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<mods:title id="ABBEB4571269B4D861B3F63C69A520BA">The genus Cephaloleia Chevrolat, 1836 (Coleoptera, Chrysomelidae, Cassidinae)</mods:title>
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<mods:namePart id="C4065FF9A1A26DED992FB05593CB5FD5">Staines, Charles L.</mods:namePart>
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<mods:namePart id="58380BA579C307D71868EDED0DCE56D5">Garcia-Robledo, Carlos</mods:namePart>
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<mods:identifier id="306D5CA92E6ADC5FD9D00F70B435DBAE" type="DOI">http://dx.doi.org/10.3897/zookeys.436.5766</mods:identifier>
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<treatment id="6ADCB2680BBF172837CB6AC3FD21C99C" ID-GBIF-Taxon="152055275" LSID="urn:lsid:plazi:treatment:6ADCB2680BBF172837CB6AC3FD21C99C" httpUri="http://treatment.plazi.org/id/6ADCB2680BBF172837CB6AC3FD21C99C" lastPageId="240" lastPageNumber="240" pageId="239" pageNumber="240" scope_family="Chrysomelidae" scope_order="Coleoptera" scope_subFamily="Cassidinae">
<subSubSection id="DEAFCAA86B595B3739FDF7F2970DE98A" pageId="239" pageNumber="240" type="multiple">
<paragraph id="BC1401DED6BFC7745B36623D3800A32B" pageId="239" pageNumber="240">
<pageBreakToken id="0F866925DFE786C8B44B4E7086A68A39" pageId="239" pageNumber="240" start="start">Taxon</pageBreakToken>
classification Animalia Coleoptera Chrysomelidae
</paragraph>
</subSubSection>
<subSubSection id="3D0AF59C3D7E32F0F644A9F9A9620AB0" pageId="239" pageNumber="240" type="nomenclature">
<paragraph id="F8D53A300009EC410E49E37C604BE203" pageId="239" pageNumber="240">
<taxonomicName id="D6CEF74EF49CF04EFCAFD8FF01E84650" ID-CoL="SDGL" LSID="http://species-id.net/wiki/Cephaloleia_schmidti" authority="Uhmann, 1933" authorityName="Uhmann" authorityYear="1933" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia schmidti" order="Coleoptera" pageId="239" pageNumber="240" phylum="Arthropoda" rank="species" species="schmidti">Cephaloleia schmidti Uhmann, 1933</taxonomicName>
Fig. 232
</paragraph>
</subSubSection>
<subSubSection id="91ECE936C501143FC75A07332718FBFC" pageId="239" pageNumber="240" type="reference_group">
<paragraph id="BB136F343022D0923EBA36E4BA76CC4F" pageId="239" pageNumber="240">
<taxonomicName id="73CDFDC71E9E4C02259B656ABE0BBD24" genus="Cephalolia" lsidName="Cephalolia schmidti" pageId="239" pageNumber="240" rank="species" species="schmidti">Cephalolia schmidti</taxonomicName>
<bibRefCitation id="6085848F64CFF77F38CF99B10F4CF253" author="Uhmann, E" journalOrPublisher="Entomologische Blaetter" pageId="295" pageNumber="296" pagination="168 - 169" title="Eine neue Cephalolia-Art aus Costa Rica. 42. Beitrag zur Kenntnis der Hispinen (Col. Chrys.)." volume="29" year="1933">Uhmann 1933</bibRefCitation>
: 168.
</paragraph>
<paragraph id="F28FB27793F50A30A38256F1CC9CBA02" pageId="239" pageNumber="240">
<taxonomicName id="F6D588C41DE2BF06F2268F87A6CE1CA3" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia schmidti" order="Coleoptera" pageId="239" pageNumber="240" phylum="Arthropoda" rank="species" species="schmidti">Cephaloleia schmidti</taxonomicName>
Uhmann.
<bibRefCitation id="7A481D03F5975A62DD9B4D28DA23A37E" author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="284" pageNumber="285" pagination="551 - 763" title="Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1946">Blackwelder 1946</bibRefCitation>
: 720 (catalog);
<bibRefCitation id="D8F6E6C4D1D624CA637D0B6C93AA22DB" author="Papp, CS" journalOrPublisher="Portugaliae Acta Biologica (B)" pageId="291" pageNumber="292" pagination="1 - 147" title="The Hispinae of America. 3 rd Contribution for promoting the scientifical results of the International Hylean Amazon Institute in Manaos, Brazil." volume="4" year="1953">Papp 1953</bibRefCitation>
: 22 (catalog);
<bibRefCitation id="0CDC362776A5C7BFE0B801A6C82B94B4" author="Uhmann, E" journalOrPublisher="W. Junk, Gravenhage" pageId="296" pageNumber="297" pagination="1 - 153" title="Coleopterorum Catalogus. Supplementa. Chrysomelidae: Hispinae, Hispinae Americanae" year="1957 a">Uhmann 1957a</bibRefCitation>
: 25 (catalog);
<bibRefCitation id="916BB046AACBB997233ED910C1725108" author="Descarpentries, A" journalOrPublisher="Bulletin du Museum d'Histoire Naturelle Paris" pageId="286" pageNumber="287" pagination="137 - 154" title="Les types de la collection M. Pic. 1. Chrysomelinae et Hispinae du Nouveau-Monde." volume="31" year="1959">Descarpentries and Villiers 1959</bibRefCitation>
a: 139 (types);
<bibRefCitation id="70C0EE6DBE38A57AA7FD8C55E71B30B2" author="Gaedike, R" journalOrPublisher="Beitraege zur Entomologie" pageId="287" pageNumber="288" pagination="341 - 395" title="Katalog der in den Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen-VII (Coleoptera: Hispinae)." volume="21" year="1971">
Gaedike and
<normalizedToken id="9FF00F771D48B6BD901ED672E531C43F" originalValue="Döbler">Doebler</normalizedToken>
1971
</bibRefCitation>
: 359 (types);
<bibRefCitation id="4F929D24C66DFC647A056D6087CA67E6" author="Wilcox, JA" journalOrPublisher="Biological Research Institute of America, Latham, NY" pageId="297" pageNumber="298" title="Checklist of the beetles of Canada, United States, Mexico, Central America and the West Indies" year="1983">Wilcox 1983</bibRefCitation>
: 137 (catalog);
<bibRefCitation id="8CCDB813F5CB1C4867E48E8DA7DD2530" pageId="239" pageNumber="240">Staines 1996</bibRefCitation>
: 58 (Central America species),
<bibRefCitation id="C9EDBFB4B2E2BA4484310C241A8F4218" pageId="239" pageNumber="240">1997</bibRefCitation>
: 414 (Uhmann species list);
<bibRefCitation id="710205DD54A87CDE2650F32E6A8C924A" pageId="239" pageNumber="240">Staines and Staines 1997</bibRefCitation>
: 20 (types);
<bibRefCitation id="0D6E6E1FA8B0F368E067C29957029E62" author="McKenna, DD" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="290" pageNumber="291" pagination="117 - 131" title="Molecular phylogenetics and evolution of host plant use in the Neotropical rolled leaf ' hispine' beetle genus Cephaloleia (Chevrolat) (Coleoptera: Cassidinae)." url="10.1016/j.ympev.2005.06.011" volume="37" year="2005">McKenna and Farrell 2005</bibRefCitation>
: 121 (phylogeny),
<bibRefCitation id="8C543EF2A84D57671F8DC3B6EA1BD73B" author="McKenna, DD" journalOrPublisher="Proceedings of the National Academy of Sciences" pageId="290" pageNumber="291" pagination="10947 - 10951" title="Tropical forests are both evolutionary cradles and museums of leaf beetle diversity." url="10.1073/pnas.0602712103" volume="103" year="2006">2006</bibRefCitation>
: 10949 (phylogeny).
</paragraph>
</subSubSection>
<subSubSection id="B9BEE0BF01AAC5176B21B7076DF38EE3" pageId="239" pageNumber="240" type="description">
<paragraph id="8F73A7B659DAD80078629261C5004416" pageId="239" pageNumber="240">Description.</paragraph>
<paragraph id="2A88057A6CD7D8A387344D62EAEB0D51" pageId="239" pageNumber="240">
Narrowly elongate; subparallel; subconvex; head, antennae, scutellum, venter, and legs black, pronotum reddish, elytra with basal
<normalizedToken id="24CB245A51880567D392CA3EE79FB3DF" originalValue="½">1/2</normalizedToken>
reddish-brown with a black sutural vitta, apical
<normalizedToken id="B95907BE24583C1584042ECAAD417058" originalValue="½">1/2</normalizedToken>
black. Head: vertex densely punctate, medial sulcus absent; frons not projecting; keel present between antennal bases; depressed between eyes. Antenna: reaches to humerus; slender; antennomere 1 transverse, laterally compressed; 2 transverse, shortest; 3-4 elongate, cylindrical; 3 elongate, as long as 1 and 2 combined; 4
<normalizedToken id="2666AF28D4C312BF1F3A3918F03C7BC2" originalValue="¾">3/4</normalizedToken>
length of 3; 5-10 transverse, subequal in length; 11 2
<normalizedToken id="57A8084AA4C524203D337F40B88F523D" originalValue="×">x</normalizedToken>
length of 10, pointed at apex; 1-2 punctate with scattered setae; 3-11 setose. Pronotum: transverse; lateral margin sinuate then rounding to anterior angle, canaliculate; anterior angle acute, produced; posterior angle acute; anterior margin emarginate behind head; disc subconvex; surface deeply, sparsely, irregularly punctate; basal impression absent; pronotal length 0.9-1.0 mm; pronotal width 1.1-1.4 mm. Scutellum: pentagonal; slightly punctate. Elytron: lateral margin straight, smooth, margined; apex rounded, sparsely serrate; sutural angle without tooth; humerus rounded, not produced; slightly constricted behind humerus; moderately punctate-striate, rows converge and unite apically; puncture row 10 removed from lateral margin; elytral length 3.8-4.0 mm; elytral width 1.8-2.0 mm. Venter: pro-, meso-, and metasterna impunctate medially, punctate laterally; abdominal sterna punctate, each puncture with pale seta; suture between sterna 1 and 2 complete. Leg: slender; femur punctate at apex; tibia with fringe of setae on inner margin of apex. Total length: 5.0-5.3 mm.
</paragraph>
</subSubSection>
<subSubSection id="7C99BFA3707AE833BD18F93EA2E0307F" pageId="239" pageNumber="240" type="diagnosis">
<paragraph id="0DC408155F60D76A735DA0DD9853342F" pageId="239" pageNumber="240">Diagnosis.</paragraph>
<paragraph id="644F942510CC956EAD82AE4E7735F768" pageId="239" pageNumber="240">
This species is similar to
<taxonomicName id="E7020F97427563CB2CCAD28AC0A19D0B" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia atriceps" order="Coleoptera" pageId="239" pageNumber="240" phylum="Arthropoda" rank="species" species="atriceps">Cephaloleia atriceps</taxonomicName>
and
<taxonomicName id="4F3B8870F94E3C81D60AAC9760FC57CA" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia ruficollis" order="Coleoptera" pageId="239" pageNumber="240" phylum="Arthropoda" rank="species" species="ruficollis">Cephaloleia ruficollis</taxonomicName>
. It can be distinguished by the elytra being reddish-brown at the base with a black sutural vitta, by the vertex of the head being depressed between the eyes, and by the suture between abdominal sterna 1 and 2 being complete.
</paragraph>
</subSubSection>
<subSubSection id="FFB0B09ADE3D17235EA3ACEE4C8463B6" pageId="239" pageNumber="240" type="distribution">
<paragraph id="D4E8125E7BC7FA1375BAF245A75CBA5C" pageId="239" pageNumber="240">Distribution.</paragraph>
<paragraph id="446894AE81B804C6DE981CA13E064E7D" pageId="239" pageNumber="240">Costa Rica, Panama.</paragraph>
</subSubSection>
<subSubSection id="DF91EC4973B1E022314753C15352DC51" pageId="239" pageNumber="240" type="type material examined">
<paragraph id="8F37D6AC67B341F63663FDE208C09B67" pageId="239" pageNumber="240">Type material examined.</paragraph>
<paragraph id="1CEA3BE1B9A170781318E3CAE65A5F10" pageId="239" pageNumber="240">Paratype: Costa Rica, F. Nevermann [green label]/ La Caja, B. San Jose, Schmidt leg. [reversed green label]/ Paratyp [red label]/ Cephalolia schmidti Uh., E. Uhmann det. 33/ Paratype No. 54638 USNM [orange label] (USNM).</paragraph>
</subSubSection>
<subSubSection id="DEEC9414D8AAF2E4BA5BBF3646FDF269" lastPageId="240" lastPageNumber="241" pageId="239" pageNumber="240" type="specimens examined">
<paragraph id="9BE92E0A93B4DC2858537091AE19BCEE" pageId="239" pageNumber="240">Specimens examined.</paragraph>
<paragraph id="FB59336389B35CA4ACF859014BE5FCA7" lastPageId="240" lastPageNumber="241" pageId="239" pageNumber="240">
COSTA RICA: Alajuela- 20 km S Upala, 22-31 May 1991 (BYUC). Cartago-
<normalizedToken id="20B8E77D7A8AE5727239C4980D523E94" originalValue="Limón">Limon</normalizedToken>
border, 500 m, 40 km NE Turrialba, 18 May 1979 (CMNC). Guanacaste- 3 km SE R. Naranjo, 1-5 June 199- (BYUC);
<normalizedToken id="8D81B41BC3791297B311FFF3D991C045" originalValue="Volcán">Volcan</normalizedToken>
Miravelles Geothermal Area, 3 July 1991 (EMEC). Puntarenas- Monteverde, 1400 m, 23 May 1979
<pageBreakToken id="179587B4A2F888048AE695D0350722E9" pageId="240" pageNumber="241" start="start">(</pageBreakToken>
CMNC); 4-6 km S Sta Elena, June 4-7 1980 (EGRC); Garabito, Reserva Biol Carara,
<normalizedToken id="8141CA00D28E6CFEFCA6A4F37B3C3069" originalValue="Estación">Estacion</normalizedToken>
Quebrada Bonita, 0-100 m (INBIO). San
<normalizedToken id="BC5E1F6556A24C2D55942CCE13FE1A12" originalValue="José-">Jose-</normalizedToken>
La Caja, S. San
<normalizedToken id="25AD39C27DFFC2AD35B8B6B47384FD26" originalValue="José">Jose</normalizedToken>
(USNM). PANAMA:
<normalizedToken id="E21CB368B735F75A6E8927A80F9BB6F7" originalValue="Panamá-">Panama-</normalizedToken>
Cerro Campana, 11-15 May 1980 (EGRC). Total: 12.
</paragraph>
</subSubSection>
</treatment>
</document>