treatments-xml/data/6C/3D/A9/6C3DA91C51641836FF0EF9E9EC881948.xml
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<document id="AD3FE92B3877D3DC8242B309783EDE8F" ID-CLB-Dataset="25190" ID-DOI="10.11646/zootaxa.4687.1.1" ID-GBIF-Dataset="de00ca6c-2843-4ae1-a39c-a29603947612" ID-ISSN="1175-5326" ID-Zenodo-Dep="3508098" ID-ZooBank="CE73264D-C234-4B82-A634-CAD6254C5957" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571388220410" checkinUser="plazi" docAuthor="Ho, - Z." docDate="2019" docId="6C3DA91C51641836FF0EF9E9EC881948" docLanguage="en" docName="zootaxa.4687.1.1.pdf.imf" docOrigin="Zootaxa 4687 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Luciola niah Ho 2019, sp. nov." docType="treatment" docVersion="20" lastPageNumber="97" masterDocId="9004D164513B1856FF99FF9FEF381C47" masterDocTitle="The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species" masterLastPageNumber="174" masterPageNumber="1" pageNumber="96" updateTime="1698760145766" updateUser="ExternalLinkService">
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<mods:title id="AB3DE25072D844F687B552D315E50929">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="E591916063BDB466FF49A26AC57F9D4A">Ho, - Z.</mods:namePart>
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<mods:title id="05A332E18D7A9178391A70E72C3F5D3E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6B46579EC4199F0DFD68398905939430">
<mods:date id="1A1E59650B6A710005764671170A63BA">2019</mods:date>
<mods:detail id="450799FED520C91EBFC155E02F315865" type="pubDate">
<mods:number id="77B9A1BD8FB75D3203ABF566FDF84D0A">2019-10-18</mods:number>
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<mods:detail id="2B7D4195107E1CDB0FEC81E45852109A" type="volume">
<mods:number id="B2EB7BB4373F5504019CB1B1A6F57092">4687</mods:number>
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<mods:classification id="AF9C5CCD77CB1443312F09E3C35F2195">journal article</mods:classification>
<mods:identifier id="F82B365DF5DFC4BB3FFBFEED30FB904E" type="CLB-Dataset">25190</mods:identifier>
<mods:identifier id="9B1A642260495E5969291FE00445A943" type="DOI">10.11646/zootaxa.4687.1.1</mods:identifier>
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</mods:mods>
<treatment id="6C3DA91C51641836FF0EF9E9EC881948" ID-DOI="http://doi.org/10.5281/zenodo.4688885" ID-GBIF-Taxon="159757381" ID-Zenodo-Dep="4688885" LSID="urn:lsid:plazi:treatment:6C3DA91C51641836FF0EF9E9EC881948" httpUri="http://treatment.plazi.org/id/6C3DA91C51641836FF0EF9E9EC881948" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">
<subSubSection id="AC8E4B8151641809FF0EF9E9EE081AF3" pageId="95" pageNumber="96" type="nomenclature">
<paragraph id="E42B180A51641809FF0EF9E9EED01AD6" blockId="95.[151,488,1654,1716]" box="[151,488,1654,1681]" pageId="95" pageNumber="96">
<heading id="BF63AF6651641809FF0EF9E9EED01AD6" bold="true" box="[151,488,1654,1681]" fontSize="11" level="1" pageId="95" pageNumber="96" reason="1">
<emphasis id="D6E0C41851641809FF0EF9E9EED01AD6" bold="true" box="[151,488,1654,1681]" pageId="95" pageNumber="96">
<taxonomicName id="2394638951641809FF0EF9E9EEBB1AD6" ID-CoL="8NC2B" authority="Jusoh" authorityName="Ho" authorityYear="2019" box="[151,387,1654,1681]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="niah" status="sp. nov.">
<emphasis id="D6E0C41851641809FF0EF9E9EE0A1AD6" bold="true" box="[151,306,1654,1681]" italics="true" pageId="95" pageNumber="96">Luciola niah</emphasis>
Jusoh
</taxonomicName>
<taxonomicNameLabel id="CDD3796351641809FE12F9E8EED01AD6" box="[395,488,1655,1681]" pageId="95" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E42B180A51641809FF0EF905EE081AF3" blockId="95.[151,488,1654,1716]" box="[151,304,1690,1716]" pageId="95" pageNumber="96">Figs 284291</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151641809FF0EF940EA7B1B36" pageId="95" pageNumber="96" type="materials_examined">
<paragraph id="E42B180A51641809FF0EF940EE861B52" blockId="95.[151,1437,1758,1905]" pageId="95" pageNumber="96">
<emphasis id="D6E0C41851641809FF0EF940EFEA1AB0" bold="true" box="[151,210,1759,1783]" pageId="95" pageNumber="96">Type.</emphasis>
<materialsCitation id="54FC125751641809FF41F941EE861B52" ID-GBIF-Occurrence="2427485649" collectingDate="2010-09-24" collectionCode="FRIM" country="Malaysia" location="Nazeri Abghani" municipality="Lambir Hill" pageId="95" pageNumber="96" specimenCode="MYFI018-14" specimenCount="1" stateProvince="Sarawak" typeStatus="holotype">
<typeStatus id="3B2FA6A851641809FF41F941EE0E1AB0" box="[216,310,1758,1783]" pageId="95" pageNumber="96" type="holotype">Holotype</typeStatus>
male.
<emphasis id="D6E0C41851641809FEE3F941ED3F1AB0" bold="true" box="[378,519,1758,1783]" pageId="95" pageNumber="96">
<collectingCountry id="9C83589A51641809FEE3F941ED3C1AB0" box="[378,516,1758,1783]" name="Malaysia" pageId="95" pageNumber="96">MALAYSIA</collectingCountry>
.
</emphasis>
<emphasis id="D6E0C41851641809FD94F941ED5E1AB1" box="[525,614,1758,1782]" italics="true" pageId="95" pageNumber="96">
<collectingRegion id="2650D6E851641809FD94F941ED5E1AB1" box="[525,614,1758,1782]" country="Malaysia" name="Sarawak" pageId="95" pageNumber="96">
<collectingCountry id="9C83589A51641809FD94F941ED5E1AB1" box="[525,614,1758,1782]" name="Malaysia" pageId="95" pageNumber="96">Sarawak</collectingCountry>
</collectingRegion>
</emphasis>
<collectingMunicipality id="044F827051641809FDF5F941EDDB1AB0" box="[620,739,1758,1783]" pageId="95" pageNumber="96">Lambir Hill</collectingMunicipality>
<date id="902A3ECA51641809FD70F941EC601AB0" box="[745,856,1758,1783]" pageId="95" pageNumber="96" value="2010-09-24">
<collectingDate id="806EC72251641809FD70F941EC601AB0" box="[745,856,1758,1783]" pageId="95" pageNumber="96" value="2010-09-24">24.ix.2010</collectingDate>
</date>
, 19.24,
<location id="E14B4ED151641809FC3EF941EB7E1AB0" LSID="urn:lsid:plazi:treatment:6C3DA91C51641836FF0EF9E9EC881948:E14B4ED151641809FC3EF941EB7E1AB0" box="[935,1094,1758,1783]" country="Malaysia" municipality="Lambir Hill" name="Nazeri Abghani" pageId="95" pageNumber="96" stateProvince="Sarawak">Nazeri Abghani</location>
(
<collectionCode id="828580CF51641809FBCAF940EB4C1AB0" box="[1107,1140,1759,1783]" pageId="95" pageNumber="96">SK</collectionCode>
0017 |
<specimenCode id="B432B07151641809FB21F940EA791AB0" box="[1208,1345,1759,1783]" collectionCode="MYFI" pageId="95" pageNumber="96">
<collectionCode id="828580CF51641809FB21F940EBC31AB0" box="[1208,1275,1759,1783]" collectionName="MYFI" pageId="95" pageNumber="96">MYFI</collectionCode>
018-14
</specimenCode>
) (
<collectionCode id="828580CF51641809FACDF940EAAD1AB0" box="[1364,1429,1759,1783]" collectionName="Malaysia, Kuala Lumpur, Forest Research Institute" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="95" pageNumber="96">FRIM</collectionCode>
) Figs 284, 285, 288291.
</materialsCitation>
</paragraph>
<paragraph id="E42B180A51641809FF0EF883EA7B1B36" blockId="95.[151,1437,1758,1905]" pageId="95" pageNumber="96">
<materialsCitation id="54FC125751641809FF0EF883EA071B36" ID-GBIF-Occurrence="2427485653" collectingDate="2010-07-02" collectingDateMax="2011-09-02" collectingDateMin="2010-07-02" collectionCode="ANIC, FRIM" collectorName="I Polunin &amp; M. Musbah" country="Malaysia" location="Niah National Park forest" municipality="Kapit" pageId="95" pageNumber="96" specimenCount="6" specimenCount-male="6" stateProvince="Sarawak" typeStatus="paratype">
<emphasis id="D6E0C41851641809FF0EF883EE981B73" bold="true" box="[151,416,1819,1844]" pageId="95" pageNumber="96">
<typeStatus id="3B2FA6A851641809FF0EF883EE3D1B73" box="[151,261,1820,1844]" pageId="95" pageNumber="96" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="9C83589A51641809FE8BF884EEA41B73" box="[274,412,1819,1844]" name="Malaysia" pageId="95" pageNumber="96">MALAYSIA</collectingCountry>
.
</emphasis>
<emphasis id="D6E0C41851641809FE30F883ED3A1B73" box="[425,514,1820,1844]" italics="true" pageId="95" pageNumber="96">
<collectingRegion id="2650D6E851641809FE30F883ED3A1B73" box="[425,514,1820,1844]" country="Malaysia" name="Sarawak" pageId="95" pageNumber="96">
<collectingCountry id="9C83589A51641809FE30F883ED3A1B73" box="[425,514,1820,1844]" name="Malaysia" pageId="95" pageNumber="96">Sarawak</collectingCountry>
</collectingRegion>
</emphasis>
:
<collectingMunicipality id="044F827051641809FD88F884ED741B73" box="[529,588,1819,1844]" pageId="95" pageNumber="96">Kapit</collectingMunicipality>
,
<collectorName id="49617DDC51641809FDC1F883ED821B73" box="[600,698,1819,1844]" pageId="95" pageNumber="96">I Polunin</collectorName>
, no date
<specimenCount id="F292D38351641809FC82F883EC531B73" box="[795,875,1819,1844]" pageId="95" pageNumber="96" type="male">2 males</specimenCount>
(
<collectionCode id="828580CF51641809FCE2F883EC871B73" box="[891,959,1820,1844]" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="95" pageNumber="96">ANIC</collectionCode>
).
<location id="E14B4ED151641809FC48F884EBE41B73" LSID="urn:lsid:plazi:treatment:6C3DA91C51641836FF0EF9E9EC881948:E14B4ED151641809FC48F884EBE41B73" box="[977,1244,1819,1844]" country="Malaysia" municipality="Kapit" name="Niah National Park forest" pageId="95" pageNumber="96" stateProvince="Sarawak">Niah National Park forest</location>
track,
<collectorName id="49617DDC51641809FABCF883EAA51B73" box="[1317,1437,1819,1844]" pageId="95" pageNumber="96">M. Musbah</collectorName>
and colleagues,
<date id="902A3ECA51641809FEF4F8A5EEEE1B14" box="[365,470,1850,1875]" pageId="95" pageNumber="96" value="2010-07-02">
<collectingDate id="806EC72251641809FEF4F8A5EEEE1B14" box="[365,470,1850,1875]" pageId="95" pageNumber="96" value="2010-07-02">2.vii.2010</collectingDate>
</date>
,
<specimenCount id="F292D38351641809FE7BF8A4ED0A1B14" box="[482,562,1850,1875]" pageId="95" pageNumber="96" type="male">2 males</specimenCount>
(
<collectionCode id="828580CF51641809FDD8F8A4ED5A1B14" box="[577,610,1851,1875]" pageId="95" pageNumber="96">SK</collectionCode>
0023 | GenBank:
<collectionCode id="828580CF51641809FC80F8A5EC061B14" box="[793,830,1850,1875]" pageId="95" pageNumber="96">KY</collectionCode>
572917.1,
<collectionCode id="828580CF51641809FC32F8A4ECE91B14" box="[939,977,1851,1875]" pageId="95" pageNumber="96">MF</collectionCode>
948239.1;
<collectionCode id="828580CF51641809FBA4F8A5EB661B14" box="[1085,1118,1850,1875]" pageId="95" pageNumber="96">SK</collectionCode>
0064). Long Aton, Ulu Baram,
<date id="902A3ECA51641809FF5EF8C7EE101B36" box="[199,296,1880,1905]" pageId="95" pageNumber="96" value="2011-09-02">
<collectingDate id="806EC72251641809FF5EF8C7EE101B36" box="[199,296,1880,1905]" pageId="95" pageNumber="96" value="2011-09-02">2.ix.2011</collectingDate>
</date>
, Musa Musbah
<specimenCount id="F292D38351641809FE50F8C6ED2F1B36" box="[457,535,1880,1905]" pageId="95" pageNumber="96" type="male">2 males</specimenCount>
(
<collectionCode id="828580CF51641809FDBCF8C7ED7E1B36" box="[549,582,1880,1905]" pageId="95" pageNumber="96">SK</collectionCode>
0067 | GenBank:
<collectionCode id="828580CF51641809FD61F8C7EC261B36" box="[760,798,1880,1905]" pageId="95" pageNumber="96">MF</collectionCode>
063274.1;
<collectionCode id="828580CF51641809FC10F8C7EC911B36" box="[905,937,1880,1905]" pageId="95" pageNumber="96">SK</collectionCode>
0069 | GenBank:
<collectionCode id="828580CF51641809FBC2F8C7EBB81B36" box="[1115,1152,1880,1905]" pageId="95" pageNumber="96">MF</collectionCode>
063275.1) (
<collectionCode id="828580CF51641809FB6DF8C6EA001B36" box="[1268,1336,1881,1905]" collectionName="Malaysia, Kuala Lumpur, Forest Research Institute" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="95" pageNumber="96">FRIM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151641836FF0EF800EB911C90" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96" type="diagnosis">
<paragraph id="E42B180A51641836FF0EF800EB911C90" blockId="95.[151,1437,1951,2014]" lastBlockId="96.[151,1437,153,1295]" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">
<emphasis id="D6E0C41851641809FF0EF800EE2A1BFD" bold="true" box="[151,274,1951,1978]" pageId="95" pageNumber="96">Diagnosis.</emphasis>
4.86.1 mm long; one of only two
<taxonomicName id="2394638951641809FD36F83FEC3D1BFD" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[687,773,1952,1978]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="D6E0C41851641809FD36F83FEC3D1BFD" box="[687,773,1952,1978]" italics="true" pageId="95" pageNumber="96">Luciola</emphasis>
</taxonomicName>
<emphasis id="D6E0C41851641809FC97F83FEC761BFD" bold="true" box="[782,846,1952,1978]" pageId="95" pageNumber="96">
<taxonomicNameLabel id="CDD3796351641809FC97F83FEC761BFD" box="[782,846,1952,1978]" pageId="95" pageNumber="96" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
having dorsal colouration of yellowish brown with black elytral apices; other
<taxonomicName id="2394638951641809FE20F85BED371B99" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[441,527,1988,2014]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="D6E0C41851641809FE20F85BED371B99" box="[441,527,1988,2014]" italics="true" pageId="95" pageNumber="96">Luciola</emphasis>
</taxonomicName>
<emphasis id="D6E0C41851641809FD8DF85BED691B99" bold="true" box="[532,593,1988,2014]" pageId="95" pageNumber="96">
<taxonomicNameLabel id="CDD3796351641809FD8DF85BED691B99" box="[532,593,1988,2014]" pageId="95" pageNumber="96" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
species with pale yellowish or pale brown pronota have dark brown to black elytra. Distinguished from the other similarly coloured species,
<taxonomicName id="23946389515B1836FCC2FF05EC861CF4" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[859,958,154,179]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="jengai" status="sp. nov.">
<emphasis id="D6E0C418515B1836FCC2FF05EC861CF4" box="[859,958,154,179]" italics="true" pageId="96" pageNumber="97">L. jengai</emphasis>
</taxonomicName>
<emphasis id="D6E0C418515B1836FC5DFF06EB251CF4" bold="true" box="[964,1053,153,179]" pageId="96" pageNumber="97">
<taxonomicNameLabel id="CDD37963515B1836FC5DFF06EB251CF4" box="[964,1053,153,179]" pageId="96" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, most obviously by the abdominal colour in
<taxonomicName id="23946389515B1836FE9BFF21EE701C90" authorityName="Ho" authorityYear="2019" box="[258,328,190,215]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="jengai">
<emphasis id="D6E0C418515B1836FE9BFF21EE701C90" box="[258,328,190,215]" italics="true" pageId="96" pageNumber="97">jengai</emphasis>
</taxonomicName>
, and which also does not have leaf like lobes on the inner margins of the LL.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B81515B1836FF5EFF7FED591F8C" pageId="96" pageNumber="97" type="description">
<paragraph id="E42B180A515B1836FF5EFF7FEECA1F18" blockId="96.[151,1437,153,1295]" pageId="96" pageNumber="97">
<emphasis id="D6E0C418515B1836FF5EFF7FEE8D1CBC" bold="true" box="[199,437,224,251]" pageId="96" pageNumber="97">Description of male.</emphasis>
4.86.1 mm long. Colour (Figs 284, 285): (Kapit males preserved in 70% ethanol differ in colour to remainder and are described here in brackets); pronotum yellowish with diffuse median darker markings (retraction of underlying fat body gives impression of darker markings); MN, MS and elytra yellowish brown with elytral apices black (elytra pale yellow with apices quite pale brown); elytra semitransparent and underlying hind wings may confuse interpretation of colour; head black, antennae and palpi dark brown (head reddish brown, antennae and palpi light brown); venter of thorax, basal abdominal ventrites and all abdominal tergites yellowish brown except for narrow dark posterior margin of
<date id="902A3ECA515B1836FD1DFE27ED9E1D94" box="[644,678,440,467]" pageId="96" pageNumber="97">V5</date>
(all areas yellow except for thin dark marking on posterior margin of
<date id="902A3ECA515B1836FF2FFE43EFE41DB0" box="[182,220,476,503]" pageId="96" pageNumber="97">V5</date>
); legs yellowish brown with tibiae and tarsi dark brown (legs yellow, tarsi light brown); LO creamy white. Pronotum: slightly wider across middle (B&gt; A, C); width a little less than humeral width; anterolateral corners angulate obtuse, posterolateral corners angulate. Elytron: subparallel sided. Head: minimal depression between eyes; GHW
<date id="902A3ECA515B1836FF78FDD6EE201E24" box="[225,280,585,611]" pageId="96" pageNumber="97" value="2011-10">11 X</date>
SIW; ASD much less than ASW (sockets are almost contiguous). Antennae: longer than GHW but not as long as
<date id="902A3ECA515B1836FE96FDF2EE011EC0" box="[271,313,621,647]" pageId="96" pageNumber="97">2 X</date>
GHW; segments elongate slender, scape longer than FS. Mouthparts: apical labial palpomere laterally flattened with inner edge dentate. Abdomen (Fig. 285): LO occupying entire area of
<date id="902A3ECA515B1836FB12FD0EEB971EEC" box="[1163,1199,657,683]" pageId="96" pageNumber="97">V6</date>
, 7;
<date id="902A3ECA515B1836FB43FD0EEBC41EEC" box="[1242,1276,657,683]" pageId="96" pageNumber="97">V7</date>
a little longer than wide, posterior margin rounded and no MPP; T8 with anterolateral arms elongate slender slightly longer than entire posterior portion which is rounded and without median emargination. Aedeagal sheath (Figs 288, 289): apical margin of sheath sternite narrowly prolonged and medianly slightly emarginated and hairy; median anterior margin of sheath tergite strongly prolonged and apically pointed.Aedeagus (Figs 290, 291): apices of LL narrow; LL longer than ML and widely divergent.
</paragraph>
<paragraph id="E42B180A515B1836FF5EFCF6ED591F8C" blockId="96.[151,1437,153,1295]" pageId="96" pageNumber="97">
<emphasis id="D6E0C418515B1836FF5EFCF6EE261FC4" bold="true" box="[199,286,873,899]" pageId="96" pageNumber="97">Female</emphasis>
: Jusoh associated the female by DNA barcode (Process ID: MYFI179-14 | GenBank: KY572915.1). Macropterous and capable of flight. Coloured as for male except for white LO restricted to
<date id="902A3ECA515B1836FB20FC12EBE31FE0" box="[1209,1243,909,935]" pageId="96" pageNumber="97">V6</date>
(Figs 286, 287). Bursa with two wide plates on each side.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B81515B1836FF5EFC4AECFA1FA8" box="[199,962,981,1007]" pageId="96" pageNumber="97" type="etymology">
<paragraph id="E42B180A515B1836FF5EFC4AECFA1FA8" blockId="96.[151,1437,153,1295]" box="[199,962,981,1007]" pageId="96" pageNumber="97">
<emphasis id="D6E0C418515B1836FF5EFC4AEE701FA8" bold="true" box="[199,328,981,1007]" pageId="96" pageNumber="97">Etymology</emphasis>
. The locality name Niah is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B81515B1836FF5EFC66EC881948" pageId="96" pageNumber="97" type="discussion">
<paragraph id="E42B180A515B1836FF5EFC66EDC018E4" blockId="96.[151,1437,153,1295]" pageId="96" pageNumber="97">
<emphasis id="D6E0C418515B1836FF5EFC66EE0C1854" bold="true" box="[199,308,1017,1043]" pageId="96" pageNumber="97">Remarks</emphasis>
.
<bibRefCitation id="800565FB515B1836FEDDFC66ED711854" author="Ballantyne" box="[324,585,1017,1043]" etAl="et al." firstAuthor="Ballantyne" pageId="96" pageNumber="97" pagination="1 - 7" refId="ref114336" refString="Ballantyne, L. A &amp; Jusoh, W. F. A. (2016) List of genera and species in the Luciolinae from SE Asia and the Australopacific Version 2. Checklist of IndoPacific Luciolinae, 2016, 1 - 7. https: // doi. org / 10.13140 / RG. 2.1.4635.2241" type="journal article" year="2016">
Ballantyne
<emphasis id="D6E0C418515B1836FE5CFC65EEC71854" box="[453,511,1017,1043]" italics="true" pageId="96" pageNumber="97">et al.</emphasis>
(2016
</bibRefCitation>
<tableCitation id="A9162DB1515B1836FDC8FC66ED9E1853" box="[593,678,1017,1044]" captionStart="TABLE 6" captionStartId="47.[151,239,153,177]" captionTargetBox="[166,1409,225,1016]" captionText="TABLE 6. List of species of Asymmetricata Ballantyne 2009 (*References: 1 = Ballantyne &amp; Lambkin 2009; 2 = Fu et al. 2012b; 3 = Yiu 2017; 4 = Fu 2014.)" httpUri="http://table.plazi.org/id/B0EB488251141879FF0EFF06EB981C91" pageId="96" pageNumber="97" tableUuid="B0EB488251141879FF0EFF06EB981C91">Table 6</tableCitation>
) listed 30 species with this colour pattern (dorsally pale yellowish or pale brown with or without dark elytral apices) that is common among S. Asian
<taxonomicName id="23946389515B1836FBA7FB82EB811870" authorityName="Lacordaire" authorityYear="1857" box="[1086,1209,1053,1079]" class="Insecta" family="Lampyridae" genus="Luciolinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="166" phylum="Arthropoda" rank="genus">Luciolinae</taxonomicName>
. Many of these species have been assigned to
<emphasis id="D6E0C418515B1836FE5EFBDDED43181C" box="[455,635,1090,1115]" italics="true" pageId="96" pageNumber="97">species incertae</emphasis>
here. Of the remainder the only species recorded from
<collectingCountry id="9C83589A515B1836FB71FBDEEA70181C" box="[1256,1352,1089,1115]" name="Malaysia" pageId="96" pageNumber="97">Sarawak</collectingCountry>
having similar colouration is
<taxonomicName id="23946389515B1836FE17FBF9EDA21838" authority="Olivier" authorityName="Olivier" box="[398,666,1125,1151]" class="Insecta" family="Lampyridae" genus="Luciolinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="D6E0C418515B1836FE17FBF9ED791838" box="[398,577,1125,1151]" italics="true" pageId="96" pageNumber="97">L. maculipennis</emphasis>
Olivier
</taxonomicName>
, where the description of the terminal abdominal ventrite suggests a prolongation that could be typical of a
<taxonomicName id="23946389515B1836FDD3FB16EDE918E4" authorityName="Wittmer" authorityYear="1939" box="[586,721,1161,1187]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418515B1836FDD3FB16EDE918E4" box="[586,721,1161,1187]" italics="true" pageId="96" pageNumber="97">Pygoluciola</emphasis>
</taxonomicName>
sp.
</paragraph>
<paragraph id="E42B180A515B1836FF5EFB32EC881948" blockId="96.[151,1437,153,1295]" pageId="96" pageNumber="97">
We have presently little information about shape and numbers of bursa plates in this genus. Paired plates as seen here are characteristic of other non-related genera like
<taxonomicName id="23946389515B1836FCB7FB4DECA218AC" authorityName="Ho" authorityYear="2019" box="[814,922,1234,1259]" class="Insecta" family="Lampyridae" genus="Pteroptyx" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418515B1836FCB7FB4DECA218AC" box="[814,922,1234,1259]" italics="true" pageId="96" pageNumber="97">Pteroptyx</emphasis>
</taxonomicName>
and
<taxonomicName id="23946389515B1836FC56FB4EEB7518AC" authorityName="Dejean" authorityYear="1833" box="[975,1101,1233,1259]" class="Insecta" family="Lampyridae" genus="Colophotia" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418515B1836FC56FB4EEB7518AC" box="[975,1101,1233,1259]" italics="true" pageId="96" pageNumber="97">Colophotia</emphasis>
</taxonomicName>
.
<taxonomicName id="23946389515B1836FBC3FB4EEBC618AC" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1114,1278,1233,1259]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="italica">
<emphasis id="D6E0C418515B1836FBC3FB4EEBC618AC" box="[1114,1278,1233,1259]" italics="true" pageId="96" pageNumber="97">Luciola italica</emphasis>
</taxonomicName>
does not have bursa plates.
<bibRefCitation id="800565FB515B1836FEB3FB6AEEC31948" author="Jusoh" box="[298,507,1269,1295]" etAl="et al." firstAuthor="Jusoh" pageId="96" pageNumber="97" pagination="1 - 71" refId="ref118381" refString="Jusoh, W. F. A., Ballantyne, L., Lambkin, C. L., Hashim, N. R. &amp; Wahlberg, N. (2018) The firefly genus Pteroptyx Olivier revisited (Coleoptera: Lampyridae: Luciolinae). Zootaxa, 4456 (1), 1 - 71. https: // doi. org / 10.11646 / zootaxa. 4456.1.1" type="journal article" year="2018">
Jusoh
<emphasis id="D6E0C418515B1836FEE8FB69EE911948" box="[369,425,1269,1295]" italics="true" pageId="96" pageNumber="97">et al.</emphasis>
(2018)
</bibRefCitation>
scored this species as “
<taxonomicName id="23946389515B1836FC98FB6AEC611948" box="[769,857,1269,1295]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">Luciola</taxonomicName>
WFA”.
</paragraph>
</subSubSection>
</treatment>
</document>