125 lines
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125 lines
17 KiB
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<document id="21B9D0C803C13A9E568551CB3CA56396" ID-DOI="10.26879/702" ID-ISSN="1094-8074" ID-Zenodo-Dep="11063310" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1714014708037" checkinUser="felipe" docAuthor="Poole, Karen E." docDate="2016" docId="6D2D8786A2765F0D7450FE6526544AA3" docLanguage="en" docName="PalaeontolElectron.25.3.a30.pdf" docOrigin="Palaeontologia Electronica (a 30) 25 (3)" docSource="http://dx.doi.org/10.26879/702" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Rhabdodontidae Weishampel, Jianu, Csiki, and Norman 2003" docType="treatment" docVersion="4" lastPageNumber="18" masterDocId="9114FFFEA2675F1C753EFFB622124921" masterDocTitle="Phylogeny of iguanodontian dinosaurs and the evolution of quadrupedality" masterLastPageNumber="65" masterPageNumber="1" pageNumber="18" updateTime="1714997633796" updateUser="julia" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="8024261EAC7D1544E48F23BFD56AB720">Phylogeny of iguanodontian dinosaurs and the evolution of quadrupedality</mods:title>
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<mods:namePart id="7E5224F8C860B17F69725A5E411BA728">Poole, Karen E.</mods:namePart>
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<mods:title id="C9F7BA6A3A9023E2728CBD5AB55FFC1D">Palaeontologia Electronica</mods:title>
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<mods:date id="9920AF7DDE6711DA6C13A5EB5B5F88A6">2016</mods:date>
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<mods:title id="531824CE4E32E33B4216DBD8AB01DCAD">a 30</mods:title>
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<mods:number id="FB1D3E3AE317E5ABEB787FEC6A03464B">25</mods:number>
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<mods:identifier id="D21E04AF4DF5A303653F98FFF1586A95" type="DOI">10.26879/702</mods:identifier>
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<subSubSection id="AD9E651BA2765F0D7450FE6520E44B2D" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="E53B3690A2765F0D7450FE65207848CC" blockId="17.[226,758,467,525]" box="[366,618,467,493]" pageId="17" pageNumber="18">
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<heading id="BE7381FCA2765F0D7450FE65207848CC" box="[366,618,467,493]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="18" reason="8">
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<taxonomicName id="22844D13A2765F0D7450FE65207848CC" authority="Weishampel, Jianu, Csiki, and Norman, 2003" authorityName="Weishampel, Jianu, Csiki, and Norman" authorityYear="2003" box="[366,618,467,493]" family="Rhabdodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Chordata" rank="family">RHABDODONTIDAE</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="E53B3690A2765F0D75DCFE4520E44B2D" blockId="17.[226,758,467,525]" box="[226,758,499,525]" pageId="17" pageNumber="18">
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<heading id="BE7381FCA2765F0D75DCFE4520E44B2D" box="[226,758,499,525]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="18" reason="8">
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<bibRefCitation id="81154B61A2765F0D75DCFE4520E44B2D" author="Weishampel, D. B. & Jianu, C. - M. & Csiki, Z. & Norman, D. B." box="[226,758,499,525]" journalOrPublisher="Journal of Systematic Palaeontology" pageId="17" pageNumber="18" pagination="65 - 123" part="1" refId="ref23503" refString="Weishampel, D. B., Jianu, C. - M., Csiki, Z., and Norman, D. B. 2003. Osteology and phylogeny of Zalmoxes (n. g.), an unusual euornithopod dinosaur from the Latest Cretaceous of Romania. Journal of Systematic Palaeontology, 1 (2): 65 - 123. https: // doi. org / 10.1017 / s 1477201903001032" title="Osteology and phylogeny of Zalmoxes (n. g.), an unusual euornithopod dinosaur from the Latest Cretaceous of Romania" type="journal article" year="2003">Weishampel, Jianu, Csiki, and Norman, 2003</bibRefCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="AD9E651BA2765F0D75FEFDA826544AA3" pageId="17" pageNumber="18" type="description">
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<paragraph id="E53B3690A2765F0D75FEFDA823E54B96" blockId="17.[192,793,542,1911]" pageId="17" pageNumber="18">
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<emphasis id="D7F0EA82A2765F0D75FEFDA823E84B16" bold="true" box="[192,506,542,567]" pageId="17" pageNumber="18">Phylogenetic definition.</emphasis>
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A node-based taxon consisting of the most recent common ancestor of
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<taxonomicName id="22844D13A2765F0D75FEFDE823B34B56" baseAuthorityName="Nopcsa" baseAuthorityYear="1902" box="[192,417,606,631]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="robustus">
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<emphasis id="D7F0EA82A2765F0D75FEFDE823B34B56" box="[192,417,606,631]" italics="true" pageId="17" pageNumber="18">Zalmoxes robustus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="22844D13A2765F0D74E2FDE820D04B56" authorityName="Matheron" authorityYear="1869" box="[476,706,606,631]" class="Squamata" family="Psammophiidae" genus="Rhabdodon" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="priscus">
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<emphasis id="D7F0EA82A2765F0D74E2FDE820D04B56" box="[476,706,606,631]" italics="true" pageId="17" pageNumber="18">Rhabdodon priscus</emphasis>
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</taxonomicName>
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and all the descendants of this common ancestor (
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<bibRefCitation id="81154B61A2765F0D75F8FD2823FE4B96" author="Weishampel, D. B. & Jianu, C. - M. & Csiki, Z. & Norman, D. B." box="[198,492,670,695]" journalOrPublisher="Journal of Systematic Palaeontology" pageId="17" pageNumber="18" pagination="65 - 123" part="1" refId="ref23503" refString="Weishampel, D. B., Jianu, C. - M., Csiki, Z., and Norman, D. B. 2003. Osteology and phylogeny of Zalmoxes (n. g.), an unusual euornithopod dinosaur from the Latest Cretaceous of Romania. Journal of Systematic Palaeontology, 1 (2): 65 - 123. https: // doi. org / 10.1017 / s 1477201903001032" title="Osteology and phylogeny of Zalmoxes (n. g.), an unusual euornithopod dinosaur from the Latest Cretaceous of Romania" type="journal article" year="2003">Weishampel et al., 2003</bibRefCitation>
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).
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</paragraph>
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<paragraph id="E53B3690A2765F0D75FEFD08230E4DB6" blockId="17.[192,793,542,1911]" pageId="17" pageNumber="18">
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<emphasis id="D7F0EA82A2765F0D75FEFD0820714BF6" bold="true" box="[192,611,702,727]" pageId="17" pageNumber="18">Unambiguous synapomorphies.</emphasis>
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<taxonomicName id="22844D13A2765F0D7746FD0822FE4BD6" family="Rhabdodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Chordata" rank="family">Rhabdodontidae</taxonomicName>
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is characterized by the presence of apicobasally extending ridges on the cutting surfaces of unworn cheek teeth (lingual surface of maxillary teeth, labial surface of dentary teeth) (138.1), acromion process of scapula does not extend beyond the edge of the coracoid (196.0), medial condyle of the humerus transversely wider than the lateral (216.2), proximal flange present on the ulna (219.1), preacetabular process of the ilium twisted around its long axis (249.1), pubic peduncle of the ischium dorsoventrally compressed (280.1), and distal condyles of the femur expanded both cranially and caudally (304.1), metatarsal V absent (320.1).
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</paragraph>
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<paragraph id="E53B3690A2765F0D75FEFB28211C4C76" blockId="17.[192,793,542,1911]" pageId="17" pageNumber="18">
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<emphasis id="D7F0EA82A2765F0D75FEFB2823294D96" bold="true" box="[192,315,1182,1207]" pageId="17" pageNumber="18">Topology.</emphasis>
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This small clade contains two genera,
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<taxonomicName id="22844D13A2765F0D75FEFB0823214DF6" authorityName="Weishampel, Jianu, Csiki & Norman" authorityYear="2003" box="[192,307,1214,1239]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2765F0D75FEFB0823214DF6" box="[192,307,1214,1239]" italics="true" pageId="17" pageNumber="18">Zalmoxes</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="22844D13A2765F0D7447FB0820114DF6" box="[377,515,1214,1239]" class="Squamata" family="Psammophiidae" genus="Rhabdodon" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2765F0D7447FB0820114DF6" box="[377,515,1214,1239]" italics="true" pageId="17" pageNumber="18">Rhabdodon</emphasis>
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</taxonomicName>
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, both known from the Late Cretaceous of Europe. This clade is recovered in the parsimony analysis with a jackknife value of 81 and a Bremer support of 4, and in the Bayesian analysis with a posterior probability of 1.
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</paragraph>
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<paragraph id="E53B3690A2765F0D75FEFAE8203B4C56" blockId="17.[192,793,542,1911]" box="[192,553,1374,1400]" pageId="17" pageNumber="18">
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DRYOMORPHA
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<bibRefCitation id="81154B61A2765F0D74B7FAE8203B4C56" author="Sereno, P. C." box="[393,553,1374,1400]" journalOrPublisher="National Geographic Research" pageId="17" pageNumber="18" pagination="234 - 256" part="2" refId="ref22197" refString="Sereno, P. C. 1986. Phylogeny of the bird-hipped dinosaurs (Order Ornithischia). National Geographic Research, 2: 234 - 256." title="Phylogeny of the bird-hipped dinosaurs (Order Ornithischia)" type="journal article" year="1986">Sereno, 1986</bibRefCitation>
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</paragraph>
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<paragraph id="E53B3690A2765F0D75FEFAC8231E4CD6" blockId="17.[192,793,542,1911]" pageId="17" pageNumber="18">
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<emphasis id="D7F0EA82A2765F0D75FEFAC823FE4CB6" bold="true" box="[192,492,1406,1431]" pageId="17" pageNumber="18">Phylogenetic definition.</emphasis>
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The most inclusive clade containing
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<taxonomicName id="22844D13A2765F0D7474FA2820C84C96" authority="(Marsh, 1878)" baseAuthorityName="Marsh" baseAuthorityYear="1878" box="[330,730,1438,1463]" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="altus">
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<emphasis id="D7F0EA82A2765F0D7474FA2820094C96" box="[330,539,1438,1463]" italics="true" pageId="17" pageNumber="18">Dryosaurus altus</emphasis>
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(
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<bibRefCitation id="81154B61A2765F0D770CFA2820C74C96" author="Marsh, O. C." box="[562,725,1438,1463]" journalOrPublisher="American Journal of Science, s 3" pageId="17" pageNumber="18" pagination="411 - 416" part="16" refId="ref19615" refString="Marsh, O. C. 1878. Principal characters of American Jurassic dinosaurs. American Journal of Science, s 3 - 16 (95): 411 - 416." title="Principal characters of American Jurassic dinosaurs" type="journal article" year="1878">Marsh, 1878</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName id="22844D13A2765F0D75FEFA08208C4CF6" authority="Parks, 1922" authorityName="Parks" authorityYear="1922" box="[192,670,1470,1495]" class="Reptilia" family="Lambeosauridae" genus="Parasaurolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="walkeri">
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<emphasis id="D7F0EA82A2765F0D75FEFA0823E24CF6" box="[192,496,1470,1495]" italics="true" pageId="17" pageNumber="18">Parasaurolophus walkeri</emphasis>
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<bibRefCitation id="81154B61A2765F0D773AFA08208C4CF6" author="Parks, W. A." box="[516,670,1470,1495]" journalOrPublisher="University of Toronto Studies, Geological Series" pageId="17" pageNumber="18" pagination="1 - 32" part="13" refId="ref21238" refString="Parks, W. A. 1922. Parasaurolophus walkeri: a new genus and species of crested trachodont dinosaur. University of Toronto Studies, Geological Series, 13: 1 - 32." title="Parasaurolophus walkeri: a new genus and species of crested trachodont dinosaur" type="journal article" year="1922">Parks, 1922</bibRefCitation>
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</taxonomicName>
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(
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<bibRefCitation id="81154B61A2765F0D7786FA0823104CD6" author="Sereno, P. C." journalOrPublisher="TaxonSearch" pageId="17" pageNumber="18" publicationUrl="http://www.taxonsearch.org/Archive/stem-archosauria-1.0.php" refId="ref22354" refString="Sereno, P. C. 2005. Stem Archosauria - TaxonSearch. http: // www. taxonsearch. org / Archive / stem-archosauria- 1.0. php" title="Stem Archosauria" type="url" year="2005">Sereno, 2005</bibRefCitation>
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).
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</paragraph>
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<paragraph id="E53B3690A2765F0D75FEFA48264249C3" blockId="17.[192,793,542,1911]" lastBlockId="17.[840,1441,169,898]" pageId="17" pageNumber="18">
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<emphasis id="D7F0EA82A2765F0D75FEFA48204C4F36" bold="true" box="[192,606,1534,1559]" pageId="17" pageNumber="18">Unambiguous synapomorphies.</emphasis>
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In both parsimony and Bayesian analyses, Dryomorpha is characterized by a premaxilla with a posterolateral process that contacts the lacrimal (6.1), a maxilla with a rostrolateral process (18.1), a lacrimal that fits into a slot in the dorsal process of the maxilla (27.1), a quadratojugal that lacks a dorsal process (57.1), a coronoid process with subequal craniocaudal widths of dentary and surangular (109.1), cheek teeth with secondary ridges that are thin and formed only from enamel (145.1), a radius that is triangular in distal view (224.1), and an ischium with the obturator process within the proximal 25% of the ischium (287.1).
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</paragraph>
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<paragraph id="E53B3690A2765F0D76BCFF5F260F48C3" blockId="17.[840,1441,169,898]" pageId="17" pageNumber="18">In the parsimony analysis, Dryomorpha is further supported by a maxillary tooth row that is medially bowed in ventral view (26.0), a jugal that is excluded from the antorbital fenestra by lacrimal-maxilla contact (47.0), a quadratojugal that lacks a foramen through the center of the element (58.0), and absent ossified hypaxial tendons on the caudal vertebrae (322.0).</paragraph>
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<paragraph id="E53B3690A2765F0D76BCFE5F219F4BC3" blockId="17.[840,1441,169,898]" pageId="17" pageNumber="18">In the Bayesian analysis, Dryomorpha is also supported by the presence of a quadrate buttress or "hamular process" (63.1), presence of a quadrate (or paraquadratic) foramen or notch on the boundary between the quadrate and quadratojugal (65.1), and presence of a short midline process of the predentary dorsal to the dentary symphysis (90.1)</paragraph>
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<paragraph id="E53B3690A2765F0D7676FD5F26544AA3" blockId="17.[840,1441,169,898]" pageId="17" pageNumber="18">
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<emphasis id="D7F0EA82A2765F0D7676FD5F21D14A23" bold="true" box="[840,963,745,770]" pageId="17" pageNumber="18">Topology.</emphasis>
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In both parsimony and Bayesian analyses (
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<figureCitation id="7DBF2A15A2765F0D76B9FCBF21E54A03" box="[903,1015,777,802]" captionStart="FIGURE 6" captionStartId="10.[194,285,1048,1071]" captionTargetBox="[204,1427,175,1009]" captionTargetId="graphics-257@10.[204,1427,175,1010]" captionTargetPageId="10" captionText="FIGURE 6. Parsimony (left) and Bayesian (right) trees plotted together. Strict consensus of 84 MPTs after pruning Oryctodromeus, Atlascopscosaurus, Planicoxa, Cumnoria, Cedrorestes, and NHMUK R28860. Jackknife values above 20 (with 10% chance of character removal) are shown above and to the left of their respective nodes. Bremer supports above one are shown below and to the left of their respective nodes, and bolded. CI=0.272, RI=0.634. Maximum clade credibility tree produced by Bayesian analysis showing posterior probabilities below and to the right of their respective node." figureDoi="http://doi.org/10.5281/zenodo.11063322" httpUri="https://zenodo.org/record/11063322/files/figure.png" pageId="17" pageNumber="18">Figure 6</figureCitation>
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), Dryomorpha contains two sister clades:
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<taxonomicName id="22844D13A2765F0D7699FC9F26574A63" authorityName="Milner and Norman" authorityYear="1984" box="[935,1093,809,834]" family="Dryosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Chordata" rank="family">Dryosauridae</taxonomicName>
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and Ankylopollexia. Support for this node is moderate (jackknife=19, Bremmer support=4, PP=0.27).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |