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<mods:title id="04AB3B7E668CD714CC61562EDE0F1713">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<taxonomicName id="239463895161180CFF0EFB82ED221870" ID-CoL="3WCQY" authority="Olivier 1913 c" authorityName="Olivier" authorityYear="1913" box="[151,538,1053,1080]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="D6E0C4185161180CFF0EFB82ED221870" bold="true" box="[151,538,1053,1080]" pageId="90" pageNumber="91">
<emphasis id="D6E0C4185161180CFF0EFB82EE53187F" bold="true" box="[151,363,1053,1080]" italics="true" pageId="90" pageNumber="91">Luciola filiformis</emphasis>
<bibRefCitation id="800565FB5161180CFEEAFB82ED221870" author="Olivier" box="[371,538,1053,1079]" firstAuthor="Olivier" pageId="90" pageNumber="91" pagination="269 - 272" refId="ref121053" refString="Olivier, E. (1913 c) H. Sauter's Formosa-Ausbeute. Lampyridae (Col.). Entomologische Mittellungen, II (9), 269 - 272." type="journal article" year="1913" yearSuffix="c">Olivier 1913c</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="E42B180A5161180CFF0EFBFEEE4318DF" blockId="90.[151,1437,1120,1238]" pageId="90" pageNumber="91">
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<taxonomicName id="239463895161180CFF0EFBFEED28183E" ID-CoL="3WCQY" authority="Olivier 1913 c: 271" authorityName="Olivier" authorityPageNumber="271" authorityYear="1913" box="[151,528,1120,1145]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="D6E0C4185161180CFF0EFBFEEE70183E" box="[151,328,1121,1145]" italics="true" pageId="90" pageNumber="91">Luciola filiformis</emphasis>
<treatmentCitation id="65353E1B5161180CFED7FBFFED28183E" author="Olivier, E." box="[334,528,1120,1145]" page="271" pageId="90" pageNumber="91" year="1913">
<bibRefCitation id="800565FB5161180CFED7FBFFED28183E" author="Olivier" box="[334,528,1120,1145]" firstAuthor="Olivier" pageId="90" pageNumber="91" pagination="269 - 272" refId="ref121053" refString="Olivier, E. (1913 c) H. Sauter's Formosa-Ausbeute. Lampyridae (Col.). Entomologische Mittellungen, II (9), 269 - 272." type="journal article" year="1913" yearSuffix="c">Olivier 1913c: 271</bibRefCitation>
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. Chȗjô &amp; Sâtô 1971: 15.
<treatmentCitation id="65353E1B5161180CFC88FBFFEC8E183E" author="Chen, T. R." box="[785,950,1120,1145]" page="170" pageId="90" pageNumber="91" year="2003">
<bibRefCitation id="800565FB5161180CFC88FBFFEC8E183E" author="Chen" box="[785,950,1120,1145]" firstAuthor="Chen" pageId="90" pageNumber="91" refId="ref115752" refString="Chen, T. R. (2003) The fireflies of Taiwan. Field Image Publisher, Taipei, 256 pp. [in Chinese]" type="book" year="2003">Chen 2003: 170</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="65353E1B5161180CFC26FBFFEBE7183E" author="Kawashima, I. &amp; Suzuki, H. &amp; Sato, M." box="[959,1247,1120,1145]" page="247" pageId="90" pageNumber="91" year="2003">
<bibRefCitation id="800565FB5161180CFC26FBFFEBE7183E" author="Kawashima" box="[959,1247,1120,1145]" etAl="et al." firstAuthor="Kawashima" pageId="90" pageNumber="91" pagination="241 - 261" refId="ref118611" refString="Kawashima, I., Suzuki, H. &amp; Sato, M. (2003) A Check-List of Japanese Fireflies (Coleoptera, Lampyridae and Rhagophthalmidae). Japanese Journal of Systematic Entomology, 9 (2), 241 - 261." type="journal article" year="2003">
Kawashima
<emphasis id="D6E0C4185161180CFBA4FBFDEB48183E" box="[1085,1136,1121,1145]" italics="true" pageId="90" pageNumber="91">et al.</emphasis>
2003: 247
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="65353E1B5161180CFB71FBFDEFC818DF" author="Jeng, M. L. &amp; Yang, P. S. &amp; Lai, J." page="260" pageId="90" pageNumber="91" year="2003">
<bibRefCitation id="800565FB5161180CFB71FBFDEFC818DF" author="Jeng" etAl="et al." firstAuthor="Jeng" pageId="90" pageNumber="91" pagination="247 - 262" refId="ref118190" refString="Jeng, M. L., Yang, P. S. &amp; Lai, J. (2003 b) Notes on the genus Luciola (Coleoptera, Lampyridae, Luciolinae) of Taiwan. Special Bulletin of the Japanese Society of Coleopterolog y, 6, 247 - 262." type="journal article" year="2003" yearSuffix="b">
Jeng
<emphasis id="D6E0C4185161180CFA84FBFDEA72183E" box="[1309,1354,1121,1145]" italics="true" pageId="90" pageNumber="91">et al</emphasis>
. 2003b: 260
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="65353E1B5161180CFF63FB1FEE4F18DF" author="Fu, X. H." box="[250,375,1152,1176]" page="23" pageId="90" pageNumber="91" year="2014">
<bibRefCitation id="800565FB5161180CFF63FB1FEE4F18DF" author="Fu" box="[250,375,1152,1176]" firstAuthor="Fu" pageId="90" pageNumber="91" refId="ref116462" refString="Fu, X. H. (2014) An illustrated handbook of Chinese fireflies. The Commercial Press, Beijing, 167 pp. [in Chinese]" type="book" year="2014">Fu 2014: 23</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="E42B180A5161180CFF0EFB01ECFB18F0" blockId="90.[151,1437,1120,1238]" box="[151,963,1182,1207]" pageId="90" pageNumber="91">
<treatmentCitationGroup id="C4843F245161180CFF0EFB01ECFB18F0" box="[151,963,1182,1207]" pageId="90" pageNumber="91">
<taxonomicName id="239463895161180CFF0EFB01ED7E18F0" authority="Matsumura 1918: 39" authorityName="Matsumura" authorityPageNumber="39" authorityYear="1918" box="[151,582,1182,1207]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="yayeyamana">
<emphasis id="D6E0C4185161180CFF0EFB01EE5218F1" box="[151,362,1182,1206]" italics="true" pageId="90" pageNumber="91">Luciola yayeyamana</emphasis>
<treatmentCitation id="65353E1B5161180CFEE9FB00ED7E18F0" author="Matsumura, M." box="[368,582,1183,1207]" page="39" pageId="90" pageNumber="91" year="1918">
<bibRefCitation id="800565FB5161180CFEE9FB00ED7E18F0" author="Matsumura" box="[368,582,1183,1207]" firstAuthor="Matsumura" pageId="90" pageNumber="91" pagination="82 - 89" refId="ref119223" refString="Matsumura, M. (1918) The fireflies of Japan. Kyoiku-Gaho (Education Illustration Post), 6 (3), 82 - 89. [in Japanese]" type="journal article" year="1918">Matsumura 1918: 39</bibRefCitation>
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</taxonomicName>
. Chȗjô &amp; Sâtô 1971: 15 (synonymy).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B815161180CFF0EFB22EB5F1892" box="[151,1127,1212,1238]" pageId="90" pageNumber="91" type="materials_examined">
<paragraph id="E42B180A5161180CFF0EFB22EB5F1892" blockId="90.[151,1437,1120,1238]" box="[151,1127,1212,1238]" pageId="90" pageNumber="91">
<emphasis id="D6E0C4185161180CFF0EFB22EFEA1892" bold="true" box="[151,210,1213,1237]" pageId="90" pageNumber="91">Type.</emphasis>
<materialsCitation id="54FC12575161180CFF40FB22EB5B1892" ID-GBIF-Occurrence="2427485634" box="[217,1123,1212,1238]" collectionCode="MNHN" country="Taiwan" location="Examined" municipality="Jeng" pageId="90" pageNumber="91" specimenCount="1" typeStatus="syntype">
<typeStatus id="3B2FA6A85161180CFF40FB22EE141892" box="[217,300,1213,1237]" pageId="90" pageNumber="91" type="syntype">Syntype</typeStatus>
male.
<emphasis id="D6E0C4185161180CFEE9FB23EEEE1892" bold="true" box="[368,470,1212,1237]" pageId="90" pageNumber="91">
<collectingCountry id="9C83589A5161180CFEE9FB23EEEE1892" box="[368,470,1212,1237]" name="Taiwan" pageId="90" pageNumber="91">TAIWAN</collectingCountry>
</emphasis>
(as
<collectingCountry id="9C83589A5161180CFD99FB22ED651892" box="[512,605,1213,1237]" name="Taiwan" pageId="90" pageNumber="91">Formosa</collectingCountry>
).
<location id="E14B4ED15161180CFDF7FB23EDED1892" LSID="urn:lsid:plazi:treatment:6C3DA91C5161180CFF0EFB82EA2A1A25:E14B4ED15161180CFDF7FB23EDED1892" box="[622,725,1212,1237]" country="Taiwan" municipality="Jeng" name="Examined" pageId="90" pageNumber="91">Examined</location>
by M-L
<collectingMunicipality id="044F82705161180CFCB7FB21EC641891" box="[814,860,1214,1238]" pageId="90" pageNumber="91">Jeng</collectingMunicipality>
(
<collectionCode id="828580CF5161180CFCF3FB22EC811892" box="[874,953,1213,1237]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="90" pageNumber="91">MNHN</collectionCode>
Olivier Box 68)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B815161180CFF0EFA9CEB8C19CE" pageId="90" pageNumber="91" type="diagnosis">
<paragraph id="E42B180A5161180CFF0EFA9CEB8C19CE" blockId="90.[151,1436,1283,1634]" pageId="90" pageNumber="91">
<emphasis id="D6E0C4185161180CFF0EFA9CEE321959" bold="true" box="[151,266,1283,1310]" pageId="90" pageNumber="91">Diagnosis</emphasis>
. A small species (around
<quantity id="236CB5EF5161180CFDB5FA9CED551959" box="[556,621,1283,1310]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="90" pageNumber="91" unit="mm" value="5.0">5 mm</quantity>
long) with a reddish or yellow pronotum having a median dark marking which reaches the anterior but not the posterior pronotal margin, black MS and elytra, and female brachelytral; the posterior margin of the aedeagal sheath sternite is rounded entire and the median anterior margin of the sheath tergite is widely prolonged and apically rounded. Larvae have laterally explanate tergal margins.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B815161180CFF5EFA0BEA2A1A25" pageId="90" pageNumber="91" type="discussion">
<paragraph id="E42B180A5161180CFF5EFA0BEA2A1A25" blockId="90.[151,1436,1283,1634]" pageId="90" pageNumber="91">
<emphasis id="D6E0C4185161180CFF5EFA0BEE0C19E9" bold="true" box="[199,308,1428,1454]" pageId="90" pageNumber="91">Remarks</emphasis>
. We assign
<taxonomicName id="239463895161180CFE2EFA0BED2619E9" authorityName="Olivier" authorityYear="1913" box="[439,542,1428,1454]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="D6E0C4185161180CFE2EFA0BED2619E9" box="[439,542,1428,1454]" italics="true" pageId="90" pageNumber="91">filiformis</emphasis>
</taxonomicName>
to
<taxonomicName id="239463895161180CFDD9FA0BEDAE19E9" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[576,662,1428,1454]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="D6E0C4185161180CFDD9FA0BEDAE19E9" box="[576,662,1428,1454]" italics="true" pageId="90" pageNumber="91">Luciola</emphasis>
</taxonomicName>
<emphasis id="D6E0C4185161180CFD05FA0BEDE119E9" bold="true" box="[668,729,1428,1454]" pageId="90" pageNumber="91">
<taxonomicNameLabel id="CDD379635161180CFD05FA0BEDE119E9" box="[668,729,1428,1454]" pageId="90" pageNumber="91" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
on the basis of illustrations in
<bibRefCitation id="800565FB5161180CFBB3FA0BEBFE19E9" author="Fu" box="[1066,1222,1428,1454]" firstAuthor="Fu" pageId="90" pageNumber="91" refId="ref116462" refString="Fu, X. H. (2014) An illustrated handbook of Chinese fireflies. The Commercial Press, Beijing, 167 pp. [in Chinese]" type="book" year="2014">Fu (2014: 23)</bibRefCitation>
. Differing altitudes seem to affect both male colour patterns and length of the abbreviated female elytra.
<bibRefCitation id="800565FB5161180CFBFFFA28EA171995" author="Chen" box="[1126,1327,1463,1490]" firstAuthor="Chen" pageId="90" pageNumber="91" refId="ref115752" refString="Chen, T. R. (2003) The fireflies of Taiwan. Field Image Publisher, Taipei, 256 pp. [in Chinese]" type="book" year="2003">Chen (2003: 171)</bibRefCitation>
depicts a male from middle altitudes with orange red MS, while on page 170 the male from lower altitudes has a black MS with narrow paler margins. Females from middle altitudes have longer elytra (about twice as long as median pronotal length) than those from lower altitudes. In the following reference the figure of the aedeagus may involve also at least part of the aedeagal sheath and is difficult to interpret (Chȗjô &amp; Sâtô 1971 fig. 2 page 14).
</paragraph>
</subSubSection>
</treatment>
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