emargination totally visible; posterior face with several setae; tarsae pentamere.</paragraph>
<paragraph id="E73860352376FF8E992AFF06A50CCB48" blockId="7.[794,1399,233,321]" pageId="7" pageNumber="180">Aedeagus short and stout; without paramera; apical lobe in rectangular angle to basal lobe; apex moderately acute; endophallus with one torsion.</paragraph>
</subSubSection>
<subSubSection id="AF9D33BE2376FF8E992AFE86A576C828" pageId="7" pageNumber="180" type="diagnosis">
<paragraph id="E73860352376FF8E992AFE86A576C828" blockId="7.[794,1399,361,545]" pageId="7" pageNumber="180">
<emphasis id="D5F3BC272376FF8E992AFE86A2D3CB88" bold="true" box="[794,903,361,385]" pageId="7" pageNumber="180">Diagnosis</emphasis>
: The two species
<taxonomicName id="20871BB62376FF8E9E7FFE85A584CB88" authorityName="Irmler" authorityYear="2015" box="[1103,1232,361,385]" class="Insecta" family="Staphylinidae" genus="Tumboecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="180" phylum="Arthropoda" rank="species" species="verhaaghi">
<emphasis id="D5F3BC272376FF8E9E7FFE85A584CB88" box="[1103,1232,361,385]" italics="true" pageId="7" pageNumber="180">T. verhaaghi</emphasis>
</taxonomicName>
and
<taxonomicName id="20871BB62376FF8E9F3AFE85A422CB88" authorityName="Irmler" authorityYear="2015" box="[1290,1398,362,385]" class="Insecta" family="Staphylinidae" genus="Tumboecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="180" phylum="Arthropoda" rank="species" species="struyvei">
<emphasis id="D5F3BC272376FF8E9F3AFE85A422CB88" box="[1290,1398,362,385]" italics="true" pageId="7" pageNumber="180">T. struyvei</emphasis>
</taxonomicName>
are extremely similar regarding the punctation and shape of the pronotum.
<taxonomicName id="20871BB62376FF8E99D8FE45A53DCBC8" authorityName="Irmler" authorityYear="2015" box="[1000,1129,425,449]" class="Insecta" family="Staphylinidae" genus="Tumboecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="180" phylum="Arthropoda" rank="species" species="verhaaghi">
<emphasis id="D5F3BC272376FF8E99D8FE45A53DCBC8" box="[1000,1129,425,449]" italics="true" pageId="7" pageNumber="180">T. verhaaghi</emphasis>
</taxonomicName>
is slightly smaller than
<taxonomicName id="20871BB62376FF8E992AFE25A2D2CBE8" authorityName="Irmler" authorityYear="2015" box="[794,902,458,481]" class="Insecta" family="Staphylinidae" genus="Tumboecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="180" phylum="Arthropoda" rank="species" species="struyvei">
<emphasis id="D5F3BC272376FF8E992AFE25A2D2CBE8" box="[794,902,458,481]" italics="true" pageId="7" pageNumber="180">T. struyvei</emphasis>
</taxonomicName>
and the endophallus is different. Furthermore, the surface of
<taxonomicName id="20871BB62376FF8E998DFE05A56AC808" authorityName="Irmler" authorityYear="2015" box="[957,1086,489,513]" class="Insecta" family="Staphylinidae" genus="Tumboecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="180" phylum="Arthropoda" rank="species" species="verhaaghi">
<emphasis id="D5F3BC272376FF8E998DFE05A56AC808" box="[957,1086,489,513]" italics="true" pageId="7" pageNumber="180">T. verhaaghi</emphasis>
</taxonomicName>
is shinier, in particular, the surface of the pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>
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