159 lines
23 KiB
XML
159 lines
23 KiB
XML
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<mods:title id="67F217832A61B8BA0F74A78BA79A691C">Review of the perciform fish genus Symphysanodon Bleeker (Symphysanodontidae), with descriptions of three new species, S. mona, S. parini, and S. rhax.</mods:title>
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<mods:namePart id="F24044D7B45B51A0E316BB6188562556">William D. Anderson, Jr.</mods:namePart>
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<mods:namePart id="26EA2F7518CFB88D9663E104B87A8B5D">Victor G. Springer</mods:namePart>
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<mods:date id="0F29EBAB7B948BF835981A97076CB9DC">2005</mods:date>
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<treatment id="6CB7CFB3B0451F787BBA4ED09159A859" ID-DOI="http://doi.org/10.5281/zenodo.6266866" ID-GBIF-Taxon="100119269" ID-Zenodo-Dep="6266866" LSID="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" httpUri="http://treatment.plazi.org/id/6CB7CFB3B0451F787BBA4ED09159A859" lastPageNumber="22" pageNumber="16">
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<subSubSection id="2EC5AB03D9686EC7F4AE410FA6007296" type="nomenclature">
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<paragraph id="E73FC2C1548D24BB8B6F609ACA792B6D" pageNumber="16">
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<taxonomicName id="BF851BEC1184832B906ED18A13D83BFC" ID-CoL="53QBD" LSID-ZBK="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon rhax Anderson & Springer 2005:16" order="Perciformes" pageNumber="16" phylum="Chordata" rank="species" species="rhax">Symphysanodon rhax</taxonomicName>
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,
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<taxonomicNameLabel id="D5C9BD4A8D5A0644225D2EFF9A2FFEF3" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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<paragraph id="CD15E9B1A490AEEA94A42E27D650FDDF" pageNumber="16">Maldives Slopefish</paragraph>
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</subSubSection>
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<subSubSection id="A673C2179CB4ED219F7D5D8E3F7F4AF2" type="description">
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<paragraph id="7438C861065D33760AB099B7E9FE204F" pageNumber="16">(Figures 9-15; Tables 4, 5, 8, 9)</paragraph>
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</subSubSection>
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<subSubSection id="B873FC0058907C74C3F50FEF277D2DDE" type="nomenclature">
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<paragraph id="7F848944667F4B1091FDE8706A793E9D" pageNumber="16">Holotype: BMNH 2003.1.5.1, 144 mm SL; Maldive Islands, northern Indian Ocean; 04°11’ N, 72°55’ E; 218 m; R/V DR FRIDTJOF NANSEN cruise: Maldives station no. 13; 21 August 1983.</paragraph>
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</subSubSection>
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<subSubSection id="87D2448010B6EF27D519F0BCED6D1E82" type="nomenclature">
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<paragraph id="07A869ECD2A816F70688DF43DEFAE9CF" pageNumber="16">Paratype. BMNH 2003.1.5.2, 134 mm SL, from the same station as the holotype.</paragraph>
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</subSubSection>
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<subSubSection id="E2345B480D9EF69DE76D094E13E42EFA" type="materials_examined">
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<paragraph id="5CAB544B320E8E9C905EA4287FFE8C0F" pageNumber="16">Other specimens, not part of the type series. BMNH 1997.9.17:2, one specimen, 64 mm SL; R/V DR FRIDTJOF NANSEN survey of the Maldives, 04°05’ N, 73°23’ E - BMNH 1997.9.17:6-8, three specimens, 128-136 mm SL; R/V DR FRIDTJOF NANSEN survey of the Maldives, 04° 11’ N, 72°55’ E.</paragraph>
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</subSubSection>
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<subSubSection id="A4934ECE2855DA63D2A7EA91F6EEB09E" type="diagnosis">
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<paragraph id="B82D8FC199CEEFD69C20BF73F5E6ED23" pageNumber="16">
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Diagnosis. A species of
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<taxonomicName id="820795E0FB5C58E34EB4A6D9772DB8FB" LSID-ZBK="urn:lsid:zoobank.org:act:F334D443-843F-494A-AC1A-947DDCBACA9F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon Bleeker 1878:60" order="Perciformes" pageNumber="16" phylum="Chordata" rank="genus">Symphysanodon</taxonomicName>
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(Figure 10) separable from all other species of the genus, except
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<taxonomicName id="38AE63307A49300F3C842E29109CD7D8" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="16" phylum="Chordata" rank="species" species="berryi">S. berryi</taxonomicName>
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, by the following combination of characters: segmented rays in anal fin 7, tubed lateral-line scales 50, total gillrakers on first gill arch 35-38 (10 or 11 + 25-27), sum of lateral-line scales and gillrakers on individual specimens 85-88, depth of body 20.6-24.8% SL (4.0-4.9 times in SL), length of depressed anal fin 21.8-23.9 % SL, length of anal-fin base 12.6-14.8% SL, length of first anal-fin spine 3.0-4.8% SL, hypurals 1 & 2 autogenous, hypurals 3 & 4 represented by a single plate, and first caudal vertebra without parapophyses. It can be separated from the Atlantic species
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<taxonomicName id="53885A392C90C915B9B51CACDDFA07BE" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="16" phylum="Chordata" rank="species" species="berryi">S. berryi</taxonomicName>
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by a number of morphometric characters (see Table 4 and Figures 11-15). It also differs slightly from
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<taxonomicName id="A5CC280D258CEE7A74D8054CF835607A" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="16" phylum="Chordata" rank="species" species="berryi">S. berryi</taxonomicName>
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in counts of pectoral-fin rays (17 or18, mean = 17.33, for
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<taxonomicName id="70A9740D5006FD330703129F7A8ACDAA" LSID-ZBK="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon rhax Anderson & Springer 2005:16" order="Perciformes" pageNumber="16" phylum="Chordata" rank="species" species="rhax">S. rhax</taxonomicName>
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vs. 16-18, mean = 16.99, for
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<taxonomicName id="7D844E742EAD6743EC822632568D4C40" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="16" phylum="Chordata" rank="species" species="berryi">S. berryi</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="916396A2E3EE66695733A41C7810D48B" type="description">
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<paragraph id="CA78D2362230E7C48B8583D3F9F319C5" pageNumber="19">
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Description. The characters included in the combined description of
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<taxonomicName id="B8D22DDDE577F988B9167F3F6CDB8824" LSID-ZBK="urn:lsid:zoobank.org:act:225CA58F-2A6D-43EE-80D7-6A05F7A05273" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon mona Anderson & Springer 2005:7" order="Perciformes" pageNumber="19" phylum="Chordata" rank="species" species="mona">S. mona</taxonomicName>
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,
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<taxonomicName id="432EA0F0176843B18000791E70C32A73" LSID-ZBK="urn:lsid:zoobank.org:act:A50F8091-80D6-415D-8AAE-6DA1E0C34A6F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon parini Anderson & Springer 2005:11" order="Perciformes" pageNumber="19" phylum="Chordata" rank="species" species="parini">S. parini</taxonomicName>
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, and
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<taxonomicName id="A8F56CD47EB254B993BB73A0227C6025" LSID-ZBK="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon rhax Anderson & Springer 2005:16" order="Perciformes" pageNumber="19" phylum="Chordata" rank="species" species="rhax">S. rhax</taxonomicName>
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and those presented in the species diagnosis form part of the species description. Counts for the holotype are indicated by asterisks. Dorsal-fin rays IX, 10.* Pectoral-fin rays 17 or 18*. Procurrent caudal-fin rays 13 or 14 dorsally, 12 or 13 ventrally. Trisegmental pterygiophores 2 or 3* associated with dorsal fin, 2 or 3 with anal fin. Epineurals associated with first 8* or 9 vertebrae (first 9 vertebrae in 5 of 6 specimens). Uroneurals 2 pairs*. Caudal-fin lobes damaged on all available specimens. Internarial distance contained 4-6 times in snout length; other morphometric data are presented in percentages of SL in Table 5.
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</paragraph>
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</subSubSection>
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<subSubSection id="325371199ADBCB522CB3B04B1E098CFA" type="discussion">
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<paragraph id="5F780D634D70A62A3972D22816E62B1B" pageNumber="21">
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Comparisons of
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<taxonomicName id="A7FD8A16719E42AD807D1B2B4729E8C1" LSID-ZBK="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon rhax Anderson & Springer 2005:16" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="rhax">S. rhax</taxonomicName>
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with other Indian Ocean species of
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<taxonomicName id="1B6D40BEC327D3C923CC1DE63692BA60" LSID-ZBK="urn:lsid:zoobank.org:act:F334D443-843F-494A-AC1A-947DDCBACA9F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon Bleeker 1878:60" order="Perciformes" pageNumber="21" phylum="Chordata" rank="genus">Symphysanodon</taxonomicName>
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.
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<taxonomicName id="73EDFD4BB5C059BE36E47C7B9A790A54" LSID-ZBK="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon rhax Anderson & Springer 2005:16" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="rhax">Symphysanodon rhax</taxonomicName>
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can be distinguished from
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<taxonomicName id="6C1250FF1DD263ED589F8DDA4F9F6F12" LSID-ZBK="urn:lsid:zoobank.org:act:929DAE3E-D784-45C8-B3A4-36E43F287826" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon andersoni Kotthaus 1974:52" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="andersoni">S. andersoni</taxonomicName>
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in having fewer gillrakers (total on first gill arch 35-38 vs. 41or 42) and lateral-line scales (50 vs. 60 or 61), from an undescribed species from the Comoros in having more gillrakers (total on first gill arch 35-38 vs. 28), and from
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<taxonomicName id="68D1D02FCBA225A781900236AEE790DE" LSID-ZBK="urn:lsid:zoobank.org:act:2B84CC23-3EDF-44F3-BA95-63674788BB30" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon maunaloae Anderson 1970:335" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="maunaloae">S. maunaloae</taxonomicName>
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(assuming that species occurs in the Indian Ocean) in having more lateral-line scales (50 vs. 42-47).
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</paragraph>
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</subSubSection>
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<subSubSection id="C89673BC79D6A70C7809DFCDBDF32802" type="description">
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<paragraph id="6884975BB0631AE09AD170E8FC9D3C92" pageNumber="21">
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Sexuality and sexual dimorphism. Histological examination of the gonads of two specimens of
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<taxonomicName id="30D44D977CE1A26FF36FFC23D06110A6" LSID-ZBK="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon rhax Anderson & Springer 2005:16" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="rhax">S. rhax</taxonomicName>
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(134 & 136 mm SL) showed both to be females with no evidence of hermaphroditism. Four specimens have short pelvic fins (20 ->26% SL) and two have produced pelvic fins (>64 &>67% SL). Because both specimens known to be female by histological examination have short pelvic fins and because females of the apparent closest relative of this species,
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<taxonomicName id="B956029E213B086CC1225FFDCE9F5E23" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="berryi">S. berryi</taxonomicName>
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, have short pelvic fins (in contrast with males which frequently have very well-produced pelvic fins), it is reasonable to assume that the individuals of
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<taxonomicName id="571CA6D019CD3BDFED7FEAA766C8D109" LSID-ZBK="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon rhax Anderson & Springer 2005:16" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="rhax">S. rhax</taxonomicName>
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with produced pelvic fins are males.
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<taxonomicName id="2445DA0E1BDE940A1BA90717AD6DE736" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="berryi">Symphysanodon berryi</taxonomicName>
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is also sexually dimorphic with regard to lengths of caudal-fin lobes. Unfortunately, both caudal-fin lobes are damaged on all six specimens of
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<taxonomicName id="D2EFD0D8ED82265C4336986449349EC2" LSID-ZBK="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon rhax Anderson & Springer 2005:16" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="rhax">S. rhax</taxonomicName>
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, leaving the question of sexual dimorphism in this character unanswered.
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</paragraph>
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</subSubSection>
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<subSubSection id="5C48EC1D37E1B76507B71C5EC2AB9BCE" type="distribution">
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<paragraph id="DCC73CE3B4DA0FCC3DDEEE72B804AEE2" pageNumber="21">Distribution. Known only from the northern Indian Ocean off the Maldive Islands (Figure 9). Depth of capture, 218 m, is available for only one collection.</paragraph>
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</subSubSection>
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<subSubSection id="2E8A6CA45919BE4B4F5043D0B5B9E4C7" type="discussion">
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<paragraph id="12D4C3795B7F8FAE6E12B9CCB50988E9" pageNumber="21">
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Remarks. Two of the six specimens of
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<taxonomicName id="FBB144FE3AFE45D8C8A68F968161C340" LSID-ZBK="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon rhax Anderson & Springer 2005:16" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="rhax">S. rhax</taxonomicName>
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(i.e., the type specimens) examined were reported under an account of
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<taxonomicName id="C6C01D0823D056CA2FBFE41F85A7BAB4" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="berryi">S. berryi</taxonomicName>
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by Anderson (2003:1306), who wrote “2 specimens(134 and 144 mm standard length), indistinguishable from this species [
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<taxonomicName id="FA1CD5135510BCAEDEA29156C26C6511" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="berryi">S. berryi</taxonomicName>
|
||
], have been obtained off the Maldives, southwest of Sri Lanka in the Indian Ocean.” Three other specimens (BMNH 1997.9.17:6-8, 128-136 mm SL) reported herein, along with another two (MRS 0455/97, 118 & 146 mm total length) from the Maldives, are listed under
|
||
<taxonomicName id="54710B7867C11FE757267D9DEDFB6D72" LSID-ZBK="urn:lsid:zoobank.org:act:F334D443-843F-494A-AC1A-947DDCBACA9F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon Bleeker 1878:60" order="Perciformes" pageNumber="21" phylum="Chordata" rank="genus">Symphysanodon</taxonomicName>
|
||
sp. in Adam et al. (1998). Meristic and much of the morphometric data for
|
||
<taxonomicName id="C129A56D9831E425825F7C84FF50DA3D" LSID-ZBK="urn:lsid:zoobank.org:act:29BB4082-BB0D-48A2-8FDD-919107F1B13F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon rhax Anderson & Springer 2005:16" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="rhax">S. rhax</taxonomicName>
|
||
agree closely with those for specimens of
|
||
<taxonomicName id="01CBB4BB72C1E17A86E017CEA4F85B96" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="berryi">S. berryi</taxonomicName>
|
||
, but ranges for some measurements do not overlap or overlap only slightly with those of
|
||
<taxonomicName id="9D738A498511D27B05B0D9B368CB6B63" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="berryi">S. berryi</taxonomicName>
|
||
(see Table 4).
|
||
</paragraph>
|
||
<paragraph id="E773AE5EF496C059AEDBF166818FA7C8" pageNumber="21">
|
||
It can be argued that the differences noted in Table 4 are artifacts of the small sample size from the Maldives and not sufficient to recognize those specimens as representing a species distinct from
|
||
<taxonomicName id="2E17B7AB1B89F60A5EA44AE793841130" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="21" phylum="Chordata" rank="species" species="berryi">Symphysanodon berryi</taxonomicName>
|
||
, but when the data for the measurements shown in Table 4 are plotted (see Figures 11-15), it is obvious that the Atlantic and Indian Ocean populations belong to different species. Also, closure of the Tethys Sea by Late Miocene (Smith et al., 1994) would have provided the long-term geographic isolation necessary for allopatric speciation to occur. Gill and Kemp (2002) addressed the problem of the taxonomy of widely distributed Indo-Pacific shore fishes, maintaining “that well-diagnosed geographic forms and subspecies should be awarded full-species status” (p. 165). We concur with their thinking and believe that it applies also to fishes, such as species of
|
||
<taxonomicName id="628D3A1E8D44436C34637310E0CB14D6" LSID-ZBK="urn:lsid:zoobank.org:act:F334D443-843F-494A-AC1A-947DDCBACA9F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon Bleeker 1878:60" order="Perciformes" pageNumber="21" phylum="Chordata" rank="genus">Symphysanodon</taxonomicName>
|
||
, that occur over the continental shelf, upper continental slope, and submarine ridges and in insular waters of similar depths. Consequently, we have chosen to describe the Maldive form as a new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CAF4466E22FE99765402E0EBAAB48CD1" type="etymology">
|
||
<paragraph id="8F853378F1EC4861431F7D48C4F61EA8" pageNumber="22">
|
||
Etymology. The name rhax (berry or grape) is from the Greek and is an allusion to the similarity of this new species to
|
||
<taxonomicName id="33EEAA564BC668E112D2819D9BC1A166" LSID-ZBK="urn:lsid:zoobank.org:act:10736DBF-680B-4D45-ADDA-A63828ACEF46" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon berryi Anderson 1970:338" order="Perciformes" pageNumber="22" phylum="Chordata" rank="species" species="berryi">S. berryi</taxonomicName>
|
||
. The new name is a feminine noun in apposition to
|
||
<taxonomicName id="4DC82E3D2AB0395BC71507262DBB7434" LSID-ZBK="urn:lsid:zoobank.org:act:F334D443-843F-494A-AC1A-947DDCBACA9F" family="Symphysanodontidae" genus="Symphysanodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Symphysanodon Bleeker 1878:60" order="Perciformes" pageNumber="22" phylum="Chordata" rank="genus">Symphysanodon</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |