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<document id="4D4257FFFE123DFB59A7F7BB769B9CF0" ID-DOI="10.11646/zootaxa.3640.2.9" ID-GBIF-Dataset="ef11d479-21ce-4271-866f-9709cb37ce28" ID-ISSN="1175-5326" ID-Zenodo-Dep="218121" ID-ZooBank="57EFDD02-CDD5-4A13-B723-54506D5E11DA" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460593274570" checkinUser="plazi" docAuthor="Brown, John W." docDate="2013" docId="6B2887DE1F70FFA3F6D52689E287F9B7" docLanguage="en" docName="zt03640p283.pdf" docOrigin="Zootaxa 3640 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eupinivora unicolora Brown, 2013, new species" docType="treatment" docVersion="6" lastPageNumber="280" masterDocId="9711FFA61F7AFFA9F6422755E61EFFB1" masterDocTitle="A new genus of pine-feeding Cochylina from the western United States and northern Mexico (Lepidoptera: Tortricidae: Euliini)" masterLastPageNumber="283" masterPageNumber="270" pageNumber="280" updateTime="1698326356857" updateUser="plazi">
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<mods:title id="64D84D2EC1F0CED70DA804E9A565C298">A new genus of pine-feeding Cochylina from the western United States and northern Mexico (Lepidoptera: Tortricidae: Euliini)</mods:title>
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<mods:namePart id="ABF1A60173BB9BA533F04894E63212FB">Brown, John W.</mods:namePart>
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<mods:title id="ACAADC2396012EA1A65F8721E9CDDCEE">Zootaxa</mods:title>
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<mods:part id="227C89B58362B1F4804E2F23B4831830">
<mods:date id="48602678DDDF1C35A9A395778E480DCE">2013</mods:date>
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<mods:number id="FE0BD07822F8AF1F34E076573E2EEB31">3640</mods:number>
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<mods:classification id="9B44D8D852D0017AD680FF60937B8980">journal article</mods:classification>
<mods:identifier id="4781C0D74185D40E0172B552491C3CF9" type="DOI">10.11646/zootaxa.3640.2.9</mods:identifier>
<mods:identifier id="A64F7BFA6CD2D1290BA721F32EBCF057" type="GBIF-Dataset">ef11d479-21ce-4271-866f-9709cb37ce28</mods:identifier>
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<mods:identifier id="13294089CFA8B1AF3BF58168BA908186" type="Zenodo-Dep">218121</mods:identifier>
<mods:identifier id="AD33BDBED955FA8D4F28FB29BE4E7594" type="ZooBank">57EFDD02-CDD5-4A13-B723-54506D5E11DA</mods:identifier>
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<treatment id="6B2887DE1F70FFA3F6D52689E287F9B7" ID-DOI="http://doi.org/10.5281/zenodo.6153362" ID-GBIF-Taxon="119566164" ID-Zenodo-Dep="6153362" LSID="urn:lsid:plazi:treatment:6B2887DE1F70FFA3F6D52689E287F9B7" httpUri="http://treatment.plazi.org/id/6B2887DE1F70FFA3F6D52689E287F9B7" lastPageNumber="280" pageId="10" pageNumber="280">
<subSubSection id="AB9B65431F70FFA3F6D52689E729FDA7" pageId="10" pageNumber="280" type="nomenclature">
<paragraph id="E33E36C81F70FFA3F6D52689E425FE44" blockId="10.[151,571,475,534]" box="[151,571,475,502]" pageId="10" pageNumber="280">
<heading id="B87681A41F70FFA3F6D52689E425FE44" bold="true" box="[151,571,475,502]" fontSize="11" level="1" pageId="10" pageNumber="280" reason="1">
<emphasis id="D1F5EADA1F70FFA3F6D52689E425FE44" bold="true" box="[151,571,475,502]" pageId="10" pageNumber="280">
<taxonomicName id="24814D4B1F70FFA3F6D52689E782FE47" ID-CoL="96TVF" box="[151,412,476,502]" class="Insecta" family="Tortricidae" genus="Eupinivora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="280" phylum="Arthropoda" rank="species" species="unicolora" status="sp. nov.">
<emphasis id="D1F5EADA1F70FFA3F6D52689E782FE47" bold="true" box="[151,412,476,502]" italics="true" pageId="10" pageNumber="280">Eupinivora unicolora</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="CAC657A11F70FFA3F7E9268EE425FE44" box="[427,571,475,501]" pageId="10" pageNumber="280" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E33E36C81F70FFA3F6D526ABE729FDA7" blockId="10.[151,571,475,534]" box="[151,311,510,534]" pageId="10" pageNumber="280">
<figureCitation id="7BBA2A4D1F70FFA3F6D526ABE6FBFDA7" box="[151,229,510,534]" captionStart="FIGURES 3 8" captionStartId="5.[151,264,1920,1943]" captionTargetBox="[151,1433,193,1880]" captionTargetId="figure@5.[151,1436,193,1899]" captionTargetPageId="5" captionText="FIGURES 3 8. Adults of Eupinivora. 3. E. ponderosae, paratype male, USA, Arizona. 4. E. angulicosta, holotype male, Mexico, Nuevo Leon. 5. E. albolineana, holotype male, Mexico, Durango. 6. E. hamartopenis, paratype male, Mexico, Durango. 7. E. unicolora, holotype male, Mexico, Durango. 8. E. rufofascia, paratype female, Mexico, Durango." httpUri="https://zenodo.org/record/218123/files/figure.png" pageId="10" pageNumber="280">Figs. 7</figureCitation>
,
<figureCitation id="7BBA2A4D1F70FFA3F6B326ABE711FDA7" box="[241,271,510,534]" captionStart="FIGURES 9 14" captionStartId="6.[151,264,1912,1935]" captionTargetBox="[231,1366,186,1882]" captionTargetId="figure@6.[209,1385,177,1905]" captionTargetPageId="6" captionText="FIGURES 9 14. Male genitalia of Eupinivora, with phallus removed. 9. E. ponderosae, paratype, USA, Arizona (MGP slide 736). 10. E. angulicosta, holotype, Mexico, Nuevo Leon (USNM slide 92,089). 11. E. albolineana, paratype, Mexico, Durango (USNM slide 119,189). 12. E. hamartopenis, paratype, Durango (MGP slide 345). 13. E. unicolora, paratype, Mexico, Durango (USNM slide 142,194). 14. E. rufofascia, paratype, Mexico, Durango (USNM slide 142,207)." httpUri="https://zenodo.org/record/218124/files/figure.png" pageId="10" pageNumber="280">13</figureCitation>
,
<figureCitation id="7BBA2A4D1F70FFA3F75926ABE729FDA7" box="[283,311,510,534]" captionStart="FIGURES 15 18" captionStartId="8.[151,264,1834,1857]" captionTargetBox="[152,1433,334,1834]" captionTargetId="figure@8.[151,1436,331,1838]" captionTargetPageId="8" captionText="FIGURES 15 18. Female genitalia of Eupinivora. 15. E. ponderosae, paratype, USA, Arizona (USNM slide 23,462). 16. E. hamartopenis, paratype, Mexico, Durango (USNM slide 119,175). 17. E. unicolora, paratype, Mexico, Durango (USNM slide 142,195). 18. E. rufofascia, paratype, Mexico, Durango (USNM slide 142,208)." httpUri="https://zenodo.org/record/218125/files/figure.png" pageId="10" pageNumber="280">17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="AB9B65431F70FFA3F6D52513E74EFD5E" pageId="10" pageNumber="280" type="diagnosis">
<paragraph id="E33E36C81F70FFA3F6D52513E74EFD5E" blockId="10.[151,1436,582,1543]" pageId="10" pageNumber="280">
<emphasis id="D1F5EADA1F70FFA3F6D52513E714FDEE" bold="true" box="[151,266,582,607]" pageId="10" pageNumber="280">Diagnosis</emphasis>
.
<taxonomicName id="24814D4B1F70FFA3F75C2512E408FDEF" box="[286,534,582,606]" class="Insecta" family="Tortricidae" genus="Eupinivora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="280" phylum="Arthropoda" rank="species" species="unicolora">
<emphasis id="D1F5EADA1F70FFA3F75C2512E408FDEF" box="[286,534,582,606]" italics="true" pageId="10" pageNumber="280">Eupinivora unicolora</emphasis>
</taxonomicName>
is easily distinguished from its congeners by the nearly uniform dark rust forewing. The male genitalia are most similar to those of
<taxonomicName id="24814D4B1F70FFA3F57D2539E5E5FD32" box="[831,1019,619,643]" class="Insecta" family="Tortricidae" genus="Eupinivora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="280" phylum="Arthropoda" rank="species" species="hamartopenis">
<emphasis id="D1F5EADA1F70FFA3F57D2539E5E5FD32" box="[831,1019,619,643]" italics="true" pageId="10" pageNumber="280">E. hamartopenis</emphasis>
</taxonomicName>
, especially in the apically bifurcate transtilla, densely spined sacculus, and overall shape of the valva, but those of
<taxonomicName id="24814D4B1F70FFA3F24625C5E291FD17" box="[1028,1167,654,679]" class="Insecta" family="Tortricidae" genus="Eupinivora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="280" phylum="Arthropoda" rank="species" species="unicolora">
<emphasis id="D1F5EADA1F70FFA3F24625C5E291FD17" box="[1028,1167,654,679]" italics="true" pageId="10" pageNumber="280">E. unicolora</emphasis>
</taxonomicName>
are easily distinguished by a long phallus (ca. 1.5 times the length of the valva) that lacks the characteristic external spines present in all other congeners.
</paragraph>
</subSubSection>
<subSubSection id="AB9B65431F70FFA3F68525AFE4F7FAC7" pageId="10" pageNumber="280" type="description">
<paragraph id="E33E36C81F70FFA3F68525AFE224FB73" blockId="10.[151,1436,582,1543]" pageId="10" pageNumber="280">
<emphasis id="D1F5EADA1F70FFA3F68525AFE74FFCA2" bold="true" box="[199,337,762,787]" pageId="10" pageNumber="280">Description</emphasis>
. Head: Vertex and frons pale cream; scaling on antenna cream; labial palpus cream medially, pale orange laterally. Thorax: Nota pale cream-orange, paler at prothoracic collar, tegula rust-orange; legs mostly pale orange brown. Forewing length 7.09.0 (mean = 8.1; n = 6) in males, 8.09.0 (mean = 8.5; n = 2) in females; forewing (
<figureCitation id="7BBA2A4D1F70FFA3F7492433E750FCCF" box="[267,334,870,894]" captionStart="FIGURES 3 8" captionStartId="5.[151,264,1920,1943]" captionTargetBox="[151,1433,193,1880]" captionTargetId="figure@5.[151,1436,193,1899]" captionTargetPageId="5" captionText="FIGURES 3 8. Adults of Eupinivora. 3. E. ponderosae, paratype male, USA, Arizona. 4. E. angulicosta, holotype male, Mexico, Nuevo Leon. 5. E. albolineana, holotype male, Mexico, Durango. 6. E. hamartopenis, paratype male, Mexico, Durango. 7. E. unicolora, holotype male, Mexico, Durango. 8. E. rufofascia, paratype female, Mexico, Durango." httpUri="https://zenodo.org/record/218123/files/figure.png" pageId="10" pageNumber="280">Fig. 7</figureCitation>
) nearly uniform rust, a narrow, ill-defined, cream blotch in distal portion of discal cell, and diffuse area of cream scales at mid-termen. Fringe cream-orange. Hindwing pale grayish brown. Fringe pale gray mixed with cream. Abdomen: Cream-orange. Male genitalia (
<figureCitation id="7BBA2A4D1F70FFA3F55924FBE56FFC77" box="[795,881,942,966]" captionStart="FIGURES 9 14" captionStartId="6.[151,264,1912,1935]" captionTargetBox="[231,1366,186,1882]" captionTargetId="figure@6.[209,1385,177,1905]" captionTargetPageId="6" captionText="FIGURES 9 14. Male genitalia of Eupinivora, with phallus removed. 9. E. ponderosae, paratype, USA, Arizona (MGP slide 736). 10. E. angulicosta, holotype, Mexico, Nuevo Leon (USNM slide 92,089). 11. E. albolineana, paratype, Mexico, Durango (USNM slide 119,189). 12. E. hamartopenis, paratype, Durango (MGP slide 345). 13. E. unicolora, paratype, Mexico, Durango (USNM slide 142,194). 14. E. rufofascia, paratype, Mexico, Durango (USNM slide 142,207)." httpUri="https://zenodo.org/record/218124/files/figure.png" pageId="10" pageNumber="280">Fig. 13</figureCitation>
) with uncus subrectangular, short; socii broad, pendant, attached basally to tegumen by tiny sclerotized rod, creating the appearance of a small bifurcate uncus; median part of transtilla large, rectangular-rounded, with a pair of tiny teeth at middle; valva subrectangular, sacculus strongly sclerotized, with dense long, spines of variable length; phallus ca. 1.5 times length of valva, paired terminal processes long and evenly curved, lacking patch of spines near junction of aedeagus and phallobase. Female genitalia (
<figureCitation id="7BBA2A4D1F70FFA3F7AC2337E45CFBCB" box="[494,578,1122,1146]" captionStart="FIGURES 15 18" captionStartId="8.[151,264,1834,1857]" captionTargetBox="[152,1433,334,1834]" captionTargetId="figure@8.[151,1436,331,1838]" captionTargetPageId="8" captionText="FIGURES 15 18. Female genitalia of Eupinivora. 15. E. ponderosae, paratype, USA, Arizona (USNM slide 23,462). 16. E. hamartopenis, paratype, Mexico, Durango (USNM slide 119,175). 17. E. unicolora, paratype, Mexico, Durango (USNM slide 142,195). 18. E. rufofascia, paratype, Mexico, Durango (USNM slide 142,208)." httpUri="https://zenodo.org/record/218125/files/figure.png" pageId="10" pageNumber="280">Fig. 17</figureCitation>
) with disc-shaped antevaginal sclerite and two broadly rectanglular lateral posterior lobes; lamellae postvaginalis with a narrow sclerotized band; ductus bursae short, not differentiated from corpus bursae, ductus seminalis lateral, originating before middle of corpus bursae.
</paragraph>
<paragraph id="E33E36C81F70FFA3F685239BE23BFB57" blockId="10.[151,1436,582,1543]" box="[199,1061,1230,1255]" pageId="10" pageNumber="280">
<typeStatus id="3C3A886A1F70FFA3F685239BE731FB57" box="[199,303,1230,1254]" pageId="10" pageNumber="280" type="holotype">Holotype</typeStatus>
3,
<collectingCountry id="9B9676581F70FFA3F71B239BE7AFFB57" box="[345,433,1230,1254]" name="Mexico" pageId="10" pageNumber="280">Mexico</collectingCountry>
, Durango, El Salto,
<date id="973F10081F70FFA3F4D6239BE532FB57" box="[660,812,1230,1255]" pageId="10" pageNumber="280" value="1984-05-30">30 May 1984</date>
. Deposited in USNM.
</paragraph>
<paragraph id="E33E36C81F70FFA3F68523A6E4F7FAC7" blockId="10.[151,1436,582,1543]" pageId="10" pageNumber="280">
<typeStatus id="3C3A886A1F70FFA3F68523A6E72AFABA" box="[199,308,1267,1291]" pageId="10" pageNumber="280" type="paratype">Paratypes</typeStatus>
(183, 3Ƥ).
<collectingCountry id="9B9676581F70FFA3F7FC23A7E408FABB" box="[446,534,1266,1290]" name="Mexico" pageId="10" pageNumber="280">Mexico</collectingCountry>
, Durango, La Ciudad,
<date id="973F10081F70FFA3F55423A7E5B2FABB" box="[790,940,1266,1291]" pageId="10" pageNumber="280" value="1984-05-20">20 May 1984</date>
(13)
<date id="973F10081F70FFA3F5B223A7E275FABB" box="[1008,1131,1266,1290]" pageId="10" pageNumber="280" value="1984-06-02">2 Jun 1984</date>
(13),
<date id="973F10081F70FFA3F2F423A7E32CFABB" box="[1206,1330,1266,1290]" pageId="10" pageNumber="280" value="1984-06-06">6 Jun 1984</date>
(13), r.f.
<taxonomicName id="24814D4B1F70FFA3F6D52242E72DFA9F" box="[151,307,1303,1326]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="280" phylum="Tracheophyta" rank="species" species="cooperi">
<emphasis id="D1F5EADA1F70FFA3F6D52242E72DFA9F" box="[151,307,1303,1326]" italics="true" pageId="10" pageNumber="280">Pinus cooperi</emphasis>
</taxonomicName>
, M. E. Guerrero (USNM). La Ciudad, Pueblo Nuevo,
<date id="973F10081F70FFA3F5CC2242E5E6FA9F" box="[910,1016,1302,1327]" pageId="10" pageNumber="280" value="1984-04">Apr 1984</date>
(13), M. E. Perez (USNM). El Salto,
<date id="973F10081F70FFA3F6D5226FE738FAE3" box="[151,294,1338,1363]" pageId="10" pageNumber="280" value="1985-04-15">15 Apr 1985</date>
(23),
<date id="973F10081F70FFA3F731226FE41CFAE3" box="[371,514,1338,1363]" pageId="10" pageNumber="280" value="1985-04-27">27 Apr 1985</date>
(13, 1Ƥ),
<date id="973F10081F70FFA3F4C3226FE506FAE3" box="[641,792,1338,1363]" pageId="10" pageNumber="280" value="1984-05-30">30 May 1984</date>
(43),
<date id="973F10081F70FFA3F526226FE5E2FAE3" box="[868,1020,1338,1363]" pageId="10" pageNumber="280" value="1986-05-27">27 May 1986</date>
(33),
<date id="973F10081F70FFA3F20A226FE2DBFAE3" box="[1096,1221,1338,1362]" pageId="10" pageNumber="280" value="1985-06-09">9 Jun 1985</date>
(13),
<date id="973F10081F70FFA3F353226FE382FAE3" box="[1297,1436,1338,1362]" pageId="10" pageNumber="280" value="1986-06-23">23 Jun 1986</date>
(23, 2Ƥ),
<date id="973F10081F70FFA3F751220BE785FAC7" box="[275,411,1374,1398]" pageId="10" pageNumber="280" value="1986-06-27">27 Jun 1986</date>
(13) [no collector] (USNM).
</paragraph>
</subSubSection>
<subSubSection id="AB9B65431F70FFA3F68522D7E783FA53" pageId="10" pageNumber="280" type="distribution">
<paragraph id="E33E36C81F70FFA3F68522D7E783FA53" blockId="10.[151,1436,582,1543]" pageId="10" pageNumber="280">
<emphasis id="D1F5EADA1F70FFA3F68522D7E7F7FA2A" bold="true" box="[199,489,1410,1435]" pageId="10" pageNumber="280">Distribution and biology</emphasis>
.
<taxonomicName id="24814D4B1F70FFA3F7BA22D1E4F4FA2A" box="[504,746,1411,1435]" class="Insecta" family="Tortricidae" genus="Eupinivora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="280" phylum="Arthropoda" rank="species" species="unicolora">
<emphasis id="D1F5EADA1F70FFA3F7BA22D1E4F4FA2A" box="[504,746,1411,1435]" italics="true" pageId="10" pageNumber="280">Eupinivora unicolora</emphasis>
</taxonomicName>
is known only from La Ciudad and El Salto in the mountains of Durango,
<collectingCountry id="9B9676581F70FFA3F76C22F3E799FA0F" box="[302,391,1446,1470]" name="Mexico" pageId="10" pageNumber="280">Mexico</collectingCountry>
. At least three of the
<typeStatus id="3C3A886A1F70FFA3F4D322F2E4E2FA0E" box="[657,764,1447,1471]" pageId="10" pageNumber="280" type="paratype">paratypes</typeStatus>
were reared from
<taxonomicName id="24814D4B1F70FFA3F59F22F2E28CFA0F" box="[989,1170,1447,1470]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="280" phylum="Tracheophyta" rank="species" species="arizonica">
<emphasis id="D1F5EADA1F70FFA3F59F22F2E28CFA0F" box="[989,1170,1447,1470]" italics="true" pageId="10" pageNumber="280">Pinus arizonica</emphasis>
</taxonomicName>
<taxonomicName id="24814D4B1F70FFA3F2DF22F2E336FA0F" box="[1181,1320,1447,1470]" pageId="10" pageNumber="280" rank="variety" variety="copperi">
var.
<emphasis id="D1F5EADA1F70FFA3F29022F2E336FA0F" box="[1234,1320,1447,1470]" italics="true" pageId="10" pageNumber="280">copperi</emphasis>
</taxonomicName>
, and it is suspected that all were.
</paragraph>
</subSubSection>
<subSubSection id="AB9B65431F70FFA3F68522BAE287F9B7" box="[199,1177,1518,1543]" pageId="10" pageNumber="280" type="etymology">
<paragraph id="E33E36C81F70FFA3F68522BAE287F9B7" blockId="10.[151,1436,582,1543]" box="[199,1177,1518,1543]" pageId="10" pageNumber="280">
<emphasis id="D1F5EADA1F70FFA3F68522BAE758F9B6" bold="true" box="[199,326,1519,1543]" pageId="10" pageNumber="280">Etymology</emphasis>
. The specific epithet refers to the nearly uniform rust color of the forewing.
</paragraph>
</subSubSection>
</treatment>
</document>