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<document id="789E8482BCEDA89020EF7AB97110674D" ID-DOI="10.5281/zenodo.178993" ID-GBIF-Dataset="47088b53-41a2-4c58-b39e-2904ac4369a8" ID-ISSN="1175-5326" ID-Zenodo-Dep="178993" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459934776494" checkinUser="plazi" docAuthor="Pesic, Vladimir &amp; Smit, Harry" docDate="2007" docId="6D381B73FFD4F34935A20504FB97FD8C" docLanguage="en" docName="zt01613p056.pdf" docOrigin="Zootaxa 1613" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Torrenticola (Torrenticola) bhutana Pesic &amp; Smit, 2007, sp. nov." docType="treatment" docVersion="9" lastPageNumber="53" masterDocId="9101630BFFD1F34135350369FFBCFFE6" masterDocTitle="First records of water mites (Acari: Hydrachnidia) from Bhutan, with description of two new species" masterLastPageNumber="56" masterPageNumber="45" pageNumber="50" updateTime="1698230036454" updateUser="plazi">
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<mods:title id="3CD745C4355230FB923E2B570A88A160">First records of water mites (Acari: Hydrachnidia) from Bhutan, with description of two new species</mods:title>
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<mods:namePart id="7B10FC11ABAC101ED496CE12DD0EFD14">Pesic, Vladimir</mods:namePart>
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<mods:namePart id="EACF9240DD29D6C548620D852CCA7514">Smit, Harry</mods:namePart>
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<mods:typeOfResource id="180D1622897B40B6983BB84B80A218A9">text</mods:typeOfResource>
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<mods:date id="99D66A6FA389C380B183F4727F297E0D">2007</mods:date>
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<treatment id="6D381B73FFD4F34935A20504FB97FD8C" ID-DOI="http://doi.org/10.5281/zenodo.6242748" ID-GBIF-Taxon="119375067" ID-Zenodo-Dep="6242748" LSID="urn:lsid:plazi:treatment:6D381B73FFD4F34935A20504FB97FD8C" httpUri="http://treatment.plazi.org/id/6D381B73FFD4F34935A20504FB97FD8C" lastPageId="8" lastPageNumber="53" pageId="5" pageNumber="50">
<subSubSection id="AD8BF9EEFFD4F34435A20504FE88F94C" pageId="5" pageNumber="50" type="nomenclature">
<paragraph id="E52EAA65FFD4F34435A20504FD10F961" blockId="5.[151,684,1645,1706]" box="[151,684,1645,1671]" pageId="5" pageNumber="50">
<heading id="BE661D09FFD4F34435A20504FD10F961" bold="true" box="[151,684,1645,1671]" fontSize="11" level="1" pageId="5" pageNumber="50" reason="1">
<emphasis id="D7E57677FFD4F34435A20504FD10F961" bold="true" box="[151,684,1645,1671]" pageId="5" pageNumber="50">
<taxonomicName id="2291D1E6FFD4F34435A20504FDFBF961" box="[151,583,1645,1671]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="bhutana" status="sp. nov." subGenus="Torrenticola">
<emphasis id="D7E57677FFD4F34435A20504FDFBF961" bold="true" box="[151,583,1645,1671]" italics="true" pageId="5" pageNumber="50">Torrenticola (Torrenticola) bhutana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCD6CB0CFFD4F344377B0504FD10F961" box="[590,684,1645,1671]" pageId="5" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E52EAA65FFD4F34435A205F9FE88F94C" blockId="5.[151,684,1645,1706]" box="[151,308,1680,1706]" pageId="5" pageNumber="50">
(
<figureCitation id="7DAAB6E0FFD4F34435AA05F9FE90F94C" box="[159,300,1680,1706]" captionStart-0="FIGURES 21 24" captionStart-1="FIGURES 25 31. 25 30" captionStartId-0="6.[151,270,1490,1514]" captionStartId-1="7.[151,270,1895,1919]" captionTargetBox-0="[164,1381,489,1437]" captionTargetBox-1="[163,1415,202,1847]" captionTargetId-0="figure@6.[151,1436,474,1466]" captionTargetId-1="figure@7.[151,1436,194,1871]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 21 24. Torrenticola bhutana sp. nov., male: 21 = dorsal shield, 22 = ventral shield; 23 = palp, lateral view; 24 = palp, medial view. Scale bars = 100 m." captionText-1="FIGURES 25 31. 25 30 Torrenticola bhutana sp. nov. (25 26, 28 30 = female, 27 = male): 25 = dorsal shield, 26 = ventral shield; 28 = capitulum, 27 = ejaculatory complex, 29 = palp, lateral view; 30 = palp, medial view. 31 Torrenticola semisuta (Halík), male, India (Garhwal Himalaya): 31 = ejaculatory complex. Scale bars = 100 m." httpUri-0="https://zenodo.org/record/178996/files/figure.png" httpUri-1="https://zenodo.org/record/178997/files/figure.png" pageId="5" pageNumber="50">Figs. 2130</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="AD8BF9EEFFD4F34435A205B6FD54F8AC" pageId="5" pageNumber="50" type="materials_examined">
<paragraph id="E52EAA65FFD4F34435A205B6FD54F8AC" blockId="5.[151,1437,1759,2026]" pageId="5" pageNumber="50">
<emphasis id="D7E57677FFD4F34435A205B6FE93F91F" bold="true" box="[151,303,1759,1785]" pageId="5" pageNumber="50">
<typeStatus id="3A2A14C7FFD4F34435A205B6FF68F91F" box="[151,212,1759,1785]" pageId="5" pageNumber="50">Type</typeStatus>
series.
</emphasis>
<typeStatus id="3A2A14C7FFD4F34434080589FE13F91C" box="[317,431,1760,1786]" pageId="5" pageNumber="50" type="holotype">Holotype</typeStatus>
: male (
<collectionCode id="838032A0FFD4F344373A0589FDD0F91C" LSID="urn:lsid:biocol.org:col:34975" box="[527,620,1760,1786]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="5" pageNumber="50">ZMAN</collectionCode>
), dissected and slide mounted in Hoyer's fluid.
<collectingCountry id="9D86EAF5FFD4F34431FE0589FA98F91C" box="[1227,1316,1760,1786]" name="Bhutan" pageId="5" pageNumber="50">Bhutan</collectingCountry>
: Pele La,
<date id="912F8CA5FFD4F34435A20461FEA2F8C4" box="[151,286,1800,1826]" pageId="5" pageNumber="50" value="2002-01-05">01.05.2002</date>
,
<quantity id="22690780FFD4F344341E0461FE3AF8C4" box="[299,390,1800,1826]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="50" unit="m" value="3200.0">3200 m</quantity>
asl.
<typeStatus id="3A2A14C7FFD4F344348A0461FD8DF8C4" box="[447,561,1800,1826]" pageId="5" pageNumber="50" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="838032A0FFD4F34437770461FD21F8C4" LSID="urn:lsid:biocol.org:col:34975" box="[578,669,1800,1826]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="5" pageNumber="50">ZMAN</collectionCode>
,
<collectionCode id="838032A0FFD4F344379E0461FCBAF8C4" box="[683,774,1800,1826]" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" pageId="5" pageNumber="50">MNHP</collectionCode>
): four females, same data and place as
<typeStatus id="3A2A14C7FFD4F34431EF0461FAF8F8C4" box="[1242,1348,1800,1826]" pageId="5" pageNumber="50" type="holotype">holotype</typeStatus>
, one of them dissected and slide mounted in Hoyer's fluid.
</paragraph>
</subSubSection>
<subSubSection id="AD8BF9EEFFD4F34435F2043EFC2BF87C" pageId="5" pageNumber="50" type="diagnosis">
<paragraph id="E52EAA65FFD4F34435F2043EFC2BF87C" blockId="5.[151,1437,1759,2026]" pageId="5" pageNumber="50">
<emphasis id="D7E57677FFD4F34435F2043EFEFBF897" bold="true" box="[199,327,1879,1905]" pageId="5" pageNumber="50">Diagnosis.</emphasis>
Cx-4 posteriorly extended beyond genital field in both sexes; P-2 long and slender, ventral projection on P-2 and P-3 blunt, one (of 4) P-4 ventral setae long.
</paragraph>
</subSubSection>
<subSubSection id="AD8BF9EEFFD4F34735F204CEFD8AF897" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50" type="description">
<paragraph id="E52EAA65FFD4F34735F204CEFB8AFE44" blockId="5.[151,1437,1759,2026]" lastBlockId="6.[151,1436,152,418]" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
<emphasis id="D7E57677FFD4F34435F204CEFEE3F827" bold="true" box="[199,351,1959,1985]" pageId="5" pageNumber="50">Description.</emphasis>
<emphasis id="D7E57677FFD4F344345304C1FE1EF827" box="[358,418,1960,1985]" italics="true" pageId="5" pageNumber="50">Male</emphasis>
: Idiosoma (ventral view:
<figureCitation id="7DAAB6E0FFD4F34437FA04C1FC9FF824" box="[719,803,1960,1986]" captionStart="FIGURES 21 24" captionStartId="6.[151,270,1490,1514]" captionTargetBox="[164,1381,489,1437]" captionTargetId="figure@6.[151,1436,474,1466]" captionTargetPageId="6" captionText="FIGURES 21 24. Torrenticola bhutana sp. nov., male: 21 = dorsal shield, 22 = ventral shield; 23 = palp, lateral view; 24 = palp, medial view. Scale bars = 100 m." httpUri="https://zenodo.org/record/178996/files/figure.png" pageId="5" pageNumber="50">Fig. 22</figureCitation>
) L 875, W 625; dorsal shield (
<figureCitation id="7DAAB6E0FFD4F34431BB04C1FB5EF824" box="[1166,1250,1960,1986]" captionStart="FIGURES 21 24" captionStartId="6.[151,270,1490,1514]" captionTargetBox="[164,1381,489,1437]" captionTargetId="figure@6.[151,1436,474,1466]" captionTargetPageId="6" captionText="FIGURES 21 24. Torrenticola bhutana sp. nov., male: 21 = dorsal shield, 22 = ventral shield; 23 = palp, lateral view; 24 = palp, medial view. Scale bars = 100 m." httpUri="https://zenodo.org/record/178996/files/figure.png" pageId="5" pageNumber="50">Fig. 21</figureCitation>
) L 706, W 531, L/W ratio; dorsal plate 656; shoulder plate L 209, W 81, L/W ratio 2.6; frontal plate L 153, W 63, L/W ratio 2.4; shoulder/frontal plate L ratio 1.37; gnathosomal bay L 157, Cx-1 total L 344, Cx-1 medial L 187, Cx-2+3 medial 61; ratio Cx-1 L/Cx-2+3 medial L 1.84; Cx-1 medial L/Cx-2+3 medial L 5.6; genital field L/W 172/ 116, L/W ratio 1.48, ejaculatory complex (
<figureCitation id="7DAAB6E0FFD7F34737A00381FD50FEE4" box="[661,748,232,258]" captionStart="FIGURES 25 31. 25 30" captionStartId="7.[151,270,1895,1919]" captionTargetBox="[163,1415,202,1847]" captionTargetId="figure@7.[151,1436,194,1871]" captionTargetPageId="7" captionText="FIGURES 25 31. 25 30 Torrenticola bhutana sp. nov. (25 26, 28 30 = female, 27 = male): 25 = dorsal shield, 26 = ventral shield; 28 = capitulum, 27 = ejaculatory complex, 29 = palp, lateral view; 30 = palp, medial view. 31 Torrenticola semisuta (Halík), male, India (Garhwal Himalaya): 31 = ejaculatory complex. Scale bars = 100 m." httpUri="https://zenodo.org/record/178997/files/figure.png" pageId="6" pageNumber="51">Fig. 27</figureCitation>
) L 253; distance genital fieldexcretory pore 188, genital fieldcaudal idiosoma margin 288; capitulum ventral L 369; chelicera L 327; palp (
<figureCitation id="7DAAB6E0FFD7F34731410279FABEFECC" box="[1140,1282,272,298]" captionStart="FIGURES 21 24" captionStartId="6.[151,270,1490,1514]" captionTargetBox="[164,1381,489,1437]" captionTargetId="figure@6.[151,1436,474,1466]" captionTargetPageId="6" captionText="FIGURES 21 24. Torrenticola bhutana sp. nov., male: 21 = dorsal shield, 22 = ventral shield; 23 = palp, lateral view; 24 = palp, medial view. Scale bars = 100 m." httpUri="https://zenodo.org/record/178996/files/figure.png" pageId="6" pageNumber="51">Figs. 2324</figureCitation>
) total L 287, dL and %L (in parentheses, given as % of total L): P-1 29 (10.1), P-2 98 (34.1), P-3 60 (20.9), P-4 85 (29.6), P-5 15 (5.2); P-2/P-4 ratio 1.15; ventral margin of P-2 slightly convex, ventral projections of P-2 and P-3 blunt, ventral protuberances of P-4 ending in two tips separated by a concavity.
</paragraph>
<caption id="B1EEFAEDFFD7F34735A206BBFDD7F9ED" httpUri="https://zenodo.org/record/178996/files/figure.png" pageId="6" pageNumber="51" targetBox="[164,1381,489,1437]" targetPageId="6">
<paragraph id="E52EAA65FFD7F34735A206BBFDD7F9ED" blockId="6.[151,1435,1490,1547]" pageId="6" pageNumber="51">
<emphasis id="D7E57677FFD7F34735A206BBFED0FA0C" bold="true" box="[151,364,1490,1514]" pageId="6" pageNumber="51">FIGURES 21 24.</emphasis>
<taxonomicName id="2291D1E6FFD7F347344706BBFDE8FA0F" box="[370,596,1490,1513]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="bhutana" status="sp. nov.">
<emphasis id="D7E57677FFD7F347344706BBFDE8FA0F" box="[370,596,1490,1513]" italics="true" pageId="6" pageNumber="51">Torrenticola bhutana</emphasis>
</taxonomicName>
<emphasis id="D7E57677FFD7F347376E06BBFD0DFA0C" bold="true" box="[603,689,1490,1514]" pageId="6" pageNumber="51">
<taxonomicNameLabel id="CCD6CB0CFFD7F347376E06BBFD0DFA0C" box="[603,689,1490,1514]" pageId="6" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male: 21 = dorsal shield, 22 = ventral shield; 23 = palp, lateral view; 24 = palp, medial view. Scale bars = 100 m.
</paragraph>
</caption>
<paragraph id="E52EAA65FFD7F34735F20529FD8AF897" blockId="6.[151,1436,1599,2025]" pageId="6" pageNumber="51">
<emphasis id="D7E57677FFD7F34735F20529FEA3F9BF" box="[199,287,1600,1625]" italics="true" pageId="6" pageNumber="51">Female</emphasis>
: Idiosoma (ventral view:
<figureCitation id="7DAAB6E0FFD7F34737650556FD19F9BF" box="[592,677,1599,1625]" captionStart="FIGURES 25 31. 25 30" captionStartId="7.[151,270,1895,1919]" captionTargetBox="[163,1415,202,1847]" captionTargetId="figure@7.[151,1436,194,1871]" captionTargetPageId="7" captionText="FIGURES 25 31. 25 30 Torrenticola bhutana sp. nov. (25 26, 28 30 = female, 27 = male): 25 = dorsal shield, 26 = ventral shield; 28 = capitulum, 27 = ejaculatory complex, 29 = palp, lateral view; 30 = palp, medial view. 31 Torrenticola semisuta (Halík), male, India (Garhwal Himalaya): 31 = ejaculatory complex. Scale bars = 100 m." httpUri="https://zenodo.org/record/178997/files/figure.png" pageId="6" pageNumber="51">Fig. 26</figureCitation>
) L 938, W 719, dorsal shield (
<figureCitation id="7DAAB6E0FFD7F347313B0556FBD8F9BF" box="[1038,1124,1599,1625]" captionStart="FIGURES 25 31. 25 30" captionStartId="7.[151,270,1895,1919]" captionTargetBox="[163,1415,202,1847]" captionTargetId="figure@7.[151,1436,194,1871]" captionTargetPageId="7" captionText="FIGURES 25 31. 25 30 Torrenticola bhutana sp. nov. (25 26, 28 30 = female, 27 = male): 25 = dorsal shield, 26 = ventral shield; 28 = capitulum, 27 = ejaculatory complex, 29 = palp, lateral view; 30 = palp, medial view. 31 Torrenticola semisuta (Halík), male, India (Garhwal Himalaya): 31 = ejaculatory complex. Scale bars = 100 m." httpUri="https://zenodo.org/record/178997/files/figure.png" pageId="6" pageNumber="51">Fig. 25</figureCitation>
) L 756, W 588, L/W ratio 1.29; dorsal plate 700; shoulder plate L 268, W 81, L/W ratio 3.3; frontal plate L 166, W 78, L/W ratio 2.1; shoulder/frontal plate L ratio 1.6; gnathosomal bay L 181, Cx-1 total L 356, Cx-1 medial L 175, Cx-2+3 medial 41; ratio Cx-1 L/Cx-2+3 medial L 2.0; Cx-1 medial L/Cx-2+3 medial L 8.7; genital field L/W 197/ 167, L/W ratio 1.18; distance genital fieldexcretory pore 194, genital fieldcaudal idiosoma margin 325; capitulum (
<figureCitation id="7DAAB6E0FFD7F3473429046EFECCF8C7" box="[284,368,1799,1825]" captionStart="FIGURES 25 31. 25 30" captionStartId="7.[151,270,1895,1919]" captionTargetBox="[163,1415,202,1847]" captionTargetId="figure@7.[151,1436,194,1871]" captionTargetPageId="7" captionText="FIGURES 25 31. 25 30 Torrenticola bhutana sp. nov. (25 26, 28 30 = female, 27 = male): 25 = dorsal shield, 26 = ventral shield; 28 = capitulum, 27 = ejaculatory complex, 29 = palp, lateral view; 30 = palp, medial view. 31 Torrenticola semisuta (Halík), male, India (Garhwal Himalaya): 31 = ejaculatory complex. Scale bars = 100 m." httpUri="https://zenodo.org/record/178997/files/figure.png" pageId="6" pageNumber="51">Fig. 28</figureCitation>
) ventral L 381; chelicera L 438; palp (
<figureCitation id="7DAAB6E0FFD7F3473607046EFC03F8C7" box="[818,959,1799,1825]" captionStart="FIGURES 25 31. 25 30" captionStartId="7.[151,270,1895,1919]" captionTargetBox="[163,1415,202,1847]" captionTargetId="figure@7.[151,1436,194,1871]" captionTargetPageId="7" captionText="FIGURES 25 31. 25 30 Torrenticola bhutana sp. nov. (25 26, 28 30 = female, 27 = male): 25 = dorsal shield, 26 = ventral shield; 28 = capitulum, 27 = ejaculatory complex, 29 = palp, lateral view; 30 = palp, medial view. 31 Torrenticola semisuta (Halík), male, India (Garhwal Himalaya): 31 = ejaculatory complex. Scale bars = 100 m." httpUri="https://zenodo.org/record/178997/files/figure.png" pageId="6" pageNumber="51">Figs. 2930</figureCitation>
) total L 306, dL and %L (in parentheses, given as % of total L): P-1 33 (10.8), P-2 108 (35.3), P-3 62 (20.3), P-4 89 (29.1), P-5 14 (4.6); P-2/P-4 ratio 1.21; shape and setation as in male.
</paragraph>
</subSubSection>
<subSubSection id="AD8BF9EEFFD7F34935F20416FAC6FE14" lastPageId="8" lastPageNumber="53" pageId="6" pageNumber="51" type="discussion">
<paragraph id="E52EAA65FFD7F34935F20416FD45FE9C" blockId="6.[151,1436,1599,2025]" lastBlockId="8.[151,1437,152,618]" lastPageId="8" lastPageNumber="53" pageId="6" pageNumber="51">
<emphasis id="D7E57677FFD7F34735F20416FEEEF87F" bold="true" box="[199,338,1919,1945]" pageId="6" pageNumber="51">Discussion.</emphasis>
Due to sub-apical Cx-4, one (of 4) P-4 ventral setae long, a long rostrum, a short median suture line of Cx-2+3 and Cx-4 extended posterior to the genital flaps in both sexes,
<taxonomicName id="2291D1E6FFD7F347319A04C1FA93F827" box="[1199,1327,1960,1985]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="bhutana" status="sp. nov.">
<emphasis id="D7E57677FFD7F347319A04C1FA93F827" box="[1199,1327,1960,1985]" italics="true" pageId="6" pageNumber="51">T. bhutana</emphasis>
</taxonomicName>
<emphasis id="D7E57677FFD7F347300E04CEFA27F827" bold="true" box="[1339,1435,1959,1985]" pageId="6" pageNumber="51">
<taxonomicNameLabel id="CCD6CB0CFFD7F347300E04CEFA27F827" box="[1339,1435,1959,1985]" pageId="6" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="2291D1E6FFD7F347342604B9FDD6F80F" authority="Halik" authorityName="Halik" box="[275,618,1999,2025]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="semisuta">
<emphasis id="D7E57677FFD7F347342604B9FDAEF80F" box="[275,530,2000,2025]" italics="true" pageId="6" pageNumber="51">Torrenticola semisuta</emphasis>
(Halík)
</taxonomicName>
.
<taxonomicName id="2291D1E6FFD7F347374304B9FD4FF80F" box="[630,755,2000,2025]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="bhutana" status="sp. nov.">
<emphasis id="D7E57677FFD7F347374304B9FD4FF80F" box="[630,755,2000,2025]" italics="true" pageId="6" pageNumber="51">T. bhutana</emphasis>
</taxonomicName>
<emphasis id="D7E57677FFD7F34737CF04A6FCE4F80F" bold="true" box="[762,856,1999,2025]" pageId="6" pageNumber="51">
<taxonomicNameLabel id="CCD6CB0CFFD7F34737CF04A6FCE4F80F" box="[762,856,1999,2025]" pageId="6" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="2291D1E6FFD7F34731A304B9FAA6F80F" box="[1174,1306,2000,2025]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="semisuta">
<emphasis id="D7E57677FFD7F34731A304B9FAA6F80F" box="[1174,1306,2000,2025]" italics="true" pageId="6" pageNumber="51">T. semisuta</emphasis>
</taxonomicName>
by the shape of the ejaculatory complex (conventional in shape with well-developed proximal horns in
<taxonomicName id="2291D1E6FFD9F349303203F1FA34FF57" box="[1287,1416,152,177]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="semisuta">
<emphasis id="D7E57677FFD9F349303203F1FA34FF57" box="[1287,1416,152,177]" italics="true" pageId="8" pageNumber="53">T. semisuta</emphasis>
</taxonomicName>
- compare
<figureCitation id="7DAAB6E0FFD9F349343003A9FEE5FF3C" box="[261,345,192,218]" captionStart="FIGURES 25 31. 25 30" captionStartId="7.[151,270,1895,1919]" captionTargetBox="[163,1415,202,1847]" captionTargetId="figure@7.[151,1436,194,1871]" captionTargetPageId="7" captionText="FIGURES 25 31. 25 30 Torrenticola bhutana sp. nov. (25 26, 28 30 = female, 27 = male): 25 = dorsal shield, 26 = ventral shield; 28 = capitulum, 27 = ejaculatory complex, 29 = palp, lateral view; 30 = palp, medial view. 31 Torrenticola semisuta (Halík), male, India (Garhwal Himalaya): 31 = ejaculatory complex. Scale bars = 100 m." httpUri="https://zenodo.org/record/178997/files/figure.png" pageId="8" pageNumber="53">Fig. 27</figureCitation>
with
<figureCitation id="7DAAB6E0FFD9F349349503A9FE4BFF3C" box="[416,503,192,218]" captionStart="FIGURES 25 31. 25 30" captionStartId="7.[151,270,1895,1919]" captionTargetBox="[163,1415,202,1847]" captionTargetId="figure@7.[151,1436,194,1871]" captionTargetPageId="7" captionText="FIGURES 25 31. 25 30 Torrenticola bhutana sp. nov. (25 26, 28 30 = female, 27 = male): 25 = dorsal shield, 26 = ventral shield; 28 = capitulum, 27 = ejaculatory complex, 29 = palp, lateral view; 30 = palp, medial view. 31 Torrenticola semisuta (Halík), male, India (Garhwal Himalaya): 31 = ejaculatory complex. Scale bars = 100 m." httpUri="https://zenodo.org/record/178997/files/figure.png" pageId="8" pageNumber="53">Fig. 31</figureCitation>
), and the blunt P-2 ventral projection not strongly curved distally (P-2 ventral extension strongly curved distally in
<taxonomicName id="2291D1E6FFD9F34937720381FD76FEE7" box="[583,714,232,257]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="semisuta">
<emphasis id="D7E57677FFD9F34937720381FD76FEE7" box="[583,714,232,257]" italics="true" pageId="8" pageNumber="53">T. semisuta</emphasis>
</taxonomicName>
). According to
<bibRefCitation id="8100D794FFD9F34936B70381FBA7FEE4" author="Wiles" box="[898,1051,232,258]" pageId="8" pageNumber="56" refString="Wiles, P. R. (1997) Asian and Oriental Torrenticolidae Piersig, 1902 (Acari: Hydrachnidia: Lebertioidea): a revision of the family and description of new species of Torrenticola Piersig and Pseudotorrenticola Walter, from Southeast Asia. Journal of Natural History, 31, 191 - 236." type="journal article" year="1997">Wiles (1997)</bibRefCitation>
the projections of P-2 are probably variable in shape and therefore the blunt projections may not be a good character for separating species. Further differences in
<taxonomicName id="2291D1E6FFD9F349349F0251FD8FFEB7" box="[426,563,312,337]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="semisuta">
<emphasis id="D7E57677FFD9F349349F0251FD8FFEB7" box="[426,563,312,337]" italics="true" pageId="8" pageNumber="53">T. semisuta</emphasis>
</taxonomicName>
regard the shortened postgenital area in male and Cx-4 more strongly extended posterior to the genital flaps in the female.
</paragraph>
<caption id="B1EEFAEDFFD6F34635A2040EFB13F825" httpUri="https://zenodo.org/record/178997/files/figure.png" pageId="7" pageNumber="52" targetBox="[163,1415,202,1847]" targetPageId="7">
<paragraph id="E52EAA65FFD6F34635A2040EFB13F825" blockId="7.[151,1436,1895,1988]" pageId="7" pageNumber="52">
<emphasis id="D7E57677FFD6F34635A2040EFE7EF899" bold="true" box="[151,450,1895,1919]" pageId="7" pageNumber="52">FIGURES 25 31. 25 30</emphasis>
<taxonomicName id="2291D1E6FFD6F34634FC040EFD17F898" box="[457,683,1895,1918]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="bhutana" status="sp. nov.">
<emphasis id="D7E57677FFD6F34634FC040EFD17F898" box="[457,683,1895,1918]" italics="true" pageId="7" pageNumber="52">Torrenticola bhutana</emphasis>
</taxonomicName>
<emphasis id="D7E57677FFD6F3463787040EFCBBF899" bold="true" box="[690,775,1895,1919]" pageId="7" pageNumber="52">
<taxonomicNameLabel id="CCD6CB0CFFD6F3463787040EFCBBF899" box="[690,775,1895,1919]" pageId="7" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(2526, 2830 = female, 27 = male): 25 = dorsal shield, 26 = ventral shield; 28 = capitulum, 27 = ejaculatory complex, 29 = palp, lateral view; 30 = palp, medial view.
<emphasis id="D7E57677FFD6F34631CD04E0FAAFF847" bold="true" box="[1272,1299,1929,1953]" pageId="7" pageNumber="52">31</emphasis>
<taxonomicName id="2291D1E6FFD6F346302C04E3FEF7F825" authority="Halik" authorityName="Halik" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="semisuta">
<emphasis id="D7E57677FFD6F346302C04E3FA20F847" box="[1305,1436,1930,1953]" italics="true" pageId="7" pageNumber="52">Torrenticola</emphasis>
<emphasis id="D7E57677FFD6F34635A204C4FF48F822" box="[151,244,1965,1988]" italics="true" pageId="7" pageNumber="52">semisuta</emphasis>
(Halík)
</taxonomicName>
, male, India (Garhwal Himalaya): 31 = ejaculatory complex. Scale bars = 100 m.
</paragraph>
</caption>
<paragraph id="E52EAA65FFD9F34935F202E1FAC6FE14" blockId="8.[151,1437,152,618]" pageId="8" pageNumber="53">
From
<taxonomicName id="2291D1E6FFD9F349343802E1FDE0FE44" authority="Lundblad" authorityName="Lundblad" box="[269,604,392,418]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="dolichorhyncha">
<emphasis id="D7E57677FFD9F349343802E1FE5DFE47" box="[269,481,392,417]" italics="true" pageId="8" pageNumber="53">T. dolichorhyncha</emphasis>
Lundblad
</taxonomicName>
, a species similar in the shape of the palp (P-2 long and slender, ventral projections on P-2 and P-3 blunt),
<taxonomicName id="2291D1E6FFD9F349370C02D9FD04FE2F" box="[569,696,432,457]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="bhutana" status="sp. nov.">
<emphasis id="D7E57677FFD9F349370C02D9FD04FE2F" box="[569,696,432,457]" italics="true" pageId="8" pageNumber="53">T. bhutana</emphasis>
</taxonomicName>
<emphasis id="D7E57677FFD9F34937F702C6FC9EFE2F" bold="true" box="[706,802,431,457]" pageId="8" pageNumber="53">
<taxonomicNameLabel id="CCD6CB0CFFD9F34937F702C6FC9EFE2F" box="[706,802,431,457]" pageId="8" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by the shape of the ejaculatory complex (conventional in shape see
<bibRefCitation id="8100D794FFD9F349376302B1FCB6FE14" author="Lundblad" box="[598,778,472,498]" pageId="8" pageNumber="55" refString="Lundblad, O. (1969) Indische Wassermilben, hauptschlich aus Hinterindien. Arkiv for Zoology, 22, 1 - 126." type="journal article" year="1969">Lundblad 1969</bibRefCitation>
) and Cx-4 are extended posterior to the genital flaps.
</paragraph>
</subSubSection>
<subSubSection id="AD8BF9EEFFD9F34935F20296FBBBFDFC" box="[199,1031,511,538]" pageId="8" pageNumber="53" type="etymology">
<paragraph id="E52EAA65FFD9F34935F20296FBBBFDFC" blockId="8.[151,1437,152,618]" box="[199,1031,511,538]" pageId="8" pageNumber="53">
<emphasis id="D7E57677FFD9F34935F20296FEE8FDFF" bold="true" box="[199,340,511,537]" pageId="8" pageNumber="53">Etymology.</emphasis>
The species is named after country where it was collected.
</paragraph>
</subSubSection>
<subSubSection id="AD8BF9EEFFD9F34935F2014EFC3BFDA4" box="[199,903,551,578]" pageId="8" pageNumber="53" type="distribution">
<paragraph id="E52EAA65FFD9F34935F2014EFC3BFDA4" blockId="8.[151,1437,152,618]" box="[199,903,551,578]" pageId="8" pageNumber="53">
<emphasis id="D7E57677FFD9F34935F2014EFEDBFDA7" bold="true" box="[199,359,551,577]" pageId="8" pageNumber="53">Distribution.</emphasis>
Known only from the
<typeStatus id="3A2A14C7FFD9F34937460141FD1AFDA4" box="[627,678,552,578]" pageId="8" pageNumber="53">type</typeStatus>
locality in
<collectingCountry id="9D86EAF5FFD9F349361F0141FC3FFDA4" box="[810,899,552,578]" name="Bhutan" pageId="8" pageNumber="53">Bhutan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="AD8BF9EEFFD9F34935F20126FB97FD8C" box="[199,1067,591,618]" pageId="8" pageNumber="53" type="biology_ecology">
<paragraph id="E52EAA65FFD9F34935F20126FB97FD8C" blockId="8.[151,1437,152,618]" box="[199,1067,591,618]" pageId="8" pageNumber="53">
<emphasis id="D7E57677FFD9F34935F20126FE97FD8F" bold="true" box="[199,299,591,617]" pageId="8" pageNumber="53">Biology.</emphasis>
<taxonomicName id="2291D1E6FFD9F34934070139FD96FD8F" box="[306,554,592,617]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="bhutana" status="sp. nov.">
<emphasis id="D7E57677FFD9F34934070139FD96FD8F" box="[306,554,592,617]" italics="true" pageId="8" pageNumber="53">Torrenticola bhutana</emphasis>
</taxonomicName>
<emphasis id="D7E57677FFD9F34937040126FD33FD8F" bold="true" box="[561,655,591,617]" pageId="8" pageNumber="53">
<taxonomicNameLabel id="CCD6CB0CFFD9F34937040126FD33FD8F" box="[561,655,591,617]" pageId="8" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is probably a rhitrobiontic species.
</paragraph>
</subSubSection>
</treatment>
</document>