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<mods:title id="B79FFA380E32F90B73AEBFA0DCF650DE">A revision of the Australian digger wasps in the genus Sphex (Hymenoptera, Sphecidae)</mods:title>
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<mods:namePart id="423AA6AB18CD6F7B9A8B6F6828998D56">Doerfel, Thorleif H.</mods:namePart>
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<mods:namePart id="1C14C43E16418F4CF3D140F14EB56D77">Ohl, Michael</mods:namePart>
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<treatment id="6B75A0E607ACCBE49EEB410B2CD32E68" ID-GBIF-Taxon="152062392" LSID="urn:lsid:plazi:treatment:6B75A0E607ACCBE49EEB410B2CD32E68" httpUri="http://treatment.plazi.org/id/6B75A0E607ACCBE49EEB410B2CD32E68" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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<paragraph id="8ECEDA1BAAA224270D2123A9DBE215C8" pageId="14" pageNumber="15">Taxon classification Animalia Hymenoptera Sphecidae</paragraph>
</subSubSection>
<subSubSection id="0DFDD2814AF0ED81C68361FE2826D5FC" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="40E575F65B8EBB0255D194C2AAD73B40" pageId="14" pageNumber="15">
<taxonomicName id="09D9B8D58596ECD1F9B5DE735FC822D0" ID-CoL="4YYGW" authority="van der Vecht, 1973" authorityName="van der Vecht" authorityYear="1973" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex carbonicolor" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carbonicolor">Sphex carbonicolor van der Vecht, 1973</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F71F09EB0FD572547348B346B30A46DD" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="605EE03363EF675AD16460ED63499464" pageId="14" pageNumber="15">
<taxonomicName id="8C88987FB45268D2F6E174A514263282" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex carbonarius" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carbonarius">Sphex carbonarius</taxonomicName>
F. Smith, 1856: 247, ♀ (as
<taxonomicName id="31D6DE98CB1D00114FCA8E1C5965E429" lsidName="carbonaria" pageId="14" pageNumber="15" rank="species" species="carbonaria">carbonaria</taxonomicName>
, incorrect original termination), junior primary homonym of
<taxonomicName id="BE66AC67AEA8F4853293FA4C52392ED5" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex carbonarius" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carbonarius">Sphex carbonarius</taxonomicName>
Scopoli, 1763. Holotype or syntypes: ♀, Australia: Sydney (BMNH). Not examined.
</paragraph>
<paragraph id="5AD7419D200F1ADD535648D5F6AFA7E2" pageId="14" pageNumber="15">
<taxonomicName id="1CDD81FB39D5B81AEC19666B60BDA91B" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex carbonicolor" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carbonicolor">Sphex carbonicolor</taxonomicName>
van der Vecht, 1973: 342 Substitute name for
<taxonomicName id="8CC2FE5D62ED4DF1D93A2102293FCC17" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex carbonarius" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carbonarius">Sphex carbonarius</taxonomicName>
F. Smith.
</paragraph>
</subSubSection>
<subSubSection id="8991F219B5EFE6C9479F73A1E625FBA9" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="2C7FFF44AE9450251A12FA24C13BDA5F" pageId="14" pageNumber="15">Material examined.</paragraph>
<paragraph id="9A36E596C55C3BBD87F2513C1194FCAD" pageId="14" pageNumber="15">
[COUNTRY UNKNOWN]:[state unknown]: [no specific locality], 2♀ (AMS). AUSTRALIA:NSW: Barraba, 1♀, March 37, M. Showers (AMS); Blue Mountains, 1♂, 08.01.1983, N. W. Rodd (AMS); Caldwell, 1♀, 1♂, 12.01.1952, V. Robb (AMS); Casula, 1♂, 11.02.1958, M. I. Nikitin (BMNH), 1♂, 24.02.1958, M. I. Nikitin (BMNH); 40 km E of Coonabarabran, 1♀, 18.11.1990, N. W. Rodd (AMS); Glen Innes, 1♀, 26.12.1979, A. W. Cameron (AMS); Tamarama near Sydney, 1♀, 19.12.2003 (AMS); NT: Central Australia, 1♀, 23.06.1927, G. Horns (ANIC); Finke Gorge National Park, Alice Springs, 1♀, 20.10.1973, G. Griffin (ANIC); QLD: Blackdown Tableland Expedition Range, 1♂, 08.01.1976, G. Daniels (AMS); Burnett River, 1♀, 1891/1892, R. Lennon (BMNH); Cairo Station, 1♀, 15.01.-31.01.1954, C. Mc.C. (ANIC); Captain Billy Landing, Cape York,
<geoCoordinate id="6D23769E5DBD4ACBE9B274B3CB8FB292" direction="south" orientation="latitude" precision="925" value="-11.633333">11°38'S</geoCoordinate>
,
<geoCoordinate id="6AE5F61D8FE445D64A2CBD57A9519F71" direction="east" orientation="longitude" precision="925" value="142.85">142°51'E</geoCoordinate>
, 1♂, 11.03.1992, G. Daniels &amp; M. A. Schneider (ANIC); Charters Towers, 1♀, 20.07.1902, W. W. Froggatt (ANIC); Division of Dawson, 1♀, Rothschild &amp; Bequest (BMNH); Kensington Downs, 2♀ (AMS); Rockhampton, 1♀ (ZMB); SA: Adelaide, 1♀, 08.01.1988, A. D. Austin (BMNH), 1♀, 1♂ (ZMB); Urrbrae, 1♀, 1944 (ANIC); Wilpena Pound Resort, Flinders Ranges, 1♂, 24.01.1995, L. Packer (ZMB); WA: 30 km N of Carnavon, Blow Holes Road, 1♂, 21.07.1978, G. A. Holloway (AMS); Carnarvon, 1♀, 01.08.1953, A. Snell (ANIC); Geraldton, 1♂, 1917, J. Clark (ANIC); Glen Forrest, 1♂, 11.12.1949, I. M. (ANIC); Lyndon Station, NW Basin, 1♀, 01.07.1950, G. Thomas (ANIC); Marloo Station, 1♂, 01.01.1935, Gebr. Goerling (ZMB), 1♂, 02.03.1935, A. Goerling (ZMB), 1♂, 01.02.1937, Gebr. Goerling (ZMB); Ongerup,
<geoCoordinate id="1DEFEB2362539B5F68D8E299FF58DB7F" direction="south" orientation="latitude" precision="92" value="-33.965">33°57.9'S</geoCoordinate>
,
<geoCoordinate id="09DF71278E399FC75D51ACDB4214EFE5" direction="east" orientation="longitude" precision="92" value="118.48">118°28.8'E</geoCoordinate>
, 1♂, 28.11.2008, D. M. Bray &amp; W. J. Pulawski (CAS).
</paragraph>
</subSubSection>
<subSubSection id="89C2FBDF7E575F3A3108A9D1EC7C8477" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="C3113BAEE391AC5DE8DA779DC33A3ECD" pageId="14" pageNumber="15">Diagnosis.</paragraph>
<paragraph id="B1DF9F0B0F6D7EED8CD90C3F25A5DEAF" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
This species differs from other members of the
<taxonomicName id="5E520908ADE23A48B3CC436FA8D793F9" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex argentatus" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="argentatus">Sphex argentatus</taxonomicName>
group in having distinct tubercles on the metanotum, combined with completely hyaline
<pageBreakToken id="55D206C2CFDFF2D77D744D78E115958F" pageId="15" pageNumber="16" start="start">wings</pageBreakToken>
except for a slight brown tinge at the apical margin of the forewing in some specimens. Specimens of
<taxonomicName id="5ADFAF63034BFE7B59F1B16EACEFA217" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex sericeus" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sericeus">Sphex sericeus</taxonomicName>
that may have the same color pattern are identifiable by the conspicuous transverse ridges on their propodeum, which are lacking in
<taxonomicName id="55629711BE1096FD50AD7A90066BE955" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex carbonicolor" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carbonicolor">Sphex carbonicolor</taxonomicName>
.
<taxonomicName id="3FA1A1BEF6B7B8FB192A6D07EDED3E36" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex argentatus" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="argentatus">Sphex argentatus</taxonomicName>
and
<taxonomicName id="83291DD2C5F0008BA67D73EB8D94E10F" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex finschii" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="finschii">Sphex finschii</taxonomicName>
have a darkened wing base, while
<taxonomicName id="F8DA580ABA16ABB84E735FEB35EBCE8E" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex decoratus" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="decoratus">Sphex decoratus</taxonomicName>
differs by having orange legs (which are dark brown or black in
<taxonomicName id="05C65C4F9860D5C3C5047C6B988B151E" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex carbonicolor" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carbonicolor">Sphex carbonicolor</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="87ACA24CD2EC255DC5D652D7FE1087BE" pageId="15" pageNumber="16" type="description">
<paragraph id="0912273B2C1834E9A3335D614645D2B6" pageId="15" pageNumber="16">Description.</paragraph>
<paragraph id="10770AB695CFCFE6EADB8D6DFEC2F672" pageId="15" pageNumber="16">
Body length 27.6-35.6 mm. Body black. Wing membrane hyaline, forewing with fuscous spot beyond marginal cell. Wing veins brown. Clypeus bulging directly above free margin. Appressed pubescence and erect setae on clypeus and frons silvery. Clypeus with medial glabrous stripe. Distance between hind- ocelli nearly equal to their shortest distance to compound eyes. Pubescence on collar and scutum silvery, on scutum denser laterally than medially. Scutellum slightly convex, with medial impression. Tubercles on metanotum distinct. Propodeal enclosure densely covered with long, erect silvery-white setae, leaving sculpture visible. Length of petiole 1.1
<normalizedToken id="3CF10068B29A55C2DA830E53A7231552" originalValue="×">x</normalizedToken>
length of flagellomere II. Tomentum moderately dense on metasomal tergum I and II, but very short.
</paragraph>
<paragraph id="780264B10A949CD432DC3B1FAE635D48" pageId="15" pageNumber="16">Female: Foretarsal rake with 12 long spines. Free clypeal margin with small notch medially.</paragraph>
<paragraph id="D1A440398C065F340FFF268B685DCC43" pageId="15" pageNumber="16">
Male: Free clypeal margin entire. Metasomal sterna
<normalizedToken id="0139A17BE84D3676A5D8BB306FE9247C" originalValue="IIVII">II-VII</normalizedToken>
mostly glabrous. Metasomal terga V and VI with few bristles. Metasomal sternum VIII notched apically, its lateral margin straight.
</paragraph>
<caption id="F4423DBCB47EFFD65E2A28CD52BC5CDA" pageId="15" pageNumber="16">
<paragraph id="6C96A22C33A37B32AEFE987602538B32" pageId="15" pageNumber="16">
Figure 9.
<taxonomicName id="26C5AFF5DAF662E1D3A1DE49F11EA601" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex carbonicolor" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carbonicolor">Sphex carbonicolor</taxonomicName>
. A ♀, habitus B collecting localities.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="01F6F6DB26C458EC18852E22F8C9D7B2" pageId="15" pageNumber="16" type="type material">
<paragraph id="EBEFE36C9AFDEAF0DD203C9B2C6411D1" pageId="15" pageNumber="16">Notes on type material.</paragraph>
<paragraph id="522E3EE714220DEDA1AFD10C4FD477B3" pageId="15" pageNumber="16">
The type of
<taxonomicName id="83DF74FEE96FC2DA8BD40D6CB192B2C2" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex carbonicolor" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="carbonicolor">Sphex carbonicolor</taxonomicName>
was not examined, because the character combination listed in the original description (black body, a glabrous area on the clypeus, a bituberculate metanotum, hyaline wings) is almost unique.
<taxonomicName id="C68BD27200DFF0A8E4BCF862975204DC" class="Insecta" family="Sphecidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex ephippium" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ephippium">Sphex ephippium</taxonomicName>
, which shares these features, is sufficiently differentiated by its character combination (see below).
</paragraph>
</subSubSection>
</treatment>
</document>