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<document id="ACEF85CDD90D0F93962328B9C1D76224" ID-CLB-Dataset="25190" ID-DOI="10.11646/zootaxa.4687.1.1" ID-GBIF-Dataset="de00ca6c-2843-4ae1-a39c-a29603947612" ID-ISSN="1175-5326" ID-Zenodo-Dep="3508098" ID-ZooBank="CE73264D-C234-4B82-A634-CAD6254C5957" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571388220410" checkinUser="plazi" docAuthor="Ho, - Z." docDate="2019" docId="6C3DA91C51A918C2FF0EFA60EC7B1DE8" docLanguage="en" docName="zootaxa.4687.1.1.pdf.imf" docOrigin="Zootaxa 4687 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Triangulara Pimpasalee 2016" docType="treatment" docVersion="19" lastPageNumber="149" masterDocId="9004D164513B1856FF99FF9FEF381C47" masterDocTitle="The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species" masterLastPageNumber="174" masterPageNumber="1" pageNumber="147" updateTime="1698760145766" updateUser="ExternalLinkService">
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<mods:title id="108C0F4CDF0D8A96D71F7CBB1A5D1518">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<treatment id="6C3DA91C51A918C2FF0EFA60EC7B1DE8" ID-DOI="http://doi.org/10.5281/zenodo.4688983" ID-GBIF-Taxon="159757319" ID-Zenodo-Dep="4688983" LSID="urn:lsid:plazi:treatment:6C3DA91C51A918C2FF0EFA60EC7B1DE8" httpUri="http://treatment.plazi.org/id/6C3DA91C51A918C2FF0EFA60EC7B1DE8" lastPageId="148" lastPageNumber="149" pageId="146" pageNumber="147">
<subSubSection id="AC8E4B8151A918C4FF0EFA60ED3A1A5D" box="[151,514,1535,1562]" pageId="146" pageNumber="147" type="nomenclature">
<paragraph id="E42B180A51A918C4FF0EFA60ED3A1A5D" blockId="146.[151,514,1535,1598]" box="[151,514,1535,1562]" pageId="146" pageNumber="147">
<heading id="BF63AF6651A918C4FF0EFA60ED3A1A5D" bold="true" box="[151,514,1535,1562]" fontSize="11" level="1" pageId="146" pageNumber="147" reason="1">
<taxonomicName id="2394638951A918C4FF0EFA60ED3A1A5D" ID-CoL="8NC7W" authority="Pimpasalee 2016" authorityName="Pimpasalee" authorityYear="2016" box="[151,514,1535,1562]" class="Insecta" family="Lampyridae" genus="Triangulara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851A918C4FF0EFA60ED3A1A5D" bold="true" box="[151,514,1535,1562]" pageId="146" pageNumber="147">
<emphasis id="D6E0C41851A918C4FF0EFA60EE121A5D" bold="true" box="[151,298,1535,1562]" italics="true" pageId="146" pageNumber="147">Triangulara</emphasis>
Pimpasalee 2016
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="AC8E4B8151A918C4FF0EF9BBEDC71AC7" pageId="146" pageNumber="147" type="description">
<paragraph id="E42B180A51A918C4FF0EF9BBEE2C1A79" blockId="146.[151,514,1535,1598]" box="[151,276,1572,1598]" pageId="146" pageNumber="147">
<figureCitation id="7CAF048F51A918C4FF0EF9BBEE2C1A79" box="[151,276,1572,1598]" captionStart="FIGURES 1721" captionStartId="13.[151,264,1607,1632]" captionTargetBox="[151,1435,174,1573]" captionTargetId="figure@13.[151,1435,174,1588]" captionTargetPageId="13" captionText="FIGURES 1721. Key to genera characters 3. 17, 18 Triangulara frontoflava; 19 Pygatyphella obsoleta; 20, 21 Pygat. huonensis. 17 ventral whole body; 18, 19, 21 ventral terminal abdomen; 20 right lateral terminal abdomen." figureDoi="http://doi.org/10.5281/zenodo.3508106" httpUri="https://zenodo.org/record/3508106/files/figure.png" pageId="146" pageNumber="147">Figs 17, 18</figureCitation>
</paragraph>
<paragraph id="E42B180A51A918C4FF0EF9F7EDC71AC7" blockId="146.[151,801,1639,1695]" box="[151,767,1639,1664]" pageId="146" pageNumber="147">
<taxonomicName id="2394638951A918C4FF0EF9F7EDC31AC7" authority="Pimpasalee 2016 in Ballantyne et al. 2016: 238" box="[151,763,1639,1664]" class="Insecta" family="Lampyridae" genus="Triangulara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851A918C4FF0EF9F7EE291AC7" box="[151,273,1640,1664]" italics="true" pageId="146" pageNumber="147">Triangulara</emphasis>
Pimpasalee
<quantity id="236CB5EF51A918C4FE0BF9F8EEE71AC7" box="[402,479,1639,1664]" metricMagnitude="1" metricUnit="m" metricValue="5.12064" pageId="146" pageNumber="147" unit="in" value="2016.0">2016 in</quantity>
<bibRefCitation id="800565FB51A918C4FE7CF9F8EDC31AC7" author="Ballantyne" box="[485,763,1639,1664]" etAl="et al." firstAuthor="Ballantyne" pageId="146" pageNumber="147" pagination="1 - 7" refId="ref114336" refString="Ballantyne, L. A &amp; Jusoh, W. F. A. (2016) List of genera and species in the Luciolinae from SE Asia and the Australopacific Version 2. Checklist of IndoPacific Luciolinae, 2016, 1 - 7. https: // doi. org / 10.13140 / RG. 2.1.4635.2241" type="journal article" year="2016">
Ballantyne
<emphasis id="D6E0C41851A918C4FDC0F9F6EDB41AC7" box="[601,652,1640,1664]" italics="true" pageId="146" pageNumber="147">et al.</emphasis>
2016: 238
</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151A918C4FF0EF918EC191AD8" box="[151,801,1670,1695]" pageId="146" pageNumber="147" type="type_taxon">
<paragraph id="E42B180A51A918C4FF0EF918EC191AD8" blockId="146.[151,801,1639,1695]" box="[151,801,1670,1695]" pageId="146" pageNumber="147">
<emphasis id="D6E0C41851A918C4FF0EF918EE1B1AD8" bold="true" box="[151,291,1670,1695]" pageId="146" pageNumber="147">
<typeStatus id="3B2FA6A851A918C4FF0EF918EFF41AD8" box="[151,204,1671,1695]" pageId="146" pageNumber="147">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="2394638951A918C4FEB3F918EDAC1AD8" ID-CoL="8N9FJ" authority="Pimpasalee" authorityName="Pimpasalee" box="[298,660,1670,1695]" class="Insecta" family="Lampyridae" genus="Triangulara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="147" phylum="Arthropoda" rank="species" species="frontoflava">
<emphasis id="D6E0C41851A918C4FEB3F918ED231AD8" box="[298,539,1671,1695]" italics="true" pageId="146" pageNumber="147">Triangulara frontoflava</emphasis>
Pimpasalee
</taxonomicName>
by monotypy
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151A918C2FF0EF953EC7B1DE8" lastPageId="148" lastPageNumber="149" pageId="146" pageNumber="147" type="diagnosis">
<paragraph id="E42B180A51A918C2FF0EF953EE671DCC" blockId="146.[151,1437,1740,2020]" lastBlockId="148.[151,1437,153,431]" lastPageId="148" lastPageNumber="149" pageId="146" pageNumber="147">
<emphasis id="D6E0C41851A918C4FF0EF953EE321AA0" bold="true" box="[151,266,1740,1767]" pageId="146" pageNumber="147">Diagnosis</emphasis>
.
<taxonomicName id="2394638951A918C4FE83F952EE991AA0" box="[282,417,1741,1767]" class="Insecta" family="Lampyridae" genus="Triangulara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851A918C4FE83F952EE991AA0" box="[282,417,1741,1767]" italics="true" pageId="146" pageNumber="147">Triangulara</emphasis>
</taxonomicName>
belongs to that group of S. E. Asian
<taxonomicName id="2394638951A918C4FCD5F952ECFC1AA0" authorityName="Lacordaire" authorityYear="1857" box="[844,964,1741,1767]" class="Insecta" family="Lampyridae" genus="Luciolinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="166" phylum="Arthropoda" rank="genus">Luciolinae</taxonomicName>
having pale yellowish dorsal surface (including a pale pronotum), with black tipped elytral apices. In the male the aedeagal LL are visible beside the ML when viewed from beneath in common with 17 other genera. Males are most obviously distinguished by the triangular outline of the LO in
<date id="902A3ECA51A918C4FE05F8A6EE861B14" box="[412,446,1849,1875]" pageId="146" pageNumber="147">V7</date>
and the separation of the LO from the sides and posterior margin of
<date id="902A3ECA51A918C4FB2AF8A6EBE01B14" box="[1203,1240,1849,1875]" pageId="146" pageNumber="147">V7</date>
; it is most similar to
<taxonomicName id="2394638951A918C4FF2CF8C2EEA31B30" authority="Ballantyne" authorityName="Ballantyne No" authorityYear="2016" box="[181,411,1885,1911]" class="Insecta" family="Lampyridae" genus="Sclerotia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851A918C4FF2CF8C2EE211B30" box="[181,281,1885,1911]" italics="true" pageId="146" pageNumber="147">Sclerotia</emphasis>
Ballantyne
</taxonomicName>
from which it is distinguished by the triangular outline of the LO in
<date id="902A3ECA51A918C4FB37F8C2EBEA1B30" box="[1198,1234,1885,1911]" pageId="146" pageNumber="147">V7</date>
, no anteromedian emargination of the
<date id="902A3ECA51A918C4FEECF81EEEAF1BDC" box="[373,407,1921,1947]" pageId="146" pageNumber="147">V7</date>
LO, and no sclerites surrounding the aedeagal sheath. (In
<taxonomicName id="2394638951A918C4FB80F81EEB451BDC" authorityName="Ballantyne No" authorityYear="2016" box="[1049,1149,1921,1947]" class="Insecta" family="Lampyridae" genus="Sclerotia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="146" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851A918C4FB80F81EEB451BDC" box="[1049,1149,1921,1947]" italics="true" pageId="146" pageNumber="147">Sclerotia</emphasis>
</taxonomicName>
the LO may occupy all of
<date id="902A3ECA51A918C4FF0EF83AEF841BF8" box="[151,188,1957,1983]" pageId="146" pageNumber="147">V7</date>
, or may appear heart shaped, with an anterior median emargination often so deep that the LO appears U shaped; a set of 3 sclerites surround the posterior area of the aedeagal sheath). It is distinguished from the similarly coloured
<taxonomicName id="2394638951AF18C2FF0EFF06EEAC1CF4" authority="Ballantyne" authorityName="Ballantyne Lambkin &amp; Fu" authorityYear="2013" box="[151,404,153,179]" class="Insecta" family="Lampyridae" genus="Abscondita" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851AF18C2FF0EFF06EE2D1CF4" box="[151,277,153,179]" italics="true" pageId="148" pageNumber="149">Abscondita</emphasis>
Ballantyne
</taxonomicName>
most obviously by the triangular shape of both the LO and
<date id="902A3ECA51AF18C2FBBDFF06EB711CF4" box="[1060,1097,153,179]" pageId="148" pageNumber="149">V7</date>
, the separation of the aedeagal LL along most of their dorsal length and the absence of bulbous projections from the lateral margins of the LL. (In
<taxonomicName id="2394638951AF18C2FF0EFF7EEE2D1CBC" authorityName="Ballantyne Lambkin &amp; Fu" authorityYear="2013" box="[151,277,225,251]" class="Insecta" family="Lampyridae" genus="Abscondita" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851AF18C2FF0EFF7EEE2D1CBC" box="[151,277,225,251]" italics="true" pageId="148" pageNumber="149">Abscondita</emphasis>
</taxonomicName>
the LO in
<date id="902A3ECA51AF18C2FE0AFF7EEE8D1CBC" box="[403,437,225,251]" pageId="148" pageNumber="149">V7</date>
occupies most if not all the area of
<date id="902A3ECA51AF18C2FCC9FF7EEC4A1CBC" box="[848,882,225,251]" pageId="148" pageNumber="149">V7</date>
which is broadly rounded behind and often very flat; the LL are fused along almost all of their dorsal length, and bulbous projections arise from the lateral margins of the LL). Confused in collections with
<taxonomicName id="2394638951AF18C2FDF4FEB6EC391D04" authorityName="Thancharoen" authorityYear="2007" box="[621,769,297,323]" class="Insecta" family="Lampyridae" genus="Sclerotia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="species" species="aquatilis">
<emphasis id="D6E0C41851AF18C2FDF4FEB6EC391D04" box="[621,769,297,323]" italics="true" pageId="148" pageNumber="149">Scl. aquatilis</emphasis>
</taxonomicName>
and
<taxonomicName id="2394638951AF18C2FCA2FEB6ECE51D04" box="[827,989,297,323]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="species" species="substriata">
<emphasis id="D6E0C41851AF18C2FCA2FEB6ECE51D04" box="[827,989,297,323]" italics="true" pageId="148" pageNumber="149">Scl. substriata</emphasis>
</taxonomicName>
. Female macropterous and coloured as for male except for pale terminal Vs 6 and 7. Larva unknown but preliminary observations by Pimpasalee indicate it may be aquatic.
</paragraph>
<caption id="B0EB488251A818C5FF0EF8F8EEFE1BAB" ID-DOI="http://doi.org/10.5281/zenodo.3508277" ID-Zenodo-Dep="3508277" httpUri="https://zenodo.org/record/3508277/files/figure.png" pageId="147" pageNumber="148" startId="147.[151,264,1895,1920]" targetBox="[213,1375,166,1875]" targetPageId="147">
<paragraph id="E42B180A51A818C5FF0EF8F8EEFE1BAB" blockId="147.[151,1436,1895,2028]" pageId="147" pageNumber="148">
<emphasis id="D6E0C41851A818C5FF0EF8F8EE491BC7" bold="true" box="[151,369,1895,1920]" pageId="147" pageNumber="148">FIGURES 463473.</emphasis>
<taxonomicName id="2394638951A818C5FEE1F8F7ED0E1B38" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[376,566,1895,1919]" class="Gammaproteobacteria" family="Enterobacteriaceae" genus="Serratia" kingdom="Bacteria" order="Enterobacteriales" pageId="147" pageNumber="148" phylum="Proteobacteria" rank="species" species="sibuyania" status="gen. et sp. nov.">
<emphasis id="D6E0C41851A818C5FEE1F8F7ED0E1B38" box="[376,566,1895,1919]" italics="true" pageId="147" pageNumber="148">Serratia sibuyania</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDD3796351A818C5FDA4F8F7EDE51BC7" box="[573,733,1896,1920]" pageId="147" pageNumber="148" rank="species">
<emphasis id="D6E0C41851A818C5FDA4F8F7ED521BC7" bold="true" box="[573,618,1896,1920]" pageId="147" pageNumber="148">gen.</emphasis>
<emphasis id="D6E0C41851A818C5FDE8F8F7EDBB1B38" box="[625,643,1896,1919]" italics="true" pageId="147" pageNumber="148">et</emphasis>
<emphasis id="D6E0C41851A818C5FD12F8F7EDE51BC7" bold="true" box="[651,733,1896,1920]" pageId="147" pageNumber="148">sp. nov.</emphasis>
</taxonomicNameLabel>
463 holotype (USNM); 464 Romblon (USNM); 465468, 470473 Luzon H.S.B. (USNM); 469 Macassar (USNM). 463, 464, 465 habitus dorsal; 466, 468 habitus ventral (468 after removal of abdomen); 467 dorsal head. Pronotum and anterior part of elytra; 470 tergite 8 ventral; 471473 abdomen: 471 intact, ventral; dissected, ventral; 473 dorsal.
</paragraph>
</caption>
<paragraph id="E42B180A51AF18C2FF5EFE0AEC7B1DE8" blockId="148.[151,1437,153,431]" box="[199,835,405,431]" pageId="148" pageNumber="149">
<bibRefCitation id="800565FB51AF18C2FF5EFE0AEE591DE8" author="Fu" box="[199,353,405,431]" firstAuthor="Fu" pageId="148" pageNumber="149" refId="ref116462" refString="Fu, X. H. (2014) An illustrated handbook of Chinese fireflies. The Commercial Press, Beijing, 167 pp. [in Chinese]" type="book" year="2014">Fu (2014: 41)</bibRefCitation>
illustrated a
<taxonomicName id="2394638951AF18C2FE6AFE0AED421DE8" box="[499,634,405,431]" class="Insecta" family="Lampyridae" genus="Triangulara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851AF18C2FE6AFE0AED421DE8" box="[499,634,405,431]" italics="true" pageId="148" pageNumber="149">Triangulara</emphasis>
</taxonomicName>
sp. as
<taxonomicName id="2394638951AF18C2FD5FFE0AEC241DE8" authorityName="in McDermott" authorityYear="1966" box="[710,796,405,431]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851AF18C2FD5FFE0AEC241DE8" box="[710,796,405,431]" italics="true" pageId="148" pageNumber="149">Luciola</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
</treatment>
</document>