305 lines
39 KiB
XML
305 lines
39 KiB
XML
<document ID-CLB-Dataset="21464" ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="6AC79A167563596F81F932D26E56B9E7" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole volontany Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="007D00C1-ED9C-462D-B6F2-F566494EE806" docUuidSource="ZooBank" docVersion="6" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1732651982216" updateUser="ExternalLinkService">
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<mods:titleInfo id="81C2ED8299D45F3E048C879EE9F19762">
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<mods:title id="5B0163F078A42374EBA36183FDD9EE07">Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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<mods:name id="0370D69474925765B430CC97BABEE0DF" type="personal">
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<mods:roleTerm id="E8CAC86BA4A576B15EF675818A1D2C6C">Author</mods:roleTerm>
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<mods:namePart id="B70C109607D3F478C5DF6395D5C52475">Salata, Sebastian</mods:namePart>
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<mods:affiliation id="4B978D45EF8A51B515DC12BDFCFC812D">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="584AA32851D957CD783555141ABC9757" type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier id="457E23D7B63937DF2215ABE17FEAFCD6" type="email">sdsalata@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E926E271B4781E9D3FE7E52E30F15918">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="F154B12FE700706B082B79B9B20E4EDD">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="627ABB8B6434EE08988ED95A082E82F7" type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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<mods:title id="11387873DD80B4BD4E0EB925E86F12DB">ZooKeys</mods:title>
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<mods:date id="DC7481ABAB4C439E4DD6D4AFD48FA5EE">2020</mods:date>
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<mods:detail id="B026F1664DFE91C7F10C8C4FAE16ADC2" type="volume">
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<mods:number id="35E3ECEB057A652C64359387D525823A">949</mods:number>
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<mods:url id="C149BE3494714F2B3AC49DE8133F8BC0">http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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<mods:classification id="EC2F183CA446F49B1F91066C05702A2E">journal article</mods:classification>
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<mods:identifier id="6829EAAEDC392798F9058D05066A9273" type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<treatment id="6AC79A167563596F81F932D26E56B9E7" ID-GBIF-Taxon="165090179" LSID="urn:lsid:zoobank.org:act:007D00C1-ED9C-462D-B6F2-F566494EE806" httpUri="http://treatment.plazi.org/id/6AC79A167563596F81F932D26E56B9E7" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="A9F8ED8534A060763BC4862376DA2E02" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="996D6845491CBFCF97DCE4C0EBF92A34" pageId="0" pageNumber="1">
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<taxonomicName id="A76584D44F25592CC80E0201536E4147" ID-CoL="CMM3X" LSID="6AC79A16-7563-596F-81F9-32D26E56B9E7" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole volontany" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="volontany" status="sp. nov.">Pheidole volontany</taxonomicName>
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<taxonomicNameLabel id="05FC02608D8E090592982A97D20A346B" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="40713B31A4855BF708F5FB70DE06A713" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">Figs 61A-F</figureCitation>
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<figureCitation id="1C311E9585382ACDDF74D8912EC2FBE2" captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">, 64S</figureCitation>
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<figureCitation id="2FB761AB0A56C5AFFB1D461B971562AE" captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Distribution. Pheidole renirano sp. nov. (A). Pheidole sava sp. nov. (B). Pheidole sikorae Forel (C). Pheidole sofia sp. nov. (D). Pheidole sparsa sp. nov. (E). Pheidole tampony sp. nov. (F). Pheidole trichotos sp. nov. (G). Pheidole tsaravoniana sp. nov. (H). Pheidole vadum sp. nov. (I). Pheidole veteratrix Forel (J). Pheidole volontany sp. nov. (K). Pheidole vony sp. nov. (L)." figureDoi="10.3897/zookeys.949.51269.figure67" httpUri="https://binary.pensoft.net/fig/432236" pageId="0" pageNumber="1">, 67K</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4D709D32B76D431C4E2A3B9FFAE409A1" pageId="0" pageNumber="1" type="type material">
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<paragraph id="699B66F63BAC7016947A13182A64F663" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="CFCEED64A4A0FEC874B46BAA0EA12C78" pageId="0" pageNumber="1">
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<emphasis id="6AD64DECB6982EEDF2FFE8792403A49C" bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Toliara;
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<normalizedToken id="4C3511661B2A6C037624810539CD4892" originalValue="Forêt">Foret</normalizedToken>
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||
<normalizedToken id="CF5ECD166960E149EE2A4EE6AB8AB423" originalValue="Classée">Classee</normalizedToken>
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<normalizedToken id="5B06EE477146D49D961DA8AFA544B3A0" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
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, 33.2 km 344°NNW Mahaboboka;
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<geoCoordinate id="1681CFCB615C33E70DC58CF0075B720C" degrees="22.64333" direction="south" orientation="latitude" precision="1" value="-22.64333">-22.64333</geoCoordinate>
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,
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<geoCoordinate id="F3BE149EEAA92399C23DF3E0EB69BEC1" degrees="44.17167" direction="east" orientation="longitude" precision="1" value="44.17167">44.17167</geoCoordinate>
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; alt. 1300 m; 22 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07935; CASENT0491861, bottom specimen on the pin (CASC).
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<emphasis id="3CE7CD86E236FC07B8FE98F2552BFC22" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
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• 9w., 3s.; same data as for holotype; CASENT0491860, CASENT0491863, CASENT0491862, CASENT0491864, CASENT0872247 (CASC, MHNG, PBZT).
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</paragraph>
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<caption id="79180D1DE57CE83643B1F78ED1016228" doi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1" start="Figure 61" startId="F61">
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<paragraph id="D0149F50A18FA8425133A6E1B247FAFE" pageId="0" pageNumber="1">
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<emphasis id="5A4F614D550D773E3BCF1F156D4567A0" bold="true" pageId="0" pageNumber="1">Figure 61.</emphasis>
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<taxonomicName id="2F1BF0E5AEE0EF0550584803EF464D19" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole volontany" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="volontany">
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<emphasis id="D2E459EFAA1FAC7AFB3A95C6280EDEC1" italics="true" pageId="0" pageNumber="1">Pheidole volontany</emphasis>
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</taxonomicName>
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Forel, full-face view (
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<emphasis id="BBE4CF9B17CD8E0CBBBB8B552F8F9A35" bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis id="923DB4F5E2DCE0677574FE9A4165A9AF" bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis id="C002B1D2514DEAD9340D294F8D16F8C9" bold="true" pageId="0" pageNumber="1">E</emphasis>
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||
) of minor paratype worker (CASENT0491862) and full-face view (
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||
<emphasis id="740FE0D0302236B322C4EFEEA308D054" bold="true" pageId="0" pageNumber="1">B</emphasis>
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||
), profile (
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||
<emphasis id="C6E50476EB5A9224AEC2C5A26B2A445F" bold="true" pageId="0" pageNumber="1">D</emphasis>
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||
), and dorsal view (
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||
<emphasis id="B8E0EC3B1BD84EC8541299F8298F0980" bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0491861).
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</paragraph>
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</caption>
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</subSubSection>
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||
<subSubSection id="D92E74EAD1E3DE97D4D7787154431702" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="170614C4CE90E81EB25935F2FFD9939B" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="2CE8D4ED16AE5E3E51D34D4A2445F164" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="8BE0730A4A05C2C2D11E3946C01F2D22" bold="true" pageId="0" pageNumber="1">Toliara</emphasis>
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: • 3w., 2s.;
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<normalizedToken id="E9DA913166432DA532DA09C409E96163" originalValue="Forêt">Foret</normalizedToken>
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||
<normalizedToken id="04FE6CA7F9F0DEA7A68C6DAE5B019311" originalValue="Classée">Classee</normalizedToken>
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||
<normalizedToken id="332E8089AEA99506F3FB376B0CB11ADB" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
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||
, 33.2 km 344°NNW Mahaboboka;
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||
<geoCoordinate id="53776E692C5320EE5838D547170297C8" degrees="22.64333" direction="south" orientation="latitude" precision="1" value="-22.64333">-22.64333</geoCoordinate>
|
||
,
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||
<geoCoordinate id="F362C5F48DA5FBE17348BDEAB4E2CBBA" degrees="44.17167" direction="east" orientation="longitude" precision="1" value="44.17167">44.17167</geoCoordinate>
|
||
; alt. 1300 m; 22 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07925 (CASC). • 3w.;
|
||
<normalizedToken id="152016BA6BE8D0F37FFDDB2F4BA793FE" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="C7A1D1D5F5CAC046EFC921A45979AB24" originalValue="Classée">Classee</normalizedToken>
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||
<normalizedToken id="F227D65801490BAD74F400CF00149A58" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
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||
, 33.2 km 344°NNW Mahaboboka;
|
||
<geoCoordinate id="5B8EDAFA45C5762587F5EA1E5AA51E69" degrees="22.64333" direction="south" orientation="latitude" precision="1" value="-22.64333">-22.64333</geoCoordinate>
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,
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<geoCoordinate id="3830859D4BB456060898C5F8544A3F12" degrees="44.17167" direction="east" orientation="longitude" precision="1" value="44.17167">44.17167</geoCoordinate>
|
||
; alt. 1300 m; 22 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07926 (CASC). • 3w., 2s.;
|
||
<normalizedToken id="3EEE540D7F04C095D2509420413BFFAC" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="D7C7D6CC5E9AA579E6FBB26853374CCC" originalValue="Classée">Classee</normalizedToken>
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||
<normalizedToken id="E937CEFF7CFA917C1753FDA43046CD13" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
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||
, 33.2 km 344°NNW Mahaboboka;
|
||
<geoCoordinate id="59CA50654989A9CB34158E80D19D5F94" degrees="22.64333" direction="south" orientation="latitude" precision="1" value="-22.64333">-22.64333</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F13CDF08F93C7894B21033E52EE54E50" degrees="44.17167" direction="east" orientation="longitude" precision="1" value="44.17167">44.17167</geoCoordinate>
|
||
; alt. 1300 m; 22 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07927 (CASC). • 1w.;
|
||
<normalizedToken id="1B91B36EA82BD3887D9C0DEA7A59A6DF" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="6818F73CA36C5044669085D739324C41" originalValue="Classée">Classee</normalizedToken>
|
||
<normalizedToken id="AE73D021DEB0C1A2402E745404F9863C" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||
, 33.2 km 344°NNW Mahaboboka;
|
||
<geoCoordinate id="0B7707B0098E10C83A7FE1E18E9ADB6B" degrees="22.64333" direction="south" orientation="latitude" precision="1" value="-22.64333">-22.64333</geoCoordinate>
|
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,
|
||
<geoCoordinate id="46D2DD8329E13F8BA0B840D03E08E17C" degrees="44.17167" direction="east" orientation="longitude" precision="1" value="44.17167">44.17167</geoCoordinate>
|
||
; alt. 1300 m; 22 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07928 (CASC). • 6w., 3s.;
|
||
<normalizedToken id="F4DC725324D46DDFCB67649983D4A530" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="4BEAF894DC1EBF261398E2B1BD1F6834" originalValue="Classée">Classee</normalizedToken>
|
||
<normalizedToken id="E4DEA86894CB603F68E807F6A93FDC64" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||
, 33.2 km 344°NNW Mahaboboka;
|
||
<geoCoordinate id="3A502C962418FFE0BC659735EB1980FF" degrees="22.64333" direction="south" orientation="latitude" precision="1" value="-22.64333">-22.64333</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0D4B109AFBC69B64D4FE709A85B3FD04" degrees="44.17167" direction="east" orientation="longitude" precision="1" value="44.17167">44.17167</geoCoordinate>
|
||
; alt. 1300 m; 22 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07931 (CASC). • 6w.;
|
||
<normalizedToken id="07CAA36489D6E5F55CEDD83A9142CC78" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="3D046018682E981C8B44D5A745BE5820" originalValue="Classée">Classee</normalizedToken>
|
||
<normalizedToken id="2BB276092907A05DC9EE367DF0A1F200" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||
, 33.2 km 344°NNW Mahaboboka;
|
||
<geoCoordinate id="4C4B60EAE245C9CA669C5418C35F2EF3" degrees="22.64333" direction="south" orientation="latitude" precision="1" value="-22.64333">-22.64333</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1E0FA5C56F8D1785516861F1A02BB65F" degrees="44.17167" direction="east" orientation="longitude" precision="1" value="44.17167">44.17167</geoCoordinate>
|
||
; alt. 1300 m; 22 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07939 (CASC). • 3w., 1s.;
|
||
<normalizedToken id="656961B40C07C03B37A4952F49DC5246" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="9525E69EC662B2F18074C8A4432FE32D" originalValue="Classée">Classee</normalizedToken>
|
||
<normalizedToken id="4E1F9ED3CC418B1C9B5ECF80D36ACBF4" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||
, 33.2 km 344°NNW Mahaboboka;
|
||
<geoCoordinate id="9A042BDAF7CBBF44C6DC6501E3F7E14E" degrees="22.64333" direction="south" orientation="latitude" precision="1" value="-22.64333">-22.64333</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="27B19476061F2D3192A401F7A79CC857" degrees="44.17167" direction="east" orientation="longitude" precision="1" value="44.17167">44.17167</geoCoordinate>
|
||
; alt. 1300 m; 22 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07952 (CASC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="943FD0454B5771E058547305C5348209" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="49D711B262A654505A7750A6E2A03D21" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="C8AF9781F5F8F1779159CE30DE5FA373" pageId="0" pageNumber="1">
|
||
Minute species.
|
||
<emphasis id="C747BB2CFBBC8B422F73D01A0DEBCB4D" bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||
HL <1.05 mm and WL <0.9 mm; head in full-face view sub-oval, slightly widening posteriorly, with anterior and posterior sides convex, in lateral view sub-oval; ventral and dorsal faces convex; body dark brown; sides of head with dense, very long, suberect to erect pilosity; entire head distinctly sculptured, medial part of frons with moderately dense, thick, longitudinal, and interrupted rugae, interspaces shiny and smooth; scape, when laid back, exceeding the midlength of head by one-fifth of its length; promesonotal dorsum with rugofoveolae and additional thick transverse to irregular rugae; lateral sides of pronotum, anepisternum, katepisternum, and propodeum with dense and moderately thick to thin rugoreticulae; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed slightly inward; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth, and approximately the same height, apex directed outward; inner and outer hypostomal teeth closely spaced and connected by concavity; base of first gastral tergite smooth.
|
||
<emphasis id="5CE1C3DBF5D5EF1B4607911C7C6A5DAA" bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
HL <0.5 mm and WL <0.6 mm, scape, when laid back, surpassing the posterior head margin by one-fifth of its length; propodeal spines minute, triangular; head relatively oval, smooth, and shiny with a few short, longitudinal rugae on the anterolateral sides of head; body brown; mesosoma smooth, indistinct foveolae can occur on propodeal dorsum and lateral sides of pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0AB2648A0BC17034D522B911F8C74038" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="FA297873F2F300E0B96B34DA3C7C43A1" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="AEF86452613BD4E742DC7C17282CFCAE" pageId="0" pageNumber="1">
|
||
<emphasis id="2BED69EE41368611434E84A5C37B41A5" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="8A6EB680A9E3962D2952BEAD16BD219E" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.91-1.05 (0.98); HW: 0.97-1.06 (1.02); SL: 0.55-0.6 (0.57); EL: 0.12-0.15 (0.14); WL: 0.82-0.87 (0.85); PSL: 0.13-0.18 (0.15); MTL: 0.51-0.56 (0.53); PNW: 0.43-0.5 (0.46); PTW: 0.14-0.16 (0.15); PPW: 0.26-0.34 (0.3); CI: 93.0-101.0 (96.2); SI: 51.4-61.6 (56.3); PSLI: 13.3-17.7 (14.9); PPI: 44.0-58.0 (50.2); PNI: 44.0-58.0 (50.2); MTI: 49.2-53.6 (51.9).
|
||
</paragraph>
|
||
<paragraph id="24ECAB83A57AE57297332B1E9617EB52" pageId="0" pageNumber="1">
|
||
<emphasis id="AB37A454518A5E2E8869AC68159456CA" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
In full-face view sub-oval, slightly widening posteriorly, with anterior and posterior sides convex (Fig.
|
||
<figureCitation id="2F731F947EB48DCA256AB0C5534473AF" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61B</figureCitation>
|
||
). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with dense, very long, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with moderately dense, thick, longitudinal and interrupted rugae, interspaces shiny and smooth; anterolateral sides with longitudinal, thick, and dense rugae; interspaces shiny with relatively dense and distinct rugofoveolae; posterolateral sides with thick, dense, and irregular rugae with relatively dense and distinct rugofoveolae. Occipital lobes with sparse, thick, and irregular rugae; interspaces rugofoveolate. Gena with relatively dense, thick, longitudinal rugae and rugofoveolate interspaces. Area posterolateral from eyes shiny, with dense, thick rugofoveolae and smooth interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by one-fifth of its length; pilosity subdecumbent to erect (Fig.
|
||
<figureCitation id="5C3D5FAE171458B6792BFBEF7D2DA4EC" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61B, D</figureCitation>
|
||
). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed slightly inward; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth closely spaced and connected by concavity (Fig.
|
||
<figureCitation id="90731AE908686E0E598F7F09D7DE8D44" captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">64S</figureCitation>
|
||
).
|
||
<emphasis id="FFA304F5BC97BEBA8247191C4B48C770" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like or absent; promesonotal groove absent; metanotal groove absent; propodeal spines moderately short, with wide base and acute apex; humeral area laterally weakly produced (Fig.
|
||
<figureCitation id="EA62AD6CD2DA714F788F05CB3393CBED" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61D</figureCitation>
|
||
). Surface shiny; promesonotal dorsum with rugofoveolae and additional thick transverse to irregular rugae; lateral sides of pronotum, anepisternum, katepisternum, and propodeum with dense and moderately thick to thin rugoreticulae. Pilosity sparse, long, and erect (Fig.
|
||
<figureCitation id="5DB9C43771BC3230F73DEE1E3E453A99" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61D, F</figureCitation>
|
||
).
|
||
<emphasis id="D86F78EA804A55795B9E60F2AF3995EB" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
|
||
Shiny with fine foveolae; node smooth to finely foveolate, low, triangular, with rounded and thin apex, in rear view node dorsoventrally straight to slightly convex; pilosity moderately sparse and erect (Fig.
|
||
<figureCitation id="C12795A3CA4E5D664F0C00D5A019A9F3" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61D, F</figureCitation>
|
||
).
|
||
<emphasis id="27D935E14A051FB98C0E70EA4D7A1403" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
|
||
Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
|
||
<figureCitation id="EFB6DD5A510A7CBA07157FC0705AA6B3" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61D, F</figureCitation>
|
||
).
|
||
<emphasis id="B21E41C0FC70B1549120E69A14C1C60F" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
Shiny and smooth; pilosity dense, long, and erect (Fig.
|
||
<figureCitation id="F21B567E9FCD73FF830104DED6C240F0" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61D, F</figureCitation>
|
||
).
|
||
<emphasis id="19B30E8D392A02BC5A02F70FDF074329" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Dark brown; legs and antennae yellow (Fig.
|
||
<figureCitation id="5293F55BA949495C2ACF633543EDC9FC" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="48C20EC054FB40532F8662B4232BDF5A" pageId="0" pageNumber="1">
|
||
<emphasis id="F9C215FD486B4555C5AC3C25AE4C24B3" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="585BE99C518F83FDBD6450CAA2975EA7" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.48-0.51 (0.5); HW: 0.44-0.46 (0.45); SL: 0.47-0.51 (0.49); EL: 0.09-0.11 (0.1); WL: 0.57-0.62 (0.59); PSL: 0.06-0.08 (0.07); MTL: 0.37-0.39 (0.38); PNW: 0.27-0.31 (0.29); PTW: 0.06-0.08 (0.07); PPW: 0.11-0.13 (0.12); CI: 108.8-116.0 (112.0); SI: 105.4-111.4 (108.4); PSLI: 11.2-14.6 (13.0); PPI: 56.9-70.2 (62.7); PNI: 61.4-68.6 (65.3); MTI: 81.3-88.2 (84.3).
|
||
</paragraph>
|
||
<paragraph id="090E2B0603110F4C011A58AAD2C5FA95" pageId="0" pageNumber="1">
|
||
<emphasis id="F0F62E6E66DCFBD21BE881C69BD918A6" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
Cephalic margin indistinctly concave or straight (Fig.
|
||
<figureCitation id="07F969C3C394844E0217BAC97D77DC02" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61A</figureCitation>
|
||
). Pilosity relatively sparse, long, decumbent to suberect. Sculpture smooth and shiny with a few short, longitudinal rugae on the anterolateral sides of head; antennal sockets with few thick, curved outward rugae and smooth interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity dense, subdecumbent to erect (Fig.
|
||
<figureCitation id="F2225475350BB7230D8350B533839D6E" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61A, C</figureCitation>
|
||
).
|
||
<emphasis id="A77AABCC69BDD5A566343A88DC2C2FFB" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum moderately high and short, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines minute, triangular (Fig.
|
||
<figureCitation id="B5FF49D2737C57F74B2AED1FF744B2C6" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61C</figureCitation>
|
||
). Sculpture shiny and smooth; indistinct foveolae can occur on propodeal dorsum and lateral sides of pronotum. Pilosity very sparse, moderately long, and erect (Fig.
|
||
<figureCitation id="ECEFD6FBA0724410E49502E396BF2B92" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61C, E</figureCitation>
|
||
).
|
||
<emphasis id="655F21F84D97B400036173744AF8079D" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
|
||
Short, low, and relatively flat; with few short, erect setae (Fig.
|
||
<figureCitation id="59C19E5EE0BF253209EF0B30B9DAA980" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61C, E</figureCitation>
|
||
).
|
||
<emphasis id="8C5BC483C202EF37F7554CBFF0301308" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
With sparse, erect pilosity (Fig.
|
||
<figureCitation id="2E3329FB72C84D1DA3D563DDC01E4013" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61C, E</figureCitation>
|
||
).
|
||
<emphasis id="98E25E21C7B9B16435D3B124147510E9" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Brown, legs and antenna yellow (Fig.
|
||
<figureCitation id="ECB0481C0EE5BC584D1C7E44EFA7521B" captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole volontany Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0491862) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491861)." figureDoi="10.3897/zookeys.949.51269.figure61" httpUri="https://binary.pensoft.net/fig/432230" pageId="0" pageNumber="1">61C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3BB75C6AA85B0BDE41448D29BC7D8044" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="7E9FB511DFC27AC8F5C51E9948DA8E2B" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="2BA928526507BEFB3B2BF27687CB6729" pageId="0" pageNumber="1">Malagasy for brown in reference to body colouration.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CAA2500634CC2676461D39570AA4D322" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="16293AEA3C08BD48B898C185459EFD6B" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="FC09EC7C0A3EEF18A29346665993D231" pageId="0" pageNumber="1">The species was collected at 1300 m in elevation, in montane rainforest. Nests were located in rotten logs.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39DB8C90C661FBDC89677B2185FB1AA2" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="F98570D94AB722AE4FAB0668DAB53F5F" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="FF415C2AC44718F5A954AAB37D344009" pageId="0" pageNumber="1">
|
||
<taxonomicName id="7F500F6C2D5208509FE3140B604FEF1D" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole volontany" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="volontany">
|
||
<emphasis id="B149682172AB3F37A24F434ED917BDEE" italics="true" pageId="0" pageNumber="1">Pheidole volontany</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. belongs to the group of species characterised by small body size (major workers: HL <1.05 mm, WL <0.9 mm and minor workers HL <0.5 mm, WL <0.6 mm), sub-oval, slightly widening posteriorly head with anterior and posterior sides convex in major workers, and minor workers with yellow to brown body colouration and foveolate head or head predominantly smooth and relatively oval. The group includes six species:
|
||
<taxonomicName id="D97B140EABEF45F2218D695714C36B9F" lsidName="P. havoana" pageId="0" pageNumber="1" rank="species" species="havoana">
|
||
<emphasis id="7FE638481FBB4FDB9CB0B1BB3A0473A8" italics="true" pageId="0" pageNumber="1">P. havoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="3BA631501EBACECE8499A9FD961CBA62" lsidName="P. kely" pageId="0" pageNumber="1" rank="species" species="kely">
|
||
<emphasis id="6601C0E0755E473888746E7D204CD540" italics="true" pageId="0" pageNumber="1">P. kely</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="D9721719AFF70B49232628CAF25C7360" lsidName="P. parvula" pageId="0" pageNumber="1" rank="species" species="parvula">
|
||
<emphasis id="229AF59727D937A811C33A7C3369EF7F" italics="true" pageId="0" pageNumber="1">P. parvula</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="BA5B28B96BF9850B44DF02579A3573E2" lsidName="P. parvulogibba" pageId="0" pageNumber="1" rank="species" species="parvulogibba">
|
||
<emphasis id="22D5AAF41FD76757D98DC37AF190D25D" italics="true" pageId="0" pageNumber="1">P. parvulogibba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="36354C3F667A76ECA19D480349754301" lsidName="P. volontany" pageId="0" pageNumber="1" rank="species" species="volontany">
|
||
<emphasis id="38367C71CF7AB4ED0DF491C803E0564E" italics="true" pageId="0" pageNumber="1">P. volontany</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., and
|
||
<taxonomicName id="DAB36ED9F0E899D9DD052FF010828339" lsidName="P. midongy" pageId="0" pageNumber="1" rank="species" species="midongy">
|
||
<emphasis id="97E6C794786D9935DBA65AFF93984E89" italics="true" pageId="0" pageNumber="1">P. midongy</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Because of dark body colouration
|
||
<taxonomicName id="39E93E2861B9AEC27B33C9F9FEC0F0EC" lsidName="P. volontany" pageId="0" pageNumber="1" rank="species" species="volontany">
|
||
<emphasis id="A2F63C0CE2F745CB384339C12044A854" italics="true" pageId="0" pageNumber="1">P. volontany</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., known from
|
||
<normalizedToken id="053D18773202AF5435B85EC865F90617" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="961F9B3003169423FB97B140978AD7CC" originalValue="Classée">Classee</normalizedToken>
|
||
<normalizedToken id="1310509214A7F4FDEE85AA5B5516A616" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||
in Toliara, is most similar to
|
||
<taxonomicName id="21CB969EECBE4582CE53B9AFEE0F6DF1" lsidName="P. midongy" pageId="0" pageNumber="1" rank="species" species="midongy">
|
||
<emphasis id="F79F80C8060D60C485BD4A6B56F3F74E" italics="true" pageId="0" pageNumber="1">P. midongy</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., described form Parc National Befotaka-Midongy in Fianarantsoa. Major workers of
|
||
<taxonomicName id="BCAE26D50C7540CC5D487551EF339A1C" lsidName="P. volontany" pageId="0" pageNumber="1" rank="species" species="volontany">
|
||
<emphasis id="4B876E122062DD578EBD2CD36A6AEEB8" italics="true" pageId="0" pageNumber="1">P. volontany</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differ from
|
||
<taxonomicName id="1AD2C7EDFC57970F434C89C7D74D78C5" lsidName="P. midongy" pageId="0" pageNumber="1" rank="species" species="midongy">
|
||
<emphasis id="8EFD5CA9C27C718A5C01EB016C4F47BE" italics="true" pageId="0" pageNumber="1">P. midongy</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in frons with moderately dense, thick, longitudinal, and interrupted rugae with shiny and smooth interspaces, and sides of the head with dense, very long, suberect to erect pilosity; minor workers differ in predominantly smooth head and darker brown body colouration.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |