172 lines
25 KiB
XML
172 lines
25 KiB
XML
<document id="A722F99091AC0D719B31D29DFEEB22EA" ID-DOI="10.26879/702" ID-ISSN="1094-8074" ID-Zenodo-Dep="11063310" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1714014708037" checkinUser="felipe" docAuthor="Poole, Karen E." docDate="2016" docId="6D2D8786A2685F13741EFD4F21F34883" docLanguage="en" docName="PalaeontolElectron.25.3.a30.pdf" docOrigin="Palaeontologia Electronica (a 30) 25 (3)" docSource="http://dx.doi.org/10.26879/702" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Thescelosauridae Sternberg 1937" docType="treatment" docVersion="2" lastPageNumber="16" masterDocId="9114FFFEA2675F1C753EFFB622124921" masterDocTitle="Phylogeny of iguanodontian dinosaurs and the evolution of quadrupedality" masterLastPageNumber="65" masterPageNumber="1" pageNumber="16" updateTime="1714997633796" updateUser="julia" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="279CCD8A8D05544629A1B05085F1CE2C">Phylogeny of iguanodontian dinosaurs and the evolution of quadrupedality</mods:title>
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<mods:namePart id="004F2FED035DC197F1F03D939A5EC514">Poole, Karen E.</mods:namePart>
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<mods:title id="65F6EFE4DEBF0A4BAF597EF32FD0666B">Palaeontologia Electronica</mods:title>
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<mods:date id="ADF389AF818DC0DBB73C177F3CE71860">2016</mods:date>
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<mods:title id="7DF608110A778228C8D9455C5D329BC6">a 30</mods:title>
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<mods:number id="7F6A4CB6C6741FAA0C371ACC10BD722C">25</mods:number>
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<mods:identifier id="D7849C7B99651BEA93CEA1F9CCF31032" type="ISSN">1094-8074</mods:identifier>
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<treatment id="6D2D8786A2685F13741EFD4F21F34883" LSID="urn:lsid:plazi:treatment:6D2D8786A2685F13741EFD4F21F34883" httpUri="http://treatment.plazi.org/id/6D2D8786A2685F13741EFD4F21F34883" lastPageNumber="16" pageId="15" pageNumber="16">
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<subSubSection id="AD9E651BA2685F13741EFD4F20A54A33" box="[288,695,760,786]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="E53B3690A2685F13741EFD4F20A54A33" blockId="15.[288,695,760,786]" box="[288,695,760,786]" pageId="15" pageNumber="16">
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<heading id="BE7381FCA2685F13741EFD4F20A54A33" box="[288,695,760,786]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="16" reason="8">
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<taxonomicName id="22844D13A2685F13741EFD4F20A54A33" authority="Sternberg, 1937" authorityName="Sternberg" authorityYear="1937" box="[288,695,760,786]" family="Thescelosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="family">
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Thescelosauridae
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<bibRefCitation id="81154B61A2685F1374C4FD4E20A54A33" author="Sternberg, C. M." box="[506,695,760,786]" journalOrPublisher="Proceedings of the Geological Society of America" pageId="15" pageNumber="16" pagination="375" part="1936" refId="ref22679" refString="Sternberg, C. M. 1937. A classification of Thescelosaurus, with a description of a new species. Proceedings of the Geological Society of America, 1936: 375." title="A classification of Thescelosaurus, with a description of a new species" type="journal article" year="1937">Sternberg, 1937</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="AD9E651BA2685F1375FEFC9521F34883" pageId="15" pageNumber="16" type="description">
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<paragraph id="E53B3690A2685F1375FEFC95200F4A9D" blockId="15.[192,793,803,1916]" pageId="15" pageNumber="16">
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<emphasis id="D7F0EA82A2685F1375FEFC9520134A1D" bold="true" box="[192,513,803,828]" pageId="15" pageNumber="16">Phylogenetic definition.</emphasis>
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All neornithischians more closely related to
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<taxonomicName id="22844D13A2685F1374E1FCF5237B4A5D" authority="Gilmore, 1913" authorityName="Gilmore" authorityYear="1913" class="Reptilia" family="Hypsilophodontidae" genus="Thescelosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="neglectus">
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<emphasis id="D7F0EA82A2685F1374E1FCF5210A4A7D" box="[479,792,835,860]" italics="true" pageId="15" pageNumber="16">Thescelosaurus neglectus</emphasis>
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<bibRefCitation id="81154B61A2685F1375FEFCD5237B4A5D" author="Gilmore, C. W." box="[192,361,867,893]" journalOrPublisher="Smithsonian Miscellaneous Collections" pageId="15" pageNumber="16" pagination="1 - 5" part="61" refId="ref17689" refString="Gilmore, C. W. 1913. A new dinosaur from the Lance Formation of Wyoming. Smithsonian Miscellaneous Collections, 61 (5): 1 - 5." title="A new dinosaur from the Lance Formation of Wyoming" type="journal article" year="1913">Gilmore, 1913</bibRefCitation>
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</taxonomicName>
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than to
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<taxonomicName id="22844D13A2685F1374F1FCD522ED4ABD" authority="Huxley, 1869" authorityName="Huxley" authorityYear="1869" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="foxii">
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<emphasis id="D7F0EA82A2685F1374F1FCD520A54A5D" box="[463,695,867,892]" italics="true" pageId="15" pageNumber="16">Hypsilophodon foxii</emphasis>
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<bibRefCitation id="81154B61A2685F1377FFFCD522ED4ABD" author="Huxley, T. H." journalOrPublisher="Abstracts of the Proceedings of the Geological Society (of London)" pageId="15" pageNumber="16" pagination="3 - 4" part="204" refId="ref18591" refString="Huxley, T. H. 1869. On Hypsilophodon, a new genus of Dinosauria. Abstracts of the Proceedings of the Geological Society (of London), 204: 3 - 4." title="On Hypsilophodon, a new genus of Dinosauria" type="journal article" year="1869">Huxley, 1869</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="22844D13A2685F137434FC3520914ABD" authority="(Marsh, 1878)" baseAuthorityName="Marsh" baseAuthorityYear="1878" box="[266,643,899,924]" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="altus">
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<emphasis id="D7F0EA82A2685F137434FC3523C34ABD" box="[266,465,899,924]" italics="true" pageId="15" pageNumber="16">Dryosaurus altus</emphasis>
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(
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<bibRefCitation id="81154B61A2685F1374E1FC3520694ABD" author="Marsh, O. C." box="[479,635,899,924]" journalOrPublisher="American Journal of Science, s 3" pageId="15" pageNumber="16" pagination="411 - 416" part="16" refId="ref19615" refString="Marsh, O. C. 1878. Principal characters of American Jurassic dinosaurs. American Journal of Science, s 3 - 16 (95): 411 - 416." title="Principal characters of American Jurassic dinosaurs" type="journal article" year="1878">Marsh, 1878</bibRefCitation>
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)
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</taxonomicName>
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, or
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<taxonomicName id="22844D13A2685F137792FC3520044A9D" authority="Parks, 1922" authorityName="Parks" authorityYear="1922" class="Reptilia" family="Lambeosauridae" genus="Parasaurolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="walkeri">
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<emphasis id="D7F0EA82A2685F137792FC3523934A9D" italics="true" pageId="15" pageNumber="16">Parasaurolophus walkeri</emphasis>
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<bibRefCitation id="81154B61A2685F1374B6FC1520044A9D" author="Parks, W. A." box="[392,534,931,956]" journalOrPublisher="University of Toronto Studies, Geological Series" pageId="15" pageNumber="16" pagination="1 - 32" part="13" refId="ref21238" refString="Parks, W. A. 1922. Parasaurolophus walkeri: a new genus and species of crested trachodont dinosaur. University of Toronto Studies, Geological Series, 13: 1 - 32." title="Parasaurolophus walkeri: a new genus and species of crested trachodont dinosaur" type="journal article" year="1922">Parks, 1922</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="E53B3690A2685F1375FEFC7520B34DBD" blockId="15.[192,793,803,1916]" pageId="15" pageNumber="16">
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<emphasis id="D7F0EA82A2685F1375FEFC7520734AFD" bold="true" box="[192,609,963,988]" pageId="15" pageNumber="16">Unambiguous synapomorphies.</emphasis>
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In the parsimony analysis (
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<figureCitation id="7DBF2A15A2685F1374B1FC5520114ADD" box="[399,515,995,1020]" captionStart="FIGURE 7" captionStartId="11.[200,291,1774,1797]" captionTargetBox="[198,1439,164,1749]" captionTargetId="figure-91@11.[193,1440,164,1750]" captionTargetPageId="11" captionText="FIGURE 7. Time scaled parsimony tree, showing assigned age ranges and broad-scale geographic data. Jackknife values above 20 (with 10% chance of character removal) are shown above and to the left of their respective nodes. Bremer supports above one are shown below and to the left of their respective nodes, and bolded." figureDoi="http://doi.org/10.5281/zenodo.11063324" httpUri="https://zenodo.org/record/11063324/files/figure.png" pageId="15" pageNumber="16">Figure 7</figureCitation>
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), thescelosaurids are characterized by three unambiguous synapomorphies: dentary tooth row straight in dorsal view (98.0), 10-12 ridges on the maxillary teeth (136.3), and acromion process of scapula does not extend beyond the edge of the coracoid (196.0).
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</paragraph>
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<paragraph id="E53B3690A2685F1375C4FB1523FE4C1D" blockId="15.[192,793,803,1916]" pageId="15" pageNumber="16">
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In the Bayesian analysis (
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<figureCitation id="7DBF2A15A2685F137704FB1520B54D9D" box="[570,679,1187,1212]" captionStart="FIGURE 8" captionStartId="12.[196,286,1624,1647]" captionTargetBox="[195,1436,163,1599]" captionTargetId="graphics-57@12.[196,1437,163,1599]" captionTargetPageId="12" captionText="FIGURE 8. Maximum Clade Credibility tree produced by Bayesian analysis. Posterior probabilities are shown to the left of their nodes. The geologic timescale is shown across the top. Tips represent the average age found for each OTU across all sampled trees." figureDoi="http://doi.org/10.5281/zenodo.11063326" httpUri="https://zenodo.org/record/11063326/files/figure.png" pageId="15" pageNumber="16">Figure 8</figureCitation>
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), a more extensive
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<taxonomicName id="22844D13A2685F13747CFB7520064DFD" authorityName="Sternberg" authorityYear="1937" box="[322,532,1219,1244]" family="Thescelosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Thescelosauridae</taxonomicName>
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is characterized by two unambiguous synapomorphies: 10-12 ridges on the maxillary teeth (136.3) and 9-11 ridges on the dentary teeth (137.3).
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</paragraph>
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<paragraph id="E53B3690A2685F1375FEFAF520F34F9D" blockId="15.[192,793,803,1916]" pageId="15" pageNumber="16">
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<emphasis id="D7F0EA82A2685F1375FEFAF523294C7D" bold="true" box="[192,315,1347,1372]" pageId="15" pageNumber="16">Topology.</emphasis>
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<taxonomicName id="22844D13A2685F137474FAF5200E4C7D" authorityName="Sternberg" authorityYear="1937" box="[330,540,1347,1372]" family="Thescelosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Thescelosauridae</taxonomicName>
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is recovered in the parsimony analysis (
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<figureCitation id="7DBF2A15A2685F13748FFAD520354C5D" box="[433,551,1379,1404]" captionStart="FIGURE 6" captionStartId="10.[194,285,1048,1071]" captionTargetBox="[204,1427,175,1009]" captionTargetId="graphics-257@10.[204,1427,175,1010]" captionTargetPageId="10" captionText="FIGURE 6. Parsimony (left) and Bayesian (right) trees plotted together. Strict consensus of 84 MPTs after pruning Oryctodromeus, Atlascopscosaurus, Planicoxa, Cumnoria, Cedrorestes, and NHMUK R28860. Jackknife values above 20 (with 10% chance of character removal) are shown above and to the left of their respective nodes. Bremer supports above one are shown below and to the left of their respective nodes, and bolded. CI=0.272, RI=0.634. Maximum clade credibility tree produced by Bayesian analysis showing posterior probabilities below and to the right of their respective node." figureDoi="http://doi.org/10.5281/zenodo.11063322" httpUri="https://zenodo.org/record/11063322/files/figure.png" pageId="15" pageNumber="16">Figures 6</figureCitation>
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,
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<figureCitation id="7DBF2A15A2685F13770DFAD520514C5D" box="[563,579,1379,1404]" captionStart="FIGURE 7" captionStartId="11.[200,291,1774,1797]" captionTargetBox="[198,1439,164,1749]" captionTargetId="figure-91@11.[193,1440,164,1750]" captionTargetPageId="11" captionText="FIGURE 7. Time scaled parsimony tree, showing assigned age ranges and broad-scale geographic data. Jackknife values above 20 (with 10% chance of character removal) are shown above and to the left of their respective nodes. Bremer supports above one are shown below and to the left of their respective nodes, and bolded." figureDoi="http://doi.org/10.5281/zenodo.11063324" httpUri="https://zenodo.org/record/11063324/files/figure.png" pageId="15" pageNumber="16">7</figureCitation>
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) containing
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<emphasis id="D7F0EA82A2685F1377EAFAD520DE4CBD" italics="true" pageId="15" pageNumber="16">
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<taxonomicName id="22844D13A2685F1377EAFAD523564CBD" baseAuthorityName="Norman" baseAuthorityYear="2004" class="Reptilia" family="Hypsilophodontidae" genus="Thescelosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">Thescelosaurus</taxonomicName>
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,
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<taxonomicName id="22844D13A2685F137459FA35201F4CBD" box="[359,525,1411,1436]" family="Hypsilophodontidae" genus="Parksosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">Parksosaurus</taxonomicName>
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,
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<taxonomicName id="22844D13A2685F13770FFA3520DA4CBD" box="[561,712,1411,1436]" family="Hypsilophodontidae" genus="Orodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">Orodromeus</taxonomicName>
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,
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</emphasis>
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and
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<taxonomicName id="22844D13A2685F1375FEFA1523604C9D" authorityName="Sues" authorityYear="1980" box="[192,370,1443,1468]" family="Hypsilophodontidae" genus="Zephyrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2685F1375FEFA1523604C9D" box="[192,370,1443,1468]" italics="true" pageId="15" pageNumber="16">Zephyrosaurus</emphasis>
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</taxonomicName>
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, though the group has poor support (Jackknife value=7). Unlike in
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<bibRefCitation id="81154B61A2685F13771FFA7520A44CFD" author="Boyd, C. A." box="[545,694,1475,1500]" journalOrPublisher="The University of Texas at Austin, Austin, Texas" pageId="15" pageNumber="16" refId="ref15426" refString="Boyd, C. A. 2012. Taxonomic revision of latest Cretaceous North American basal neornithischian taxa and a phylogenetic analysis of basal ornithischian relationships. Ph. D. dissertation, The University of Texas at Austin, Austin, Texas." title="Taxonomic revision of latest Cretaceous North American basal neornithischian taxa and a phylogenetic analysis of basal ornithischian relationships" type="book" year="2012">Boyd (2012)</bibRefCitation>
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,
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<taxonomicName id="22844D13A2685F1377FEFA7520EC4CFD" box="[704,766,1475,1500]" class="Magnoliopsida" family="Thescelosauridae" genus="Haya" higherTaxonomySource="CoL" kingdom="Animalia" order="Caryophyllales" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2685F1377FEFA7520EC4CFD" box="[704,766,1475,1500]" italics="true" pageId="15" pageNumber="16">Haya</emphasis>
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</taxonomicName>
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is recovered at a node basal to thescelosaurids, rather than within the Thescelosaurinae.
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<taxonomicName id="22844D13A2685F13779EF9B5232E4F1D" authorityName="Calvo, Porfiri & Novas" authorityYear="2007" genus="Macrogryphosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2685F13779EF9B5232E4F1D" italics="true" pageId="15" pageNumber="16">Macrogryphosaurus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="22844D13A2685F137449F99520124F1D" box="[375,512,1571,1596]" class="Reptilia" family="Hadrosauridae" genus="Talenkauen" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2685F137449F99520124F1D" box="[375,512,1571,1596]" italics="true" pageId="15" pageNumber="16">Talenkauen</emphasis>
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</taxonomicName>
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, which Boyd also found within Thescelosaurinae, are recovered within iguanodontia, based on the presence of opisthocoelus cervical vertebrae, and a postacetabular process that is less than 30% of total ilium length.
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</paragraph>
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<paragraph id="E53B3690A2685F1375C4F97521F34883" blockId="15.[192,793,803,1916]" lastBlockId="15.[840,1441,168,418]" pageId="15" pageNumber="16">
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In the Bayesian analysis (
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<figureCitation id="7DBF2A15A2685F137714F97520B24FFD" box="[554,672,1731,1756]" captionStart="FIGURE 6" captionStartId="10.[194,285,1048,1071]" captionTargetBox="[204,1427,175,1009]" captionTargetId="graphics-257@10.[204,1427,175,1010]" captionTargetPageId="10" captionText="FIGURE 6. Parsimony (left) and Bayesian (right) trees plotted together. Strict consensus of 84 MPTs after pruning Oryctodromeus, Atlascopscosaurus, Planicoxa, Cumnoria, Cedrorestes, and NHMUK R28860. Jackknife values above 20 (with 10% chance of character removal) are shown above and to the left of their respective nodes. Bremer supports above one are shown below and to the left of their respective nodes, and bolded. CI=0.272, RI=0.634. Maximum clade credibility tree produced by Bayesian analysis showing posterior probabilities below and to the right of their respective node." figureDoi="http://doi.org/10.5281/zenodo.11063322" httpUri="https://zenodo.org/record/11063322/files/figure.png" pageId="15" pageNumber="16">Figures 6</figureCitation>
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,
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<figureCitation id="7DBF2A15A2685F137793F97520AD4FFD" box="[685,703,1731,1756]" captionStart="FIGURE 8" captionStartId="12.[196,286,1624,1647]" captionTargetBox="[195,1436,163,1599]" captionTargetId="graphics-57@12.[196,1437,163,1599]" captionTargetPageId="12" captionText="FIGURE 8. Maximum Clade Credibility tree produced by Bayesian analysis. Posterior probabilities are shown to the left of their nodes. The geologic timescale is shown across the top. Tips represent the average age found for each OTU across all sampled trees." figureDoi="http://doi.org/10.5281/zenodo.11063326" httpUri="https://zenodo.org/record/11063326/files/figure.png" pageId="15" pageNumber="16">8</figureCitation>
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),
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<taxonomicName id="22844D13A2685F1377EAF97523454FDD" authorityName="Sternberg" authorityYear="1937" family="Thescelosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Thescelosauridae</taxonomicName>
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is nested within a larger
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<taxonomicName id="22844D13A2685F1377B2F95523394E3D" family="Hypsilophodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Hypsilophodontidae</taxonomicName>
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. Though it only has moderate support (PP=0.59), the internal nodes have higher posterior probabilities. The genera included in this group match those recovered in the parsimony analysis; the one difference in topology is that
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<taxonomicName id="22844D13A2685F137032FF1E27B249E0" box="[1292,1440,168,193]" family="Hypsilophodontidae" genus="Orodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2685F137032FF1E27B249E0" box="[1292,1440,168,193]" italics="true" pageId="15" pageNumber="16">Orodromeus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="22844D13A2685F137641FF7F262349C3" authorityName="Sues" authorityYear="1980" box="[895,1073,201,226]" family="Hypsilophodontidae" genus="Zephyrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2685F137641FF7F262349C3" box="[895,1073,201,226]" italics="true" pageId="15" pageNumber="16">Zephyrosaurus</emphasis>
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</taxonomicName>
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form successive outgroups to
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<taxonomicName id="22844D13A2685F137676FF5F26164823" baseAuthorityName="Norman" baseAuthorityYear="2004" box="[840,1028,233,258]" class="Reptilia" family="Hypsilophodontidae" genus="Thescelosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2685F137676FF5F26164823" box="[840,1028,233,258]" italics="true" pageId="15" pageNumber="16">Thescelosaurus</emphasis>
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</taxonomicName>
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+
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<taxonomicName id="22844D13A2685F13711CFF5F26D74823" box="[1058,1221,233,258]" family="Hypsilophodontidae" genus="Parksosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2685F13711CFF5F26D74823" box="[1058,1221,233,258]" italics="true" pageId="15" pageNumber="16">Parksosaurus</emphasis>
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</taxonomicName>
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instead of forming a distinct Orodrominae (as found by
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<bibRefCitation id="81154B61A2685F137033FEBF27854803" author="Boyd, C. A." box="[1293,1431,265,290]" journalOrPublisher="The University of Texas at Austin, Austin, Texas" pageId="15" pageNumber="16" refId="ref15426" refString="Boyd, C. A. 2012. Taxonomic revision of latest Cretaceous North American basal neornithischian taxa and a phylogenetic analysis of basal ornithischian relationships. Ph. D. dissertation, The University of Texas at Austin, Austin, Texas." title="Taxonomic revision of latest Cretaceous North American basal neornithischian taxa and a phylogenetic analysis of basal ornithischian relationships" type="book" year="2012">Boyd 2012</bibRefCitation>
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). Though
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<taxonomicName id="22844D13A2685F137693FE9F21F94863" box="[941,1003,297,322]" class="Magnoliopsida" family="Thescelosauridae" genus="Haya" higherTaxonomySource="CoL" kingdom="Animalia" order="Caryophyllales" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2685F137693FE9F21F94863" box="[941,1003,297,322]" italics="true" pageId="15" pageNumber="16">Haya</emphasis>
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</taxonomicName>
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|
and
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<taxonomicName id="22844D13A2685F137110FE9F270B4863" authorityName="Calvo, Porfiri & Novas" authorityYear="2007" box="[1070,1305,297,322]" genus="Macrogryphosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="D7F0EA82A2685F137110FE9F270B4863" box="[1070,1305,297,322]" italics="true" pageId="15" pageNumber="16">Macrogryphosaurus</emphasis>
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</taxonomicName>
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|
are recovered as closely related hypsilophodontids, they are not found within
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<taxonomicName id="22844D13A2685F13712FFEDF26F148A3" authorityName="Sternberg" authorityYear="1937" box="[1041,1251,361,386]" family="Thescelosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Thescelosauridae</taxonomicName>
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as proposed in
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<bibRefCitation id="81154B61A2685F137676FE3F21CF4883" author="Boyd, C. A." box="[840,989,393,418]" journalOrPublisher="The University of Texas at Austin, Austin, Texas" pageId="15" pageNumber="16" refId="ref15426" refString="Boyd, C. A. 2012. Taxonomic revision of latest Cretaceous North American basal neornithischian taxa and a phylogenetic analysis of basal ornithischian relationships. Ph. D. dissertation, The University of Texas at Austin, Austin, Texas." title="Taxonomic revision of latest Cretaceous North American basal neornithischian taxa and a phylogenetic analysis of basal ornithischian relationships" type="book" year="2012">Boyd (2012)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |