treatments-xml/data/6B/89/EC/6B89ECAA735B50BA9E6ECAB068266314.xml

<document id="E3271FA29D3A45377E2D0F871884750D" ID-DOI="10.3897/zookeys.1223.138220" ID-ZooBank="3557A991-63DF-42D8-B8B1-F310CD52FD00" ID-publisher-id="138220" URI-arpha="7352E8C9-65CA-546F-BE46-EF55E52C51A6" URI-zoobank="http://zoobank.org/3557A991-63DF-42D8-B8B1-F310CD52FD00" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1736187739607" checkinUser="pensoft" docAuthor="Hendrich, Lars &amp; Brancucci, Michel" docDate="2025" docId="6B89ECAA735B50BA9E6ECAB068266314" docLanguage="en" docName="ZooKeys 1223: 87-147" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/138220/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Sandracottus femoralis Heller 1934" docType="treatment" docVersion="4" dtd-version="3.0" lastPageNumber="147" masterDocId="7352E8C965CA546FBE46EF55E52C51A6" masterDocTitle="Revision of the Oriental and Australasian diving beetle genus Sandracottus Sharp, 1882 (Coleoptera, Dytiscidae, Dytiscinae)" masterLastPageNumber="147" masterPageNumber="87" pageNumber="87" updateTime="1736200074934" updateUser="admin">
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<mods:title id="1E0EFFC95B857E5F5266B819E240E08F">Revision of the Oriental and Australasian diving beetle genus Sandracottus Sharp, 1882 (Coleoptera, Dytiscidae, Dytiscinae)</mods:title>
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<mods:namePart id="B4D6B986DD12B6567D87291A4A4BB734">Hendrich, Lars</mods:namePart>
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<mods:affiliation id="FA08FBEF63BD9AE783248B00AB3B69DD">SNSB - Zoologische Staatssammlung München, Münchhausenstraße 21, D – 81247 München, Germany</mods:affiliation>
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<mods:namePart id="A3A34157C15601A3CEE4F01695ED6A04">Brancucci, Michel</mods:namePart>
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<subSubSection id="3991D8F980C1074C59D7F31802CB46B0" type="nomenclature">
<paragraph id="63731FCFC4D8B428501E672F3E51F893">
<taxonomicName id="C3880166E58A7EBC769FD6D9C576FFA7" ID-CoL="79HLB" authority="Heller, 1934" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="2E57FB8FD1CB29813AD1854819DF2403" italics="true">Sandracottus femoralis</emphasis>
Heller, 1934
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="B6E84ABD20248FF8F48E96692147D900" type="description">
<paragraph id="FDE5B6916E7C5509443355CA90F73414">
<figureCitation id="CEAB4D7BE1378DFF5FBE738686E7DDB1" captionStart="Figures 4–7" captionStartId="F2" captionText="Figures 4–7. Habitus of 4 Sandracottus dejeanii (South India) 5 S. festivus (Sri Lanka) 6 S. femoralis (Papua New Guinea, Rigo) 7 S. hunteri (Thailand)." figureDoi="10.3897/zookeys.1223.138220.figures4-7" httpUri="https://binary.pensoft.net/fig/1214409">Figs 6</figureCitation>
, 
<figureCitation id="C5A845D8A79807B33FF4BF4A5B46F977" captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16–19. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b): 16 Sandracottus festivus 17 S. femoralis 18 S. hunteri and 19 S. insignis." figureDoi="10.3897/zookeys.1223.138220.figures16-19" httpUri="https://binary.pensoft.net/fig/1214412">17</figureCitation>
, 
<figureCitation id="9079B0BB4495BA15099166C321912547" captionStart="Figure 27" captionStartId="F11" captionText="Figure 27. Distribution of Sandracottus femoralis [black dots] in New Guinea and the Moluccas, and S. rotundus [red squares] on Sulawesi." figureDoi="10.3897/zookeys.1223.138220.figure27" httpUri="https://binary.pensoft.net/fig/1214418">27</figureCitation>
, 
<figureCitation id="04C8FBB525CFD50BEAD9779CF795942D" captionStart="Figures 31–36" captionStartId="F15" captionText="Figures 31–36. Habitats of Sandracottus 31 Australia, Northern Territory, 50 km S Adelaide River, creek at Scenic Route (NT 9) and 32 South Queensland, Winfield, Winfield Road, rest pool (QLD 54), habitats of S. bakewellii bakewellii (photos: L. Hendrich) 33 Northern Territory, Ormiston Gorge, habitat of S. bakewellii guttatus (photo: M. Baehr) 34 Pakistan, Sistan va Baluchestan Prov., Pir Sohrab env., rest pool in wadi, habitat of S. dejeanii (photo: J. Hájek) 35 India, southern Madhya Pradesh, Hoshangabad District, Pagara – Pachmarhi road, ca 5 km NNE Panchmarhi, Panar Pani, habitat of S. dejeanii and S. festivus (photo: M. Jäch) 36 Indonesia, Irian Jaya, Nabire, Nabire-Ilaga track, km 62, habitat of S. femoralis (photo: L. Hendrich)." figureDoi="10.3897/zookeys.1223.138220.figures31-36" httpUri="https://binary.pensoft.net/fig/1214422">36</figureCitation>
, 
<figureCitation id="C283753241A3015EAFD963A2D16987F7" captionStart="Figures 50–53" captionStartId="F19" captionText="Figures 50–53. Habitus of 50 Sandracottus dejeanii (India, Himachal Pradesh, Kangra Valley, Dharamsala) 51 S. festivus (Sri Lanka, Ratnapura, Sincharaja 52 S. femoralis (Indonesia, West Papua, Kapupaten Paniai, Nabire) 53 S. hunteri (China, Central Sichuan)." figureDoi="10.3897/zookeys.1223.138220.figures50-53" httpUri="https://binary.pensoft.net/fig/1214426">52</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="B1F87DA1E37EE32C80DA23AFFDACBCF8">
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<taxonomicName id="AC03C17ACC186041B9C69747A53F0C3F" authority="Guerin-Meneville, 1830: 61" authorityName="Guerin-Meneville" authorityPageNumber="61" authorityYear="1830" class="Insecta" family="Dytiscidae" genus="Dytiscus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="flavocinctus">
<emphasis id="73A1108FC305352D8C836EE5137106AD" italics="true">Dytiscus flavocinctus</emphasis>
<comment id="435700BCA30D52E4EB87013A3EEB91F2">
Guérin-Méneville, 1830: 61 (type locality Manokwari, 
<collectingRegion id="A3022E2EC2BFE662710A58AFD4E64BCC" country="Indonesia" name="Papua Barat">West Papua</collectingRegion>
, 
<collectingCountry id="7BD947FF29AB863456994C8836D76450" name="Indonesia">Indonesia</collectingCountry>
); 
<treatmentCitation id="32A7A8E659426CE1A45FC960DCDCF3B3" author="Boisduval JBA" page="49" year="1835">
<bibRefCitation id="203D94E8C56298C5FF7E9E371CFE8A84" author="Boisduval JBA" refId="B9" refString="Boisduval JBA (1835) Faune entomologique de l'Ocean Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage. 2. Coleopteres et autres ordres. Voyage de decouvertes de l'Astrolabe execute par ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, sous le commandement de M. J. Dumont d'Urville. Paris: J. Tastu, vii + 716 pp." url="1826-1827 - 1828 - 1829" year="1835">Boisduval, 1835: 49</bibRefCitation>
</treatmentCitation>
(preoccupied by 
<treatmentCitation id="B948763C7EA761519E73896FAC36C862">
<bibRefCitation id="019E56E1E216DEB5B287E16C98F273A6" author="Hummel" firstAuthor="Hummel" refId="B26" refString="Hummel AD (1823) Observations sur les insectes de 1823. Monographia pelophilarum. Novae species. In: Hummel AD (Ed.) Essais entomologiques. Vol. 1. Nr. 3. St. Pétersbourg: Imprimerie de la Chancellerie Privée du Ministère de l’Intérieur, 48 pp. [+ 1 pl.]" year="1823">Hummel 1823</bibRefCitation>
</treatmentCitation>
and objective synonym of 
<taxonomicName id="E5D37D2AF64760339DF178D7D12D89B4" ID-CoL="79HLC" authorityName="J. Balfour-Browne" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="C0C38FCB8F18F2CA009B2362EAD3951D" italics="true">Sandracottus guerini</emphasis>
</taxonomicName>
); 
<treatmentCitation id="CE49043159BF01C7D9FE8817C27AD31E" author="Boisduval JBA" page="49" year="1835">
<bibRefCitation id="667445692B92DC957D4D522E2745F308" author="Boisduval JBA" refId="B9" refString="Boisduval JBA (1835) Faune entomologique de l'Ocean Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage. 2. Coleopteres et autres ordres. Voyage de decouvertes de l'Astrolabe execute par ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, sous le commandement de M. J. Dumont d'Urville. Paris: J. Tastu, vii + 716 pp." url="1826-1827 - 1828 - 1829" year="1835">Boisduval, 1835: 49</bibRefCitation>
</treatmentCitation>
(descr.).
</comment>
</taxonomicName>
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</paragraph>
<paragraph id="47013D827FCD8C1E9D14863FC7E7FD0F">
<treatmentCitationGroup id="D89C9667C303E00096C8CC8073A94D3A">
<treatmentCitation id="4EDBC883287764A03502080C6B4529DC">
<taxonomicName id="D59B6B91EE90EFFAF89E1161C0124F3A" authority="(Guerin-Meneville, 1830)" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1830" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="flavocinctus">
<emphasis id="A95EFFCAC8751DA4C57504F424B5CEE1" italics="true">Sandracottus flavocinctus</emphasis>
<comment id="CBC9E9FDD06719401FACC2195ABD2776">
(Guérin-Méneville, 1830): 
<treatmentCitation id="6AAB77126D5B7AAEE4A11F17C08AA626" author="Régimbart M" page="339" year="1899">
<bibRefCitation id="90645B35A0D04187FF330A8AF347A87C" DOI="10.1080/21686351.1899.12279737" author="Régimbart M" firstAuthor="Regimbart" journalOrPublisher="Annales de la Société Entomologique de France" page="339" pagination="186-367" refId="B42" refString="Régimbart M (1899) Révision des Dytiscidae de la région Indo-Sino-Malaise. Annales de la Société Entomologique de France 68: 186–367. https://doi.org/10.1080/21686351.1899.12279737" title="Révision des Dytiscidae de la région Indo-Sino-Malaise." volume="68" year="1899">Régimbart 1899: 339</bibRefCitation>
</treatmentCitation>
(descr.); 
<treatmentCitation id="A66DC85903E9A0215A454DBE578164E9" author="Zimmermann A" page="234" year="1920">
<bibRefCitation id="9983A6AEB1F2396F0CEC2273334AC260" author="Zimmermann A" firstAuthor="Zimmermann" journalOrPublisher="In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71" page="234" refId="B60" refString="Zimmermann A (1920) Dytiscidae. In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71, 326 pp." title="Dytiscidae." year="1920">Zimmermann 1920: 234</bibRefCitation>
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(cat.).
</comment>
</taxonomicName>
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</paragraph>
<paragraph id="1AC676E92BD5674C4CE713DF689DB95C">
<treatmentCitationGroup id="368CA156DF86B7AB00D710A86EC8A4D1">
<treatmentCitation id="0D1A0E1870837F9F0CB90E157F8E9764">
<taxonomicName id="892EFAAA955C2153F162C67FBD24C580" ID-CoL="79HLB" authority="Heller, 1934 b: 3" authorityName="Heller" authorityPageNumber="3" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="88C68026F64986EE839B9F73B8E4A548" italics="true">Sandracottus femoralis</emphasis>
<comment id="A356E9AF32FF0096544FA5F3FB700997">
Heller, 1934 b: 3 (type locality: Buka, 
<collectingCountry id="B433FCDC20397BC8C8245627E79B10BC" name="Solomon Islands">Solomon Islands</collectingCountry>
); 
<treatmentCitation id="F1BAE1C9473D6EF5676EE6838B7E63E2" ID-DOI="http://doi.org/10.5281/zenodo.10557784" ID-Zenodo-Dep="10557784" author="Alarie Y &amp; Michat M &amp; Shaverdo H &amp; Hájek J" httpUri="http://treatment.plazi.org/id/03B39B0B7858371253DEFAF6FBCEF92E" page="305" year="2023">
<bibRefCitation id="9FE7EA4D711900D63C0AC3685AF47CD7" DOI="10.11646/zootaxa.5263.3.1" author="Alarie Y &amp; Michat M &amp; Shaverdo H &amp; Hájek J" etAl="et al." firstAuthor="Alarie" journalOrPublisher="Zootaxa" page="305" pagination="301-334" refId="B1" refString="Alarie Y, Michat M, Shaverdo H, Hájek J (2023) Morphology of the larvae of Sandracottus femoralis Heller, 1934, and S. mixtus (Blanchard, 1843) and phylogenetic comparison with other known Aciliini (Coleoptera: Dytiscidae, Dytiscinae). Zootaxa 5263: 301–334. https://doi.org/10.11646/zootaxa.5263.3.1" title="Morphology of the larvae of Sandracottus femoralis Heller, 1934, and S. mixtus (Blanchard, 1843) and phylogenetic comparison with other known Aciliini (Coleoptera: Dytiscidae, Dytiscinae)." volume="5263" year="2023">Alarie et al. 2023: 305</bibRefCitation>
</treatmentCitation>
(larval descr.), idem: 320 (habitat); Hájek and 
<treatmentCitation id="1324A09139B028CF3EE01063296843C2" author="Nilsson A &amp; Hájek J" page="91" year="2024">
<bibRefCitation id="5E0FEAB5F76A6B083FE385C073C64066" author="Nilsson A &amp; Hájek J" refId="B38" refString="Nilsson A, Hajek J (2024) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2024. Distributed as a PDF file via Internet, 323 pp. http: // www. waterbeetles. eu / documents / W _ CAT _ Dytiscidae _ 2024. pdf" url="http://www.waterbeetles.eu/documents/W_CAT_Dytiscidae_2024.pdf" year="2024">Nilsson 2024: 91</bibRefCitation>
</treatmentCitation>
(cat.).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="22F0D11AF12650B923724BDD38F85FB4">
<treatmentCitationGroup id="93350A2DBC1DF58FB691DAC3CACF6F36">
<taxonomicName id="D1C3ECE5FFDA707465CE7D106BABD89F" ID-CoL="79HLC" authority="J. Balfour-Browne, 1939: 114" authorityName="J. Balfour-Browne" authorityPageNumber="114" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="1202AB7FDCF3EBC0A087F5F2ED31AC54" italics="true">Sandracottus guerini</emphasis>
<comment id="B34DF286536473EE8A8AE4ACA1AE3091">
J. Balfour-Browne, 1939: 114, replacement name for 
<taxonomicName id="4B4CD80677A9B0EB491BE8566DD52220" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1830" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="flavocinctus">
<emphasis id="25FAE691970730F11E945EB380248C77" italics="true">Sandracottus flavocinctus</emphasis>
</taxonomicName>
; Hájek and 
<treatmentCitation id="57314026CFAAC974E0C01FD9ABAA64F0" author="Nilsson A &amp; Hájek J" page="91" year="2024">
<bibRefCitation id="0209613F79FC6C7A94C79CDDFC1C2664" author="Nilsson A &amp; Hájek J" refId="B38" refString="Nilsson A, Hajek J (2024) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2024. Distributed as a PDF file via Internet, 323 pp. http: // www. waterbeetles. eu / documents / W _ CAT _ Dytiscidae _ 2024. pdf" url="http://www.waterbeetles.eu/documents/W_CAT_Dytiscidae_2024.pdf" year="2024">Nilsson 2024: 91</bibRefCitation>
</treatmentCitation>
(cat.) (syn. nov.).
</comment>
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</paragraph>
</subSubSection>
<subSubSection id="SECID0ETVBI" type="comments">
<paragraph id="D359EC6BF74804260A433430B4BFFA9F">
<heading id="D2B5CD7F64A79F5552B3195C03E406D6" reason="title">Comments on classification.</heading>
</paragraph>
<paragraph id="251C4735CDBE35403F9CFA4A47AFB70F">
<taxonomicName id="0ED92ECCCA3331AD09479D326235E2AA" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="FB77EFF2129A73A78C83B5A64E91C813" italics="true">Sandracottus femoralis</emphasis>
</taxonomicName>
was described from the 
<collectingCountry id="30E6785A8838B75EC84CF8DEEB39DC68" name="Solomon Islands">Solomon Islands</collectingCountry>
. We have studied Heller’s type specimen of 
<taxonomicName id="F7EEB30D243613CF7E2FB0CC599A0218" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="D048047B890C5CCA7615915C1708AB2D" italics="true">S. femoralis</emphasis>
</taxonomicName>
, Guérin-Meneville’s type of 
<taxonomicName id="DCB86DC97BFFEA056FFEA4647A77092A" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1830" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="flavocinctus">
<emphasis id="3B2C9AA46EDD0280EB104AE5AB8B695F" italics="true">D. flavocinctus</emphasis>
</taxonomicName>
, as well as numerous specimens of 
<taxonomicName id="0518528AF89BBF3E3695A6A4F44EE11D" authorityName="J. Balfour-Browne" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="B9BD9C6178F22847A0730BE01FB0FAE7" italics="true">S. guerini</emphasis>
</taxonomicName>
over its complete range. There is no doubt that both taxa belong to the same species. 
<taxonomicName id="FF59A7B6756BBEB44A736C008D2DF111" authority="Balfour-Browne, 1939" authorityName="J. Balfour-Browne" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="D7A6A9B8D2237A55088F72A54922330E" italics="true">Sandracottus guerini</emphasis>
Balfour-Browne, 1939
</taxonomicName>
is a replacement name for 
<taxonomicName id="378436B939FF9A96A8BCD82DB4E8E6F3" authority="Guerin-Meneville" authorityName="Guerin-Meneville" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="flavocinctus">
<emphasis id="F83E0C91E97A1AD220A5E3C50855DFF9" italics="true">S. flavocinctus</emphasis>
Guérin-Méneville
</taxonomicName>
but is more recent than 
<taxonomicName id="D2D8D68A8B13ED32038F33554FE0EBA9" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="3EC75414BAD36A7F61650DE140ED964F" italics="true">Sandracottus femoralis</emphasis>
</taxonomicName>
, so the valid name for this species must be 
<taxonomicName id="9F3B0153E5C9FC10646BF4472AF5E6DF" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CB414944C11C8055F15D2BE9071DE0D9" italics="true">S. femoralis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERYBI" type="type material">
<paragraph id="78CCC53EF890D59970F0FF80415DC89F">
<heading id="634F361F442A412B2860CA7F01DC2B99" reason="title">Type material</heading>
</paragraph>
<paragraph id="3E085963F3EE35B44E5645751C23A24C">
<emphasis id="C4405B513ED301F537B5F53B1C5CC601" bold="true">
of 
<taxonomicName id="9E34C8C6E6CA4EA6709684C8BEDD4E09" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="3E8E695D1D338A0A0A980D5A01362A94" bold="true" italics="true">Sandracottus femoralis</emphasis>
</taxonomicName>
. 
<emphasis id="6B94751433F12E118AF62DE53791401C" bold="true" italics="true">
<typeStatus id="C560023AB4E85384FFDF3D00DC75B549" type="holotype">Holotype</typeStatus>
</emphasis>
</emphasis>
: Female, “ Buka 
<collectingCountry id="847A9E61650FC84DE032BB8995E459EE" name="Solomon Islands">Salomonen</collectingCountry>
<date id="A78ED62423D63C151EC93097AEB23D82" value="1930-07">Juli 1930</date>
”, “ Coll. H. Hediger ”, “ 1933 8 ” [blue label], “ 
<taxonomicName id="C1F49BAFBD1116E1D27EDC37974868C9" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="97BF1A08E66D29BE5C9D41916BC47CCA" italics="true">S. femoralis</emphasis>
</taxonomicName>
Typus ” [red label], “ Staatl. Museum für Tierkunde, Dresden ”, “ 
<taxonomicName id="44A6C20A9941D9C54331099DE88653B6" authority="Heller, 1934 Hendrich &amp; Brancucci" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="DA72FED9F6FC49263464B51243E5A8AB" italics="true">Sandracottus femoralis</emphasis>
Heller, 1934 Hendrich &amp; Brancucci
</taxonomicName>
det. 2006 ”. Examined.
</paragraph>
<paragraph id="2259BFA00237662FB3CD256753D1350B">
<emphasis id="3A03ECA27417ACA962BFCBE6BEC57077" bold="true">
<taxonomicName id="0028F0A4F2B84A8CF42B73BF478E7DA9" authorityName="Guerin-Meneville" authorityYear="1830" class="Insecta" family="Dytiscidae" genus="Dytiscus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="flavocinctus">
<emphasis id="0198A2D7231637145A6C9C884489A3D6" bold="true" italics="true">Dytiscus flavocinctus</emphasis>
</taxonomicName>
: 
<emphasis id="D156AF35B40A97FB76B54C164A1B9AE6" bold="true" italics="true">
<typeStatus id="D44F4E57EB192427375FB6712E4B4F3B" type="lectotype">Lectotype</typeStatus>
</emphasis>
</emphasis>
(herewith designated): Female, “ Mus. Paris Nouv. Guninée Dumont d´Urv. 1841 ”, “ Dory [Manokwari] N. 
<collectingCountry id="F75BA8FACB7B392C3BC13D2CEB411951" name="Guinea-Bissau">Guinee</collectingCountry>
Duvelle Durville ” [Dumont d´Urville], [round handwritten label], “ Hidaticus [sic!] flavocinctus ” [handwritten label], “ 
<typeStatus id="224591B1BA97D6A4482A5663C9F0EA22" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="9B91D4C085A12DB070B6E24BE1F7B023" authority="Guerin-Men." authorityName="Guerin-Meneville" authorityYear="1830" class="Insecta" family="Dytiscidae" genus="Dytiscus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="flavocinctus">
<emphasis id="159C828CF465EC53C951AAF9214361BE" italics="true">Dytiscus flavocinctus</emphasis>
Guérin-Men.
</taxonomicName>
des. M. Brancucci &amp; L. Hendrich 2010 ” [red printed label], “ 
<taxonomicName id="EE2CCF191DC09C908F3864FB1E4E2A07" authorityName="J. Balfour-Browne" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="7EE73F8292ECC516EC2882ECE204DE48" italics="true">Sandracottus guerini</emphasis>
</taxonomicName>
B. - Br. Det. M. Brancucci L. Hendrich 10 ” [white printed label] (
<abbrev id="ABBRID0EA2BI" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="F905D8F685DBCB57BA810EC0C1BE8882" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
). Examined.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EE2BI" type="material">
<paragraph id="2BEA6E9F5E7CEDD709075BF0116D7F0F">
<heading id="C47EA1CC9433781C75BC97BD6D46AC6B" reason="title">Additional material.</heading>
</paragraph>
<paragraph id="46E072AB61DE40C3C41B9B8A23824020">
<emphasis id="CCBFCE6D5159C55FF7139D3D041801AB" bold="true">
(
<specimenCount id="D979CA74913B8C63EBAA9ECB0F20EA69" count="118" type="generic">118 specimens</specimenCount>
)
</emphasis>
: • 
<emphasis id="F272410EA0F81008655ED1E8E363DA8A" bold="true">
<collectingCountry id="482C6F91EBA0E1F50C05FEC88B6F4FD6" name="Indonesia">Indonesia</collectingCountry>
</emphasis>
: 
<specimenCount id="B609C8513B40E3B0CE3164958EB1D35C" count="1" type="generic">1 ex.</specimenCount>
, “ Indonesia: Halmahera Isl., Khao Dist., Camp 34 
<quantity id="2229F8721A496CA28669998919154B74" metricMagnitude="4" metricUnit="m" metricValue="3.4" unit="km" value="34.0">34 km</quantity>
inland from 
<collectingCounty id="038DD228D7B0697A3428C030ADFEB151">Pan-Tunggal Lumber Co.</collectingCounty>
base camp at 
<collectorName id="2B57953BE5D5624131BEF3EED343BDA2">Tg. Loleo</collectorName>
”, “ 
<date id="436527DB90597683B493C01B71C598F6" value="1981-02-01" valueMax="1981-02-14" valueMin="1981-02-01">
<collectingDate id="2D18719548E94F3DE082D01ADED55821" value="1981-02-01" valueMax="1981-02-14" valueMin="1981-02-01">1-14 Feb. 1981</collectingDate>
</date>
AC Messer &amp; PM Taylor ” (
<abbrev id="ABBRID0EO2BI" xlink_title="United States National Museum of Natural History">
<collectionCode id="EA30D75A644F0B8133575A478F30B7BC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</abbrev>
); 
<specimenCount id="5F0DC4BFB8890778E2F033391845A90B" count="1" type="generic">1 ex.</specimenCount>
, “ Indonesia: Halmahera Isl., Jailolo Dist., Kampung Psir Putih 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EV2BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[127.683333,0.883333]}">
<geoCoordinate id="5A60866B7BCD939B7D60ECF0EC99785A" degrees="0" direction="north" minutes="53" orientation="latitude" precision="925" value="0.8833333">0 ° 53 ' N</geoCoordinate>
<geoCoordinate id="1BABCDEB9982242C2E0B4E75DFACB472" degrees="127" direction="east" minutes="41" orientation="longitude" precision="925" value="127.683334">127 ° 41 ' E</geoCoordinate>
</named-content>
”, “ 
<date id="A81FBACE702D13E85879D846267C8EF3" value="1981-08-01" valueMax="1981-08-14" valueMin="1981-08-01">
<collectingDate id="2C1C6EB135676E7395C2D529F90D4342" value="1981-08-01" valueMax="1981-08-14" valueMin="1981-08-01">1-14 Aug. 1981</collectingDate>
</date>
PM Taylor ” (
<abbrev id="ABBRID0E12BI" xlink_title="United States National Museum of Natural History">
<collectionCode id="DF5038EE7C458CEB39E2BF64C3291D3D" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</abbrev>
); 
<specimenCount id="912077D57F5E744817376098E166ED97" count="13" type="generic">13 exs.</specimenCount>
, “ Indonesia: Halmahera Isl., Jailolo Dist., Kampung Psir Putih 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EB3BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[127.683333,0.883333]}">
<geoCoordinate id="8D8845FFFEC17A25AFE56CA41A3A7AB5" degrees="0" direction="north" minutes="53" orientation="latitude" precision="925" value="0.8833333">0 ° 53 ' N</geoCoordinate>
<geoCoordinate id="CBBE00876875B28374EE0FA236F2BF05" degrees="127" direction="east" minutes="41" orientation="longitude" precision="925" value="127.683334">127 ° 41 ' E</geoCoordinate>
</named-content>
”, “ 
<date id="B477BD4A550241D355078524A8128D77" value="1981-01-15" valueMax="1981-01-31" valueMin="1981-01-15">
<collectingDate id="49EAE319D429A7FF81C0D8562B47008B" value="1981-01-15" valueMax="1981-01-31" valueMin="1981-01-15">15–31 Jan. 1981</collectingDate>
</date>
AC Messer &amp; PM Taylor ” (
<abbrev id="ABBRID0EG3BI" xlink_title="United States National Museum of Natural History">
<collectionCode id="1F55DBE911C46D3DE66DB6C63D3F6C87" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</abbrev>
); 
<specimenCount id="D16D88E64B7DC4B7079248467D53B1C8" count="18" type="generic">18 exs.</specimenCount>
, “ Indonesia: Halmahera Isl., Jailolo Dist., Kampung Psir Putih 
<geoCoordinate id="2656AB35DEB8648A1A70BD69B4286A3D" degrees="0" direction="north" minutes="53" orientation="latitude" precision="925" value="0.8833333">
0 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EN3BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[127.683333,53.000000]}">
° 53 ' N 
<geoCoordinate id="6653946B834D8F7AF0C6E023FE73C4E8" degrees="127" direction="east" minutes="41" orientation="longitude" precision="925" value="127.683334">127 ° 41 ' E</geoCoordinate>
</named-content>
</geoCoordinate>
”, “ 
<date id="D550769D5DF69B80B86FA26DDA69BF82" value="1981-05-01" valueMax="1981-05-14" valueMin="1981-05-01">
<collectingDate id="3ECA4A72C5D53B53E0350E3DA3A05CC6" value="1981-05-01" valueMax="1981-05-14" valueMin="1981-05-01">1-14 May 1981</collectingDate>
</date>
AC Messer &amp; PM Taylor ” (
<abbrev id="ABBRID0ES3BI" xlink_title="United States National Museum of Natural History">
<collectionCode id="D7468D11B780851706FBCD17F3F44FD0" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</abbrev>
); 
<specimenCount id="A2517F0728130D5DF6A6E4F618372DBA" count="2" type="generic">2 exs.</specimenCount>
, “ Indonesia: Halmahera Isl., Jailolo Dist., Kampung Psir Putih 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZ3BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[127.683333,0.883333]}">
<geoCoordinate id="DC150B987F2F3AAB8A0E3B0442855F02" degrees="0" direction="north" minutes="53" orientation="latitude" precision="925" value="0.8833333">0 ° 53 ' N</geoCoordinate>
<geoCoordinate id="4BEA24B52321C091650AD6D70FD4B666" degrees="127" direction="east" minutes="41" orientation="longitude" precision="925" value="127.683334">127 ° 41 ' E</geoCoordinate>
</named-content>
”, “ 
<date id="A3FF994A8B3D267B90C230750B1848B4" value="1981-08-14">
<collectingDate id="9868DA5EAB4AD2EE68A173585FDC6A7E" value="1981-08-14">14. Aug. 1981</collectingDate>
</date>
PM Taylor ” (
<abbrev id="ABBRID0E53BI" xlink_title="United States National Museum of Natural History">
<collectionCode id="11FE30A3542F044D7CA90784E0E1DC1A" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</abbrev>
); 
<specimenCount id="AECAAA517BD57EED2889EC31880F81C7" count="1" type="generic">1 ex.</specimenCount>
, “ Batanta Island [Pulau Batanta], Arefi, 2004, 
<collectorName id="C6E1C296D094BBE6A4EA945160CA3ACF">A. Skale</collectorName>
leg. ” (
<abbrev id="ABBRID0EC4BI" xlink_title="Collection André Skale">
<collectionCode id="DBEA8F942762CD6D475CE602FCAEEBC1" country="USA" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">CAS</collectionCode>
</abbrev>
); 
<specimenCount id="BC197B0AD1A4AD50279E3DFA045AED58" count="1" type="generic">1 ex.</specimenCount>
, “ Bernstein Morotai ”, “ flavocinctus Aubé ” (
<abbrev id="ABBRID0EG4BI" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="71653A62994DFB5ED46F70E806563BD6" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
); 
<specimenCount id="197C414E9C51CB3D299FE4021E22CCB4" count="1" type="generic">1 ex.</specimenCount>
, “ Aru Insel ”, “ 
<collectorName id="BD5F2DCDA0E15ED29B5565339EF67581">Samml. A. Zimmermann</collectorName>
” (
<abbrev id="ABBRID0EK4BI" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="5D1FF91FFFFB9B4AF1AA8F84E225E1A0" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
); 
<specimenCount id="F1351C271D840E6156C161A4EBE97CAC" count="1" type="generic">1 ex.</specimenCount>
, “ Rosenberg, Ins. Aru ” (
<abbrev id="ABBRID0EO4BI" xlink_title="Nationaal Natuurhistorisch Museum">
<collectionCode id="796D7C623B47513FF9CBF181E6110143" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</abbrev>
); 
<specimenCount id="772259E877D5CA2403B897F43BFEE82A" count="1" type="generic">1 ex.</specimenCount>
, “ Key Insel ”, “ 
<collectorName id="E8EB2C034870F3036B4001F3200B4E46">Samml. A. Zimmermann</collectorName>
” (
<abbrev id="ABBRID0ES4BI" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="E1F7E6388B35C65D7660B2C02D2DFC1C" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
); 
<specimenCount id="05F43C4A412C6190FB9652A163B71E03" count="2" type="generic">2 exs.</specimenCount>
, “ Bernstein, Morotai ” (
<abbrev id="ABBRID0EW4BI" xlink_title="Nationaal Natuurhistorisch Museum">
<collectionCode id="9AB7A602D45E1C26578D3058153FFB03" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</abbrev>
); 
<specimenCount id="D25784A6FE4A3CDA3D701660B42EF2F9" count="1" type="generic">1 ex.</specimenCount>
, “ 
<collectorName id="18F1B73ED819177DACB608C4C27C3310">N. Moluccas</collectorName>
, 
<collectingMunicipality id="275BB427A9372E43D5B5B9DAAAB66225">Bacan Island</collectingMunicipality>
, 
<location id="FB103DE202A27654CA4A7D0EEE4046DC" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:FB103DE202A27654CA4A7D0EEE4046DC" country="Indonesia" county="Pan-Tunggal Lumber Co." latitude="-0.80775255" longLatPrecision="1" longitude="133.05923" municipality="Bacan Island" name="Mt. Sibela" stateProvince="Maluku Utara">Mt. Sibela</location>
, 
<quantity id="6FDE7BE2CD78604A900B1E445BC7570B" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
<elevation id="A596C6C7D69B4FD9A8E2845528B5C763" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</elevation>
</quantity>
, 
<locationDeviation id="07DAA248013CA57920EBDCE82C40C13D" location="Labuha">
<quantity id="55B988F6A09FF6D277C5781D52C50509" metricMagnitude="4" metricUnit="m" metricValue="1.4" unit="km" value="14.0">14 km</quantity>
SE Labuha
</locationDeviation>
, primary forest, 
<date id="F5DC551FED6AAE5EC794D2BE41640CF3" value="1996-02-02" valueMax="1996-02-13" valueMin="1996-02-02">
<collectingDate id="E22213E67E4457CA61BE3F478CD5A852" value="1996-02-02" valueMax="1996-02-13" valueMin="1996-02-02">2–13. II. 1996</collectingDate>
</date>
, 
<geoCoordinate id="AF49553F0708BE5FC0A1AB07B2EAC76A" degrees="0" direction="south" minutes="38" orientation="latitude" precision="925" value="-0.6333333">0 ° 38´S</geoCoordinate>
Lat. 
<geoCoordinate id="60E041B4A35A6EEE09F8DDF4D2D53CA0" degrees="127" direction="east" minutes="32" orientation="longitude" precision="925" value="127.53333">127 ° 32´E</geoCoordinate>
, leg. 
<collectorName id="C8BEDE883BD952F2E99808D4E0D202C7">V. Siniaev</collectorName>
&amp; 
<collectorName id="0A74DD9577133AE2C955B7A0CEA37EE6">E. Afonin</collectorName>
leg. ” (
<abbrev id="ABBRID0E14BI" xlink_title="Collection Lars Hendrich">
<collectionCode id="0CF132184B9C60EC1A0E7317237E0189">CLH</collectionCode>
</abbrev>
); 
<specimenCount id="AD744CAD9CE4999069F990EC2F75A93F" count="2" type="generic">2 exs.</specimenCount>
, “ SE Moluccas Aru ISLS, WOKAM I. 
<locationDeviation id="07AE59EEC579410AD38DBA69903835DF" location="Wakua">
<quantity id="4CA4F10D9EF613373AA11F0DCD2B4653" metricMagnitude="4" metricUnit="m" metricValue="1.7" unit="km" value="17.0">17 km</quantity>
NE Wakua
</locationDeviation>
vill., 
<date id="197C73E8127F748809FFAC4CEDED503D" value="2022-02-01" valueMax="2022-02-07" valueMin="2022-02-01">
<collectingDate id="D8248B70754DECDEAE342FC8E0D389E5" value="2022-02-01" valueMax="2022-02-07" valueMin="2022-02-01">1–7 - II. 2022</collectingDate>
</date>
, 
<location id="4535D41D72CEC09278ADC1AAE8C37EE8" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:4535D41D72CEC09278ADC1AAE8C37EE8" country="Indonesia" county="Pan-Tunggal Lumber Co." latitude="-0.80775255" longLatPrecision="1" longitude="133.05923" municipality="Bacan Island" name="St. Jakl" stateProvince="Maluku Utara">
<collectorName id="A40CE4BED5B124B16AC803C170377F36">St.</collectorName>
Jakl
</location>
leg. ” (
<abbrev id="ABBRID0E54BI" xlink_title="Collection Lars Hendrich">
<collectionCode id="97893D5E31A0AFFCF95AD110CE3034B2">CLH</collectionCode>
</abbrev>
, 
<abbrev id="ABBRID0EC5BI" xlink_title="Národní muzeum, Praha">
<collectionCode id="6B80C8A20477906F9B2D2989EAC5DEAA" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
</abbrev>
); 
<specimenCount id="8C6D17B0FB3D42421AD353EE1891D1D0" count="8" type="generic">8 exs.</specimenCount>
, “ N Moluccas: Bacan, Wayaua, alt. m. 0050, 
<date id="A26D200CA3700975DF901587C9E59AAF" value="1985-07-05" valueMax="1985-07-16" valueMin="1985-07-05">
<collectingDate id="B3FDFFCBF287366C4869DA3A15F71D34" value="1985-07-05" valueMax="1985-07-16" valueMin="1985-07-05">05. - 16. vii. 1985</collectingDate>
</date>
, 
<collectorName id="5E45327E415548E508B775EC57982C43">J. Huijbregts</collectorName>
” (
<abbrev id="ABBRID0EG5BI" xlink_title="Nationaal Natuurhistorisch Museum">
<collectionCode id="7BC552B2B530ADB2BB9DA4DD13003EC0" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</abbrev>
); 
<specimenCount id="D95645C891175DABE4A155779C657E28" count="1" type="generic">1 ex.</specimenCount>
, “ Indonesia, 
<collectingRegion id="5912EBDA0E8814A7D5B86C78C6E6FCD9" country="Indonesia" name="Maluku Utara">Maluku</collectingRegion>
, 
<location id="0AF1961A3B083DD6B778BC5FB1F78F0A" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:0AF1961A3B083DD6B778BC5FB1F78F0A" country="Indonesia" county="Pan-Tunggal Lumber Co." latitude="-0.80775255" longLatPrecision="1" longitude="133.05923" municipality="Bacan Island" name="Obi Isl." stateProvince="Maluku Utara">Obi Isl.</location>
, South coast, 
<locationDeviation id="34B10AEAD4F9B518D79936976354027C" location="Tapaya">
<quantity id="742CF747BFFD310A4E920A08BC6B62A8" metricMagnitude="4" metricUnit="m" metricValue="2.2" unit="km" value="22.0">22 km</quantity>
N of Tapaya
</locationDeviation>
vill., 
<location id="024D5E5D89364A0CB7A470204F0C9F65" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:024D5E5D89364A0CB7A470204F0C9F65" country="Indonesia" county="Pan-Tunggal Lumber Co." latitude="-0.80775255" longLatPrecision="1" longitude="133.05923" municipality="Bacan Island" name="Seribu Mts." stateProvince="Maluku Utara">Seribu Mts.</location>
, 
<quantity id="2EACA6C79833620B9330A40860F781D6" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">
<elevation id="CA0317BFD5E15F8A9A46B5F3FE82221C" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">1200–1500 m</elevation>
</quantity>
, 
<collectorName id="0F362F1D0F2CD82AAC25E017F7560C3A">S. Jákl</collectorName>
leg., 
<date id="3187CED056DBA662BD4B4574A29115B9" value="2008-11-20" valueMax="2008-12-10" valueMin="2008-11-20">
<collectingDate id="8EE44FAB2A95A0AC86295D822D718F54" value="2008-11-20" valueMax="2008-12-10" valueMin="2008-11-20">20. xi- 10. xii. 2008</collectingDate>
</date>
” (
<abbrev id="ABBRID0EK5BI" xlink_title="Národní muzeum, Praha">
<collectionCode id="F53B85DB8B0B1D6FCF6A350E7149A137" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
</abbrev>
); 
<specimenCount id="BECFBADA412141255B4C213EAD2E237C" count="3" type="generic">3 exs.</specimenCount>
, “ Indonesia, 
<collectingRegion id="ECF5C1FB21346C915D56AE0053104152" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
Dekai, upper Brazza, 
<quantity id="B02CF35F88C53FC0E4F8559847E0B989" metricMagnitude="2" metricUnit="m" metricValue="2.73" unit="m" value="273.0">
<elevation id="F682278AD7C2571E73EC4D64B0CD8FBE" metricMagnitude="2" metricUnit="m" metricValue="2.73" unit="m" value="273.0">273 m</elevation>
</quantity>
, 2. / 
<date id="EDE670BA78BA7186E572CCEFDB95F03E" value="2015-06-03">
<collectingDate id="19AEA68FF936A6534DDD49E010E84CBD" value="2015-06-03">3. vi. 2015</collectingDate>
</date>
, 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ER5BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[139.654211,-4.741085]}">- 4.74108472466468 139.654211075976</named-content>
, Sumoked (Pap 044) ” (
<abbrev id="ABBRID0EW5BI" xlink_title="Museum Zoologicum Bogoriense">
<collectionCode id="2BD5904B7578771A422617D69E80DD44" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
</abbrev>
, 
<abbrev id="ABBRID0E15BI" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="D95FCDEB6BE1A1A9568E0B878D41E29F" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
); 
<specimenCount id="BA0DC4B10466170A18BF93545B4CD292" count="2" type="generic">2 exs.</specimenCount>
, “ Indonesia 
<collectingRegion id="D0B5367B9A68BAD6EE1BF82196F40EA7" country="Indonesia" name="Papua Barat">Papua Barat</collectingRegion>
, Kebar, shaded deep sandy irrigation roadside ditches, 
<quantity id="8B434082449CB5FAE57BF49FE0478BB0" metricMagnitude="2" metricUnit="m" metricValue="5.84" unit="m" value="584.0">
<elevation id="7CB32073143DF4A302522185166544F9" metricMagnitude="2" metricUnit="m" metricValue="5.84" unit="m" value="584.0">584 m</elevation>
</quantity>
, 
<date id="FED8EDE5A446D99CA3C2E1C3B2F9FE25" value="2013-11-06">
<collectingDate id="BE5DD2DA32D6F804E7A51D59652F1C34" value="2013-11-06">6. xi. 2013</collectingDate>
</date>
, 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EB6BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[133.059235,-0.807753]}">
<geoCoordinate id="12862965BB0084ED02EF144162C565F3" degrees="0.80775253" direction="south" orientation="latitude" precision="1" value="-0.80775255">- 0.80775253</geoCoordinate>
<geoCoordinate id="FC2B39E70B4C8785F08C4B00E65A33CB" degrees="133.05923529" direction="east" orientation="longitude" precision="1" value="133.05923">133.05923529</geoCoordinate>
</named-content>
, UNIPA (BH 031) ” (
<abbrev id="ABBRID0EG6BI" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="A1C0581D30B2086E5ED235E37398248A" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
); 
<specimenCount id="BDD405290B323D0039C7EC8F79454AC3" count="4" type="generic">4 exs.</specimenCount>
, “ 
<collectingRegion id="7A5CCB9C2B59E74F1B4F57A5161E6C69" country="Indonesia" name="Papua Barat">Irian Jaya</collectingRegion>
, Nabire, Nabire-Ilaga track, km 
<quantity id="3E7985E59AC11609D9346A4B55D43389" metricMagnitude="4" metricUnit="m" metricValue="6.225" unit="m" value="62250.0">
<elevation id="EA09552194D00D1E4BA87B97974B878C" metricMagnitude="4" metricUnit="m" metricValue="6.225" unit="m" value="62250.0">62, 250 m</elevation>
</quantity>
, 
<date id="56BDA303663AF161DC40BE30203C4EFE" value="1991-07-24">
<collectingDate id="314813A510F14D8274CF90BFC086D3E5" value="1991-07-24">24. VII. 1991</collectingDate>
</date>
, 
<collectorName id="15F956604B4B9E65E671110A70D4D6EE">M. Balke</collectorName>
&amp; 
<collectorName id="1BC21CBDE9A274C93CFA75536B6BB59A">L. Hendrich</collectorName>
leg. (IR 22) ” (
<abbrev id="ABBRID0EK6BI" xlink_title="Collection Lars Hendrich">
<collectionCode id="0C1339BBBCF106F526D3287CA027FED1">CLH</collectionCode>
</abbrev>
, 
<abbrev id="ABBRID0EO6BI" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="BD79897894CDBC433CE4980A4C72E9AD" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
); 
<specimenCount id="90269419255AFBDA44A5FDEDEF6AC9CC" count="1" type="generic">1 ex.</specimenCount>
, “ 
<location id="5FE7743A99F26A803C565A55854F1952" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:5FE7743A99F26A803C565A55854F1952" country="Indonesia" county="Pan-Tunggal Lumber Co." latitude="-0.80775255" longLatPrecision="1" longitude="133.05923" municipality="Bacan Island" name="Tami River" stateProvince="Maluku Utara">
Tami 
<location id="A1578FFDCA21CE30BBE64589C12092B9" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:A1578FFDCA21CE30BBE64589C12092B9" country="Indonesia" county="Pan-Tunggal Lumber Co." latitude="-0.80775255" longLatPrecision="1" longitude="133.05923" municipality="Bacan Island" name="River Hollandia" stateProvince="Maluku Utara">River Hollandia</location>
</location>
1930 
<collectorName id="F8D616A4B9AAA60CBC97A8C5724B8456">R. Voorhoeve</collectorName>
leg. ” (
<abbrev id="ABBRID0ES6BI" xlink_title="Collection Lars Hendrich">
<collectionCode id="3250B40797B1C99B63B99EAD7F3CFBB8">CLH</collectionCode>
</abbrev>
); 
<specimenCount id="6E724B3D2919C2528167B00ADCB94744" count="1" type="generic">1 ex.</specimenCount>
, “ Seram, E Wahai, 
<date id="F089EBCDAA359AF1FB76C7D69A7489D6" value="1989-02-12">
<collectingDate id="A635D0DF7C1655222C8D609821163C26" value="1989-02-12">12. II. 1989</collectingDate>
</date>
, 
<collectorName id="5A19BBCD854D1324121DA81714943CF0">M. Jäch</collectorName>
” (
<abbrev id="ABBRID0EW6BI" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="56A64E3CE25BB336A1A4D41C36132F0B" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
); 
<specimenCount id="BB3D1CC7175325F350BD46331434C324" count="1" type="generic">1 ex.</specimenCount>
, “ New Guinea, Wareo, 1933 ” (
<abbrev id="ABBRID0E16BI" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="9DBCDDC753758ABFDF02C6B2F9A89CE6" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
); 
<specimenCount id="39B889DE0FCC7B9F734F4AA245B852EF" count="6" type="generic">6 exs.</specimenCount>
, “ Dutch New Guinea: Humboldt Bay Dist. Pukusam Dist. West of Tami River 
<date id="2A45B33F5A1686C07CBE1ECF68E2C364" value="1937-06">
<collectingDate id="E5A570258D2D5DEBEDC6034F8D85FAFE" value="1937-06">vi. 1937</collectingDate>
</date>
”, “ 
<collectorName id="94352044570F5D13422B68D9DC5B27BD">B. M. Nat. Hist. London</collectorName>
” (
<abbrev id="ABBRID0E56BI" xlink_title="Collection Lars Hendrich">
<collectionCode id="B0D1DEE79AF09042FACB004A3BBC12C3">CLH</collectionCode>
</abbrev>
, 
<abbrev id="ABBRID0ECACI" xlink_title="Natural History Museum, London">
<collectionCode id="64068FBC3A8BD37BBFD4183D497BFCA3" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
). • 
<emphasis id="CCED8E99AC7E8E36F5A236B41CA1D28C" bold="true">
<collectingCountry id="7EAB11DE7682076BDD3814E4939226AD" name="Papua New Guinea">Papua New Guinea</collectingCountry>
</emphasis>
: 
<specimenCount id="14551C72EB7D733107D0490961FF461C" count="1" type="generic">1 ex.</specimenCount>
, “ Nouvelle Guinée ” (
<abbrev id="ABBRID0EIACI" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="84EE73578EB4F69BE3E01B24DC704C91" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
); 
<specimenCount id="C21A0440ED0458E178B86AC55795F6DE" count="1" type="generic">1 ex.</specimenCount>
, “ 
<collectingCounty id="F733FCF1EE6F3FD8B170AA0905DBA0CE">
New 
<collectorName id="0EC32F45B806D7C50B4AB325A8B85AE1">Guinea, NE</collectorName>
</collectingCounty>
Zenag-Lae, 
<quantity id="DA1557F95DC71482C808B012D800D95D" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation id="999C95BCA4683A5207C0C2BC87D755CB" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
, 
<date id="593B11F6C6280193CC71B5B29EB76326" value="1979-01-16">
<collectingDate id="219C7EC6BD52F440010A737D0D38F5FC" value="1979-01-16">16. I. 1979</collectingDate>
</date>
” (
<abbrev id="ABBRID0EMACI" xlink_title="Naturhistorisches Museum Basel">
<collectionCode id="2A8A3EED8DA72560599A45FC5FCBCE17" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" type="">NMB</collectionCode>
</abbrev>
); 
<specimenCount id="0E80A911FEE23E92659D33EE01986144" count="10" type="generic">10 exs.</specimenCount>
, “ 
<collectorName id="FB339CEE04473F55E3655A95ECC97EA6">N. Guinea</collectorName>
mer. 
<collectingMunicipality id="79DFE0560475F7A397F09AB680EDC368">Rigo Lugio</collectingMunicipality>
1889 
<collectorName id="EFB5BC254D0521AD61256B8D3BA687D8">L. Loria</collectorName>
” (
<abbrev id="ABBRID0EQACI" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="A3FACBB021C0336E706C3C113EDE985D" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
, 
<abbrev id="ABBRID0EUACI" xlink_title="Nationaal Natuurhistorisch Museum">
<collectionCode id="72E1D7A81221D60D6144C0FFB855D5B9" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</abbrev>
); 
<specimenCount id="F9E86F7B5E4F0434121F7A9BBED6E66D" count="5" type="generic">5 exs.</specimenCount>
, “ 
<collectorName id="EBFF8C9BDAFA96A4B358DB98BD0EDC6A">N. Guinea</collectorName>
mer. 
<location id="1C847EFF921ED29724AFA0CB20F05721" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:1C847EFF921ED29724AFA0CB20F05721" country="Papua New Guinea" county="New Guinea" latitude="-5.133333" longLatPrecision="1305" longitude="145.76666" municipality="Rigo Lugio" name="Rigo" stateProvince="Morobe">Rigo</location>
<date id="4DC9B7C212223403B4EE7DDB304076BD" value="1889-07">
<collectingDate id="FFF17584B469483EA16AE7AA5F49683C" value="1889-07">VII. 1889</collectingDate>
</date>
<collectorName id="D1627B91AF7D8D197291067FA9EAD8D8">L. Loria</collectorName>
” (
<abbrev id="ABBRID0EYACI" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="97F5D858CDCE985D57F47A8FF2565ECB" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
, 
<abbrev id="ABBRID0E3ACI" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="D36625D8DC5B1893034FCE6562C0EA40" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
); 
<specimenCount id="82ED5B4ADF19654BCC5DC4F0954FCEDF" count="1" type="generic">1 ex.</specimenCount>
, “ 
<collectingRegion id="E0CF2EE7B9160B8CAEDBD4D6BED29DFC" country="Papua New Guinea" name="Morobe">Morobe Prov.</collectingRegion>
<location id="10EDE26E0433B92A3A9EC77E395C0692" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:10EDE26E0433B92A3A9EC77E395C0692" country="Papua New Guinea" county="New Guinea" latitude="-5.133333" longLatPrecision="1305" longitude="145.76666" municipality="Rigo Lugio" name="Wau" stateProvince="Morobe">Wau</location>
, 
<quantity id="62C8DA71F7F56026BAA064A13FA3BDC4" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<elevation id="EC87F94AC3759EAE8A0C7DF4E587F48C" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
</quantity>
<date id="AF2A479018880B9C9FD1CCF31EAF6563" value="1962-05-01" valueMax="1962-05-15" valueMin="1962-05-01">
<collectingDate id="96BAB4CD330ED18A89AF362FA0BA1B4F" value="1962-05-01" valueMax="1962-05-15" valueMin="1962-05-01">1. - 15. V. 1962</collectingDate>
</date>
<collectingMethod id="9A3D5F9E1EE751D42F15A19787F57D90">light trap</collectingMethod>
<collectorName id="4D4988F18309592CF3FC776565BB1F33">J. Sedlacek</collectorName>
leg. ” (
<abbrev id="ABBRID0EABCI" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="91FF90AAE341E99E507D1DCEC1B10295" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
); 
<specimenCount id="F0D93ABBC308D82E7C2A547558B5F4F8" count="1" type="generic">1 ex.</specimenCount>
, “ 
<collectingRegion id="AD89BF3113291A542F0BCC0796EF996E" country="Papua New Guinea" name="Morobe">Morobe Prov.</collectingRegion>
<location id="35B684FD9E3CBEE350408D92099879D0" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:35B684FD9E3CBEE350408D92099879D0" country="Papua New Guinea" county="New Guinea" latitude="-5.133333" longLatPrecision="1305" longitude="145.76666" municipality="Rigo Lugio" name="Bulolo" stateProvince="Morobe">Bulolo</location>
<quantity id="F786258B377DED76A389095C109CCB8E" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">
<elevation id="566ECACE9EB30161B1B74433F7E6A2D0" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">700 m</elevation>
</quantity>
<date id="ACADD40EE2D021A753C9A48B83E62DA9" value="1962-11-09">
<collectingDate id="C95452FA17DEF0D81ABBC5CBDEB9C8B8" value="1962-11-09">9. XI. 1962</collectingDate>
</date>
<collectorName id="DD2F074BF3F9AA268DD3CED48F319462">J. &amp; M. Sedlacek</collectorName>
leg. ” (
<abbrev id="ABBRID0EEBCI" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="523EF6DDC1A2100F2A25A20C806C5054" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
); 
<specimenCount id="8A0B9F22C822F3509366B5FC89A46BEF" count="1" type="generic">1 ex.</specimenCount>
, “ 
<collectingRegion id="CFCAC87DCF0007DDBBC22FE84CF63EF2" country="Papua New Guinea" name="Morobe">Morobe</collectingRegion>
<location id="6F6B7E25D67D3DD247742B5985F6EA02" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:6F6B7E25D67D3DD247742B5985F6EA02" country="Papua New Guinea" county="New Guinea" latitude="-5.133333" longLatPrecision="1305" longitude="145.76666" municipality="Rigo Lugio" name="Kilolo Creek" stateProvince="Morobe">Kilolo Creek</location>
<locationDeviation id="FD2CE9BEBCC6BE4F499635AA2DC2D892" location="Wau">
<quantity id="46C220CC8480CD4CA612479D19777E2A" metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
W Wau
</locationDeviation>
<quantity id="DD12E72D729A89E94A358730B947BE30" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" unit="m" value="1070.0">
<elevation id="880E7DF7FFC3A3A29C04318DC5F73E82" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" unit="m" value="1070.0">1070 m</elevation>
</quantity>
<date id="7F11867C589DEFEB49FD186A9F65F566" value="1969-01-15" valueMax="1969-01-21" valueMin="1969-01-15">
<collectingDate id="70B9061FF89EBF1B29D6EAFD90CEDD53" value="1969-01-15" valueMax="1969-01-21" valueMin="1969-01-15">15. - 21. I. 1969</collectingDate>
</date>
<collectorName id="611A0A304B1A76D89CE2F3D5B602881A">J. Sedlacek</collectorName>
leg. ” (
<abbrev id="ABBRID0EIBCI" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="F8D007DC0102DE6AF912A738C9319016" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
); 
<specimenCount id="1F7C26D52DA26FF7F4265B93BBDB49BF" count="1" type="generic">1 ex.</specimenCount>
, “ 
<collectingRegion id="6030061ED20EEFF2552AF4BF4C8A77EB" country="Papua New Guinea" name="Morobe">Morobe Prov.</collectingRegion>
<location id="4C646A770194995F43F61CEE0BB747AB" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:4C646A770194995F43F61CEE0BB747AB" country="Papua New Guinea" county="New Guinea" latitude="-5.133333" longLatPrecision="1305" longitude="145.76666" municipality="Rigo Lugio" name="Garaina" stateProvince="Morobe">Garaina</location>
<quantity id="AA3E61FFBFD349C682DC517DD71B9DD1" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">
<elevation id="4D1FE2119712F741526330424F3583E2" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">700 m</elevation>
</quantity>
<date id="D335209A2A484E28B881CC35C0853D9A" value="1998-03-21">
<collectingDate id="C3EC3B6D1469A67BD20916076AAEFBA3" value="1998-03-21">21. III. 1998</collectingDate>
</date>
<collectorName id="A8E87DCB910921B06FF4852046CAFC97">A. Riedel</collectorName>
leg. ” (
<abbrev id="ABBRID0EMBCI" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="E00AD0C98C63EB92777BD706C31E7B23" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
); 
<specimenCount id="F2E6B151F4AEF2BD58D5BBA5E64882EE" count="1" type="generic">1 ex.</specimenCount>
, “ 
<collectingRegion id="A67EC80895CE953AA1B9B431738F89B7" country="Papua New Guinea" name="Morobe">Morobe Prov.</collectingRegion>
<location id="15E2E906AC08692196FDAACB4C5AA85C" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:15E2E906AC08692196FDAACB4C5AA85C" country="Papua New Guinea" county="New Guinea" latitude="-5.133333" longLatPrecision="1305" longitude="145.76666" municipality="Rigo Lugio" name="Wau" stateProvince="Morobe">Wau</location>
<date id="0F501257FFDFCC64E870860FF18BD36F" value="1968-12-15">
<collectingDate id="E80777F657AA8302B3EF079D27560311" value="1968-12-15">15. XII. 1968</collectingDate>
</date>
<collectorName id="60B5241DE49D2A4EBB67486D9073E946">H. Ohlmus</collectorName>
leg. ” (
<abbrev id="ABBRID0EQBCI" xlink_title="Australian National Insect Collection">
<collectionCode id="55C165840CBD0A540C85FE3DF2B01218" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
); 
<specimenCount id="F171F9C48E06E32D5361976DABD47B09" count="6" type="generic">6 exs.</specimenCount>
, “ 
<collectingRegion id="3CF82ECCF9D569F202401E8504EEA322" country="Papua New Guinea" name="Madang">Madang Prov.</collectingRegion>
<location id="96DB45DC9445B8243445ED0E1F603702" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:96DB45DC9445B8243445ED0E1F603702" country="Papua New Guinea" county="New Guinea" latitude="-5.133333" longLatPrecision="1305" longitude="145.76666" municipality="Rigo Lugio" name="Batabag village" stateProvince="Morobe">Batabag village</location>
<quantity id="A1088889099E31FBE0381526F4506199" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">
<elevation id="32794820344F0287A752CE291FB6AAE5" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</elevation>
</quantity>
<date id="BE0C229848AF4857AE1A9BB5F8C60765" value="2000-11" valueMax="2000-12-31" valueMin="2000-11-00">
<collectingDate id="F45298A5E8DEBB3320BBB5D34CF71AA7" value="2000-11" valueMax="2000-12-31" valueMin="2000-11-00">XI. - XII. 2000</collectingDate>
</date>
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXBCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[145.766667,-5.133333]}">
<geoCoordinate id="900097ECE873A4E1927B0ED71CC93F6A" degrees="5" direction="south" minutes="08" orientation="latitude" precision="925" value="-5.133333">5 ° 08 ' S</geoCoordinate>
<geoCoordinate id="436DED4D8446A7561B7D4B0F24E5E790" degrees="145" direction="east" minutes="46" orientation="longitude" precision="925" value="145.76666">145 ° 46 ' E</geoCoordinate>
</named-content>
<collectorName id="4D04A927BD2AACF43026B7F36C12E6A0">L. Cizek</collectorName>
leg. ” (
<abbrev id="ABBRID0E3BCI" xlink_title="Naturhistorisches Museum Basel">
<collectionCode id="8647C9428647FD3AA9F01A877DAD0F8A" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" type="">NMB</collectionCode>
</abbrev>
); 
<specimenCount id="B40E78C538AE430E0FF8D9806ECB865A" count="1" type="generic">1 ex.</specimenCount>
, “ 
<collectingRegion id="E2D1CF9650ABDBED748400268A17630E" country="Papua New Guinea" name="Madang">Madang</collectingRegion>
Alexishafen, sago swamp, 
<date id="F5A4FF9D62BDC52768D2EF2D8CFD5423" value="1991-05-22">
<collectingDate id="B1D38CDC78257115DC42D11C5059BDEA" value="1991-05-22">22. V. 1991</collectingDate>
</date>
, 
<collectorName id="96B263802BDCF589AFB8D042C67C48CF">D. J. Larson</collectorName>
leg. ” (
<abbrev id="ABBRID0EACCI" xlink_title="Australian National Insect Collection">
<collectionCode id="CFFF5B3103938F0480B2AF0BD5ABB1CC" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
); 
<specimenCount id="F405216B97E7FE37FD704ABB00C982C2" count="6" type="generic">6 exs.</specimenCount>
, “ 
<collectingRegion id="33F50C302F0187BBF973742D382ADC18" country="Papua New Guinea" name="Madang">Madang</collectingRegion>
, 
<location id="8AF7ABAEF6BC06C4B2A45858D8D484F9" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:8AF7ABAEF6BC06C4B2A45858D8D484F9" country="Papua New Guinea" county="New Guinea" latitude="-5.133333" longLatPrecision="1305" longitude="145.76666" municipality="Rigo Lugio" name="Nagada River" stateProvince="Morobe">Nagada River</location>
near Nobanob, 
<date id="A22C6DE17363F02BF3A2DDA86094B16E" value="1991-06-12">
<collectingDate id="DAB0CF5CFB350D98E53447CDB6F419F1" value="1991-06-12">12. VI. 1991</collectingDate>
</date>
, 
<collectorName id="353AD45683B63B1B9316651A3810B25F">D. J. Larson</collectorName>
leg. ” (
<abbrev id="ABBRID0EECCI" xlink_title="Australian National Insect Collection">
<collectionCode id="6AF17BF9DA1F2172D12D3D47AC52B303" country="Australia" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection">ANIC</collectionCode>
</abbrev>
, 
<abbrev id="ABBRID0EICCI" xlink_title="Queensland Museum">
<collectionCode id="0EB3E8BF641742D189F70E33BA05308C" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" type="Museum">QM</collectionCode>
</abbrev>
); 
<specimenCount id="BC5E32F85FF237ECFCD5BF46A027269A" count="1" type="generic">1 ex.</specimenCount>
, “ New Guinea Biro 96 Friedrich-Wilhelmshafen ” (
<abbrev id="ABBRID0EMCCI" xlink_title="Természettudományi Múzeum">
<collectionCode id="2EF8C32BBC3D333700F1D3094E254AF5">TDMB</collectionCode>
</abbrev>
). • 
<emphasis id="DBEF2EEDF238ADD16E640FF40F53C2E5" bold="true">
<collectingCountry id="F65B0432AAB120103A6CEBF2D7711C31" name="Solomon Islands">Solomon Islands</collectingCountry>
</emphasis>
: 
<specimenCount id="6D668A77840722B2C1E0DA6FADF36BE6" count="1" type="generic">1 ex.</specimenCount>
, “ Solomon Islands: New Georgia, 2 mls. 
<collectorName id="911AF35D40FC93C8A8CED07A3E199680">W. Of Lamberte</collectorName>
1. IX. 65. Roy. Soc. Exped. B. M. 1966-1 ”, “ bomb crater pool, coast road ” (
<abbrev id="ABBRID0ESCCI" xlink_title="Natural History Museum, London">
<collectionCode id="8366CB88409D9BC469C4A84F1D21B55B" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
); 
<specimenCount id="643D3C042D2DE4862D3166CF5F74F261" count="3" type="generic">3 exs.</specimenCount>
(+ numerous larvae), “ SOLOMON ISLANDS, 
<collectingRegion id="D3D407515995DBF4B7FF7C3C3AD964D3" country="Solomon Islands" name="Guadalcanal">GUADALCANAL</collectingRegion>
, ca. 
<quantity id="625F3FD4A3AA5A407FF0FA31FC9EA2B0" metricMagnitude="3" metricUnit="m" metricValue="3.5" unit="km" value="3.5">3.5 km</quantity>
SE of BARANA vill., (drying up stream in shaded gorge), 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZCCI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[159.991667,-9.496667]}">
<geoCoordinate id="3A1AE623DC0487FB83EDE14FF5832758" degrees="09" direction="south" minutes="29.8" orientation="latitude" precision="92" value="-9.496667">09 ° 29.8 ' S</geoCoordinate>
, 
<geoCoordinate id="B826C00CF4AF4F072ADFF1EB7B2811A8" degrees="159" direction="east" minutes="59.5" orientation="longitude" precision="92" value="159.99167">159 ° 59.5 ' E</geoCoordinate>
</named-content>
; 
<quantity id="C02F652DE3F7A0D131879A02D2AD8779" metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">
<elevation id="DB802A725867053B78AC5488952A7ECD" metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">190 m</elevation>
</quantity>
, 
<date id="A025646262A5916073BC3D9F482A2C84" value="2013-11-24" valueMax="2013-12-14" valueMin="2013-11-24">
<collectingDate id="962F47C5E8F7AF73E66C49944B604CC9" value="2013-11-24" valueMax="2013-12-14" valueMin="2013-11-24">24. xi. - 14. xii. 2013</collectingDate>
</date>
, 
<collectorName id="8C2D50D7B6298BF2170B85F4D90C1DC6">Jiří Hájek</collectorName>
leg. ” (
<abbrev id="ABBRID0E5CCI" xlink_title="Národní muzeum, Praha">
<collectionCode id="9D7729755E353F3F235E94A97952B488" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
</abbrev>
); 
<specimenCount id="8C0CA266B5F444053FE46BB233B43769" count="3" type="generic">3 exs.</specimenCount>
, “ 
<collectingCounty id="41D9DCD3F585F5E38EBB8C1F78C0D6D4">South Pacific</collectingCounty>
, Solomon Is. 
<collectingRegion id="A4CFF086FF443AF2211E5C5D1765FA77" country="Solomon Islands" name="Guadalcanal">GUADALCANAL</collectingRegion>
I., 
<quantity id="A040953C337297EB78995BDCE753E63F" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" unit="m" value="575.0" valueMax="650.0" valueMin="500.0">
<elevation id="39F1FDD089EB0D173F0C8CCE76E745B8" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" unit="m" value="575.0" valueMax="650.0" valueMin="500.0">500–650 m</elevation>
</quantity>
<collectingMunicipality id="669EDBE33F52F85EC084FC8CE4204B7A">Koso</collectingMunicipality>
vill. Env., ca. 
<quantity id="C2E9C999D71FC4CFDFF93F066184C5D5" metricMagnitude="4" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" unit="km" value="16.5" valueMax="18.0" valueMin="15.0">15–18 km</quantity>
SSE of 
<collectingRegion id="F5BEAF390DF8616350273103C00F227C" country="Solomon Islands" name="Honiara">Honiara</collectingRegion>
, 
<date id="471C6A2B6886A2FB8DE049555A10F4E7" value="2016-12-01" valueMax="2016-12-18" valueMin="2016-12-01">
<collectingDate id="84124FAB2EEBF14F34B35BFC99659021" value="2016-12-01" valueMax="2016-12-18" valueMin="2016-12-01">1–18. XII. 2016</collectingDate>
</date>
<location id="F34D70DE0303E20DCB4315C247BCCFCE" LSID="urn:lsid:plazi:treatment:6B89ECAA735B50BA9E6ECAB068266314:F34D70DE0303E20DCB4315C247BCCFCE" country="Solomon Islands" county="South Pacific" latitude="-9.496667" longLatPrecision="129" longitude="159.99167" municipality="Koso" name="St. Jakl" stateProvince="Guadalcanal">
<collectorName id="C9CA4DF4FF7C096EFB41C1673FA805BF">St.</collectorName>
Jakl
</location>
leg. ” (
<abbrev id="ABBRID0ECDCI" xlink_title="Collection Lars Hendrich">
<collectionCode id="941B2360EE14D7698C96513D55829E44">CLH</collectionCode>
</abbrev>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGDCI" type="redescription">
<paragraph id="9C8475D003C763F4F0DC6A3D06ECF70B">
<heading id="41499D0B385D8D4818AD8B56D304ADC4" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="9337D20A24B8DF22E8452DACAA2A7F65">Body broadly oval, submatt / slightly shiny, black with ferrugineus brown markings. Ventral side completely dark brown to black, legs testaceous, hind legs somewhat darker.</paragraph>
<paragraph id="CD9DEBF162D616634B80863154EF7DE0">
Head testaceous with posterior part broadly black leaving free two small testaceous spots on vertex (Figs 
<figureCitation id="A047DDE028C8B42E5049585F76182499" captionStart="Figures 4–7" captionStartId="F2" captionText="Figures 4–7. Habitus of 4 Sandracottus dejeanii (South India) 5 S. festivus (Sri Lanka) 6 S. femoralis (Papua New Guinea, Rigo) 7 S. hunteri (Thailand)." figureDoi="10.3897/zookeys.1223.138220.figures4-7" httpUri="https://binary.pensoft.net/fig/1214409">6</figureCitation>
, 
<figureCitation id="3894FD1B3998B971475653F76CA816A3" captionStart="Figures 50–53" captionStartId="F19" captionText="Figures 50–53. Habitus of 50 Sandracottus dejeanii (India, Himachal Pradesh, Kangra Valley, Dharamsala) 51 S. festivus (Sri Lanka, Ratnapura, Sincharaja 52 S. femoralis (Indonesia, West Papua, Kapupaten Paniai, Nabire) 53 S. hunteri (China, Central Sichuan)." figureDoi="10.3897/zookeys.1223.138220.figures50-53" httpUri="https://binary.pensoft.net/fig/1214426">52</figureCitation>
). Surface submatt, shagreened, consisting of dense and minute punctures not very uniform in size and of larger and much sparser ones, more numerous on frons. Clypeal grooves and punctures alongside eyes marked, punctures medium-sized but coalescent. Antennae testaceous; antennomeres slender, antennomere V 4 × as long as broad.
</paragraph>
<paragraph id="03BCDDC9F86F8D5E14AA755174C931E7">
Pronotum black with lateral margins ferrugineus brown, and narrow, testaceous, horizontal band, interrupted medially (Figs 
<figureCitation id="3B0C0221B4432CB8C437B8DF4F7B3B63" captionStart="Figures 4–7" captionStartId="F2" captionText="Figures 4–7. Habitus of 4 Sandracottus dejeanii (South India) 5 S. festivus (Sri Lanka) 6 S. femoralis (Papua New Guinea, Rigo) 7 S. hunteri (Thailand)." figureDoi="10.3897/zookeys.1223.138220.figures4-7" httpUri="https://binary.pensoft.net/fig/1214409">6</figureCitation>
, 
<figureCitation id="6EFFB16CEDF12D6C5BEA4D6B5660EED0" captionStart="Figures 50–53" captionStartId="F19" captionText="Figures 50–53. Habitus of 50 Sandracottus dejeanii (India, Himachal Pradesh, Kangra Valley, Dharamsala) 51 S. festivus (Sri Lanka, Ratnapura, Sincharaja 52 S. femoralis (Indonesia, West Papua, Kapupaten Paniai, Nabire) 53 S. hunteri (China, Central Sichuan)." figureDoi="10.3897/zookeys.1223.138220.figures50-53" httpUri="https://binary.pensoft.net/fig/1214426">52</figureCitation>
). Surface submatt, distinctly shagreened, with very dense punctation; punctures medium-sized and distant of only 1–2 × their own diameter. Anterior puncture line interrupted in middle; punctures relatively small but coalescent, forming wrinkles and becoming sparse medially and lacking in mid-length of anterior margin. Posterior puncture line with coarse and coalescent punctures in middle of each side, forming distinct wrinkles, distinctly smaller and spaced on disc.
</paragraph>
<paragraph id="34369F39FCD1A5D31B31CCBB10C4BEC2">
Elytra black with small testaceous markings consisting of basal band sometimes reduced to different small spots at base, one postmedian spot, one preapical short and reduced band, and an apical spot (Figs 
<figureCitation id="B708DB71505614D6A31CB32721D2920B" captionStart="Figures 4–7" captionStartId="F2" captionText="Figures 4–7. Habitus of 4 Sandracottus dejeanii (South India) 5 S. festivus (Sri Lanka) 6 S. femoralis (Papua New Guinea, Rigo) 7 S. hunteri (Thailand)." figureDoi="10.3897/zookeys.1223.138220.figures4-7" httpUri="https://binary.pensoft.net/fig/1214409">6</figureCitation>
, 
<figureCitation id="B3F74E3A2B14D79D723E9F0DB4772405" captionStart="Figures 50–53" captionStartId="F19" captionText="Figures 50–53. Habitus of 50 Sandracottus dejeanii (India, Himachal Pradesh, Kangra Valley, Dharamsala) 51 S. festivus (Sri Lanka, Ratnapura, Sincharaja 52 S. femoralis (Indonesia, West Papua, Kapupaten Paniai, Nabire) 53 S. hunteri (China, Central Sichuan)." figureDoi="10.3897/zookeys.1223.138220.figures50-53" httpUri="https://binary.pensoft.net/fig/1214426">52</figureCitation>
), submatt. Epipleura ferrugineus brown. Surface distinctly shagreened and covered with small and dense punctures as well as with larger and much sparser ones. Puncture lines with groups of medium-sized punctures mostly grouped in five or six punctures, groups closer together on discal line. Sutural puncture line marked only by few punctures on apical part.
</paragraph>
<paragraph id="F909055B94E969C90A17C6518B8FFBF8">Ventral side dark brown. Legs, particularly fore and mid legs testaceous, hind legs ferrugineus brown to dark brown. Prosternal process short and broad, 1.3 × longer than broad, flattened, finely but distinctly sculptured; posterior margin broadly rounded. Metatibia with sparse medium-sized punctures on whole surface. Ventrites II – VI very superficially shagreened, slightly and longitudinally wrinkled on lateral parts, densely covered with very small punctures and larger sparser ones. Posterior margins rounded, deeply bordered with some large and coalescent punctures in middle of each side along margin.</paragraph>
<paragraph id="8AE2F0CF312856FE2A69044BC0E953AC">
Measurements: 
<abbrev id="ABBRID0EMECI" xlink_title="total length">TL</abbrev>
= 
<quantity id="53287701A03A7818923182A4E779696B" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.26" metricValueMin="1.19" unit="mm" value="12.25" valueMax="12.6" valueMin="11.9">11.9–12.6 mm</quantity>
, 
<abbrev id="ABBRID0EQECI" xlink_title="total length minus head length">TL-h</abbrev>
= 11.0– 
<quantity id="B5C76D948773CD6E8C71470CA2236151" metricMagnitude="-2" metricUnit="m" metricValue="1.28" unit="mm" value="12.8">12.8 mm</quantity>
, 
<abbrev id="ABBRID0EUECI" xlink_title="maximum width">TW</abbrev>
= 7.0– 
<quantity id="DC3E3B4319B6C10B874117A54F6F83DC" metricMagnitude="-3" metricUnit="m" metricValue="7.5" unit="mm" value="7.5">7.5 mm</quantity>
.
</paragraph>
<paragraph id="17207B4FB5307EDAE0C0D61CA4384354">
<specimenCount id="CAB897500FC1784B34865327DEAC2D1C" count="1" type="male">♂</specimenCount>
. Protarsomeres I – III strongly enlarged with three larger suckers and numerous smaller ones. Mesotarsomeres I – III with two rows of small suckers. Median lobe of aedeagus, in ventral view, broad, parallel-sided up to apex, here slightly broadened and broadly rounded (Fig. 
<figureCitation id="0E250F25881544FB992897766B695B0F" captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16–19. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b): 16 Sandracottus festivus 17 S. femoralis 18 S. hunteri and 19 S. insignis." figureDoi="10.3897/zookeys.1223.138220.figures16-19" httpUri="https://binary.pensoft.net/fig/1214412">17 a</figureCitation>
). Parameres broad and pointed at apex (Fig. 
<figureCitation id="CB725AB4357C2C1C107CC9CBDB7AACCB" captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16–19. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b): 16 Sandracottus festivus 17 S. femoralis 18 S. hunteri and 19 S. insignis." figureDoi="10.3897/zookeys.1223.138220.figures16-19" httpUri="https://binary.pensoft.net/fig/1214412">17 b</figureCitation>
).
</paragraph>
<paragraph id="FD2459912BDBD39D4C12BA38640A0A1D">
<specimenCount id="A1A3448811C7C4DFB7B92A262B07A8CA" count="1" type="female">♀</specimenCount>
. Similar to male, tarsi not enlarged. Microsculpture on ventrite VI as in male.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDFCI" type="Differential diagnosis">
<paragraph id="04666E7BA0DFDF53A787710A6271AAD8">
<heading id="5CBBA5758A577FC6815A1AC4FA8CCF71" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="CD7966DD0A4588786135F137B761E5FF">
The combination of size and dorsal colour pattern (Figs 
<figureCitation id="016C54090F4594001BAAB8274671CF81" captionStart="Figures 4–7" captionStartId="F2" captionText="Figures 4–7. Habitus of 4 Sandracottus dejeanii (South India) 5 S. festivus (Sri Lanka) 6 S. femoralis (Papua New Guinea, Rigo) 7 S. hunteri (Thailand)." figureDoi="10.3897/zookeys.1223.138220.figures4-7" httpUri="https://binary.pensoft.net/fig/1214409">6</figureCitation>
, 
<figureCitation id="C5293E8EC2B0C1EF09AA85DDAD55B3ED" captionStart="Figures 50–53" captionStartId="F19" captionText="Figures 50–53. Habitus of 50 Sandracottus dejeanii (India, Himachal Pradesh, Kangra Valley, Dharamsala) 51 S. festivus (Sri Lanka, Ratnapura, Sincharaja 52 S. femoralis (Indonesia, West Papua, Kapupaten Paniai, Nabire) 53 S. hunteri (China, Central Sichuan)." figureDoi="10.3897/zookeys.1223.138220.figures50-53" httpUri="https://binary.pensoft.net/fig/1214426">52</figureCitation>
) separates 
<taxonomicName id="C0E092341ED06AF947EF0CF5804BD821" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="40D1F530D064DD939117A93F7B26C093" italics="true">S. femoralis</emphasis>
</taxonomicName>
from all other species of the genus. The very similarly coloured 
<taxonomicName id="CFB8D06659429A24ADCBBDB448EA393B" authorityName="Hendrich &amp; Brancucci" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="F9375CC8D2724C9C559036CFCB70BDEE" italics="true">S. maculatus</emphasis>
</taxonomicName>
is always larger (
<abbrev id="ABBRID0EHGCI" xlink_title="total length">TL</abbrev>
= 14.0– 
<quantity id="1399B055C65BFE535FD3F6AD9013EB63" metricMagnitude="-2" metricUnit="m" metricValue="1.61" unit="mm" value="16.1">16.1 mm</quantity>
) and more broadly oval than 
<taxonomicName id="C5372199F334AB443C0E52184310CB4F" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="98D6C1449D52BA38428367996A92BB07" italics="true">S. femoralis</emphasis>
</taxonomicName>
(
<abbrev id="ABBRID0EWGCI" xlink_title="total length">TL</abbrev>
= 
<quantity id="32211E63B3B588F1BBB0C7F9B89CC021" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.26" metricValueMin="1.19" unit="mm" value="12.25" valueMax="12.6" valueMin="11.9">11.9–12.6 mm</quantity>
). Furthermore, 
<taxonomicName id="9B840B1B0286DDA2337D66FC8289BAB6" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="1DB03C28ECE271DC403FFDB47813E6C0" italics="true">S. femoralis</emphasis>
</taxonomicName>
can be separated from all other species of the genus by the shape of the median lobe and parameres (Fig. 
<figureCitation id="6DC3E1A2D32C7813580C3857DC03FCA6" captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16–19. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b): 16 Sandracottus festivus 17 S. femoralis 18 S. hunteri and 19 S. insignis." figureDoi="10.3897/zookeys.1223.138220.figures16-19" httpUri="https://binary.pensoft.net/fig/1214412">17 a, b</figureCitation>
). All instar larvae were described by 
<bibRefCitation id="153186CB0D62FBEF6FCE70788AED45D9" DOI="10.11646/zootaxa.5263.3.1" author="Alarie Y &amp; Michat M &amp; Shaverdo H &amp; Hájek J" etAl="et al." firstAuthor="Alarie" journalOrPublisher="Zootaxa" pagination="301-334" refId="B1" refString="Alarie Y, Michat M, Shaverdo H, Hájek J (2023) Morphology of the larvae of Sandracottus femoralis Heller, 1934, and S. mixtus (Blanchard, 1843) and phylogenetic comparison with other known Aciliini (Coleoptera: Dytiscidae, Dytiscinae). Zootaxa 5263: 301–334. https://doi.org/10.11646/zootaxa.5263.3.1" title="Morphology of the larvae of Sandracottus femoralis Heller, 1934, and S. mixtus (Blanchard, 1843) and phylogenetic comparison with other known Aciliini (Coleoptera: Dytiscidae, Dytiscinae)." volume="5263" year="2023">Alarie et al. (2023)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENHCI" type="distribution">
<paragraph id="3FB8837E2B72D8730C0B027900436349">
<heading id="E16B87234875E494BF6BFB04CF433FA8" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="0EF0EBD759F84AB482E40904DBE05F57">
<collectingCountry id="03720FBAEFAB7FA78320526E7BD24F83" name="Indonesia">Indonesia</collectingCountry>
: 
<collectingCountry id="76DACFCDFAD181597CA44A5CA1B022B7" name="Indonesia">Moluccas</collectingCountry>
(Bacan, Batanta, Halmahera, Obi, Seram, Morotai), Key and Aru Islands, and 
<collectingRegion id="C8B424BCCAFD382B9CC04455BD90AAD2" country="Indonesia" name="Papua Barat">West Papua</collectingRegion>
; 
<collectingCountry id="E9C01C9EA5772BCBFC30ED26ED8EEDCA" name="Papua New Guinea">Papua New Guinea</collectingCountry>
up to the 
<collectingCountry id="47DCE096AF49141DAADA25E8AF30E560" name="Solomon Islands">Solomon Islands</collectingCountry>
(Fig. 
<figureCitation id="8D40203961B3F097582A4990AA0881AA" captionStart="Figure 27" captionStartId="F11" captionText="Figure 27. Distribution of Sandracottus femoralis [black dots] in New Guinea and the Moluccas, and S. rotundus [red squares] on Sulawesi." figureDoi="10.3897/zookeys.1223.138220.figure27" httpUri="https://binary.pensoft.net/fig/1214418">27</figureCitation>
).
</paragraph>
<caption id="1BC77D0A1AD1B77F0DD2BC80A8F6035B" ID-DOI="10.3897/zookeys.1223.138220.figure27" ID-arpha="FE66CC56-BE0D-57C7-A045-0ADBE057284E" httpUri="https://binary.pensoft.net/fig/1214418" startId="F11">
<paragraph id="186F72DC9647D22E1DC4CF3B37A1D125">
<label id="5BC8A53E360730D1E0981A1830A09CD3">Figure 27.</label>
</paragraph>
<paragraph id="D908FC3EEE46C87047B53578F1A9A06C">
Distribution of 
<taxonomicName id="AC20F5895DCF2983C8097EBFF8973486" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="EFE40D96535FD001D8A35A7A32A3ABEA" italics="true">Sandracottus femoralis</emphasis>
</taxonomicName>
[black dots] in New Guinea and the Moluccas, and 
<taxonomicName id="37200C0B69590CFB8BA228C983ED6D80" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rotundus">
<emphasis id="EBB74DF6158A1D701EBFE8C2795B6319" italics="true">S. rotundus</emphasis>
</taxonomicName>
[red squares] on Sulawesi.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E1ICI" type="habitat">
<paragraph id="7CF63767B5FD198DD63224DE481714CD">
<heading id="4D8CBA5C573871C0138E612BB6462EC0" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="0FA4696DD2071CF843FC536E690AD922">
In West Papua specimens were collected in small but mainly exposed primary rainforest pools and puddles, always enriched with decaying leaves (Fig. 
<figureCitation id="BB359BA0262BFE4568A8DAF44610CC4A" captionStart="Figures 31–36" captionStartId="F15" captionText="Figures 31–36. Habitats of Sandracottus 31 Australia, Northern Territory, 50 km S Adelaide River, creek at Scenic Route (NT 9) and 32 South Queensland, Winfield, Winfield Road, rest pool (QLD 54), habitats of S. bakewellii bakewellii (photos: L. Hendrich) 33 Northern Territory, Ormiston Gorge, habitat of S. bakewellii guttatus (photo: M. Baehr) 34 Pakistan, Sistan va Baluchestan Prov., Pir Sohrab env., rest pool in wadi, habitat of S. dejeanii (photo: J. Hájek) 35 India, southern Madhya Pradesh, Hoshangabad District, Pagara – Pachmarhi road, ca 5 km NNE Panchmarhi, Panar Pani, habitat of S. dejeanii and S. festivus (photo: M. Jäch) 36 Indonesia, Irian Jaya, Nabire, Nabire-Ilaga track, km 62, habitat of S. femoralis (photo: L. Hendrich)." figureDoi="10.3897/zookeys.1223.138220.figures31-36" httpUri="https://binary.pensoft.net/fig/1214422">36</figureCitation>
). On Bacan Island also collected at light. On the 
<collectingCountry id="84A0D5591ADFE7D0739DA0905E2CAE62" name="Solomon Islands">Solomon Islands</collectingCountry>
found in an old water-filled bomb crater near a coastal road (New 
<collectingCountry id="438B7E2CFE24F3C525A9F6560000D780" name="Georgia">Georgia</collectingCountry>
), and a pool formed by the drying up of a stream on a clay bed, covered with decaying leaves and twigs (Guadalcanal; 
<bibRefCitation id="773F7D6742E087CA8A657D513FDE197F" DOI="10.11646/zootaxa.5263.3.1" author="Alarie Y &amp; Michat M &amp; Shaverdo H &amp; Hájek J" etAl="et al." firstAuthor="Alarie" journalOrPublisher="Zootaxa" page="320" pagination="301-334" refId="B1" refString="Alarie Y, Michat M, Shaverdo H, Hájek J (2023) Morphology of the larvae of Sandracottus femoralis Heller, 1934, and S. mixtus (Blanchard, 1843) and phylogenetic comparison with other known Aciliini (Coleoptera: Dytiscidae, Dytiscinae). Zootaxa 5263: 301–334. https://doi.org/10.11646/zootaxa.5263.3.1" title="Morphology of the larvae of Sandracottus femoralis Heller, 1934, and S. mixtus (Blanchard, 1843) and phylogenetic comparison with other known Aciliini (Coleoptera: Dytiscidae, Dytiscinae)." volume="5263" year="2023">Alarie et al. 2023: 320</bibRefCitation>
). 
<taxonomicName id="70963D7DB41AD93787B886C37258BBEF" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="42B15A16BB65DC443E4FD1AEAB5517D9" italics="true">Sandracottus femoralis</emphasis>
</taxonomicName>
is distributed from lowland forest up to hilly mountain rain forests at ~ 
<quantity id="EB9F2B5977E9889143F912A6C4E9A47A" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
altitude.
</paragraph>
<caption id="9572ED5F60858B2AE1CBD32CBC6A982F" ID-DOI="10.3897/zookeys.1223.138220.figure28" ID-arpha="B0F2084D-4768-57F1-91C1-42DFBEB30168" httpUri="https://binary.pensoft.net/fig/1214419" startId="F12">
<paragraph id="63CE4AEFA7CD244AC7654B7C5AC9E86D">
<label id="3C29D0F740CD2B15A0944A5C005974B6">Figure 28.</label>
</paragraph>
<paragraph id="A966F91D9170B2EF3AB677DDB2C9F95D">
Distribution of 
<taxonomicName id="A515C0141C33AF4638F4C4C62E344DE9" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="D25F21C5D2836E0DADAD81DEDDB6DB8B" italics="true">Sandracottus insignis</emphasis>
</taxonomicName>
in the Philippines.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>