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<document id="AE18A7D6E1A9238CF4D69BEACE13600F" ID-DOI="10.11646/zootaxa.4748.2.3" ID-GBIF-Dataset="48d67ec7-59cb-4b6c-b2b0-a37142dc3aae" ID-ISSN="1175-5326" ID-Zenodo-Dep="3698777" ID-ZooBank="661CD94A-130B-4BD8-B201-28B079815618" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583482611617" checkinUser="plazi" docAuthor="Klautau, Michelle, Lopes, Matheus Vieira, Guarabyra, Bruna, Folcher, Eric, Ekins, Merrick &amp; Debitus, Cécile" docDate="2020" docId="6A74544AFFFA7245FF635898FF27FF45" docLanguage="en" docName="zootaxa.4748.2.3.pdf" docOrigin="Zootaxa 4748 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucandra tahuatae Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus 2020, sp. nov." docType="treatment" docUuid="A8A61580-4BAE-4672-8EF9-F6F120C9BF8A" docUuidSource="ZooBank" docVersion="7" lastPageNumber="286" masterDocId="964D2C32FFEC725FFFF45C5FFFF6FFBE" masterDocTitle="Calcareous sponges from the French Polynesia (Porifera: Calcarea)" masterLastPageNumber="295" masterPageNumber="261" pageNumber="283" updateTime="1698725212753" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B8F2D6FE4D2CCAFB23EA00E02920BA35">Calcareous sponges from the French Polynesia (Porifera: Calcarea)</mods:title>
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<mods:namePart id="D3EE9CC2422B0F134F0755C342CAA67F">Klautau, Michelle</mods:namePart>
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<mods:namePart id="5E59C20F7534100C57E876748AF093BC">Lopes, Matheus Vieira</mods:namePart>
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<mods:namePart id="E2297401310F588306CE9116CA68F927">Guarabyra, Bruna</mods:namePart>
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<mods:namePart id="05CCA0DEBF15F200B0759827B0CDE962">Folcher, Eric</mods:namePart>
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<mods:namePart id="8DD7697FB9E9BA29685C280058D148C4">Ekins, Merrick</mods:namePart>
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<mods:namePart id="7BADEBA9453338F22012555E019F94AA">Debitus, Cécile</mods:namePart>
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<mods:date id="7E7887783149306A574E1BD9AF911611">2020</mods:date>
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<mods:number id="132D0CC854610E250569231FD88B045F">4748</mods:number>
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<treatment id="6A74544AFFFA7245FF635898FF27FF45" ID-DOI="http://doi.org/10.5281/zenodo.3704832" ID-GBIF-Taxon="162462237" ID-Zenodo-Dep="3704832" LSID="urn:lsid:zoobank.org:act:A8A61580-4BAE-4672-8EF9-F6F120C9BF8A" httpUri="http://treatment.plazi.org/id/6A74544AFFFA7245FF635898FF27FF45" lastPageId="26" lastPageNumber="286" pageId="22" pageNumber="283">
<subSubSection id="AAC7B6D7FFFA7249FF635898FE07FB5C" box="[151,497,1223,1250]" pageId="22" pageNumber="283" type="nomenclature">
<paragraph id="E262E55CFFFA7249FF635898FE07FB5C" blockId="22.[151,497,1223,1286]" box="[151,497,1223,1250]" pageId="22" pageNumber="283">
<heading id="B92A5230FFFA7249FF635898FE07FB5C" bold="true" box="[151,497,1223,1250]" fontSize="11" level="1" pageId="22" pageNumber="283" reason="1">
<emphasis id="D0A9394EFFFA7249FF635898FE07FB5C" bold="true" box="[151,497,1223,1250]" pageId="22" pageNumber="283">
<taxonomicName id="25DD9EDFFFFA7249FF635898FE7BFB5C" ID-CoL="86X2N" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[151,397,1223,1250]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="22" pageNumber="283" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFFA7249FF635898FE7BFB5C" bold="true" box="[151,397,1223,1250]" italics="true" pageId="22" pageNumber="283">Leucandra tahuatae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CB9A8435FFFA7249FE605897FE07FB5C" box="[404,497,1224,1250]" pageId="22" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AAC7B6D7FFFA7249FF6358B3FE49FAB8" box="[151,447,1260,1286]" pageId="22" pageNumber="283" type="description">
<paragraph id="E262E55CFFFA7249FF6358B3FE49FAB8" blockId="22.[151,497,1223,1286]" box="[151,447,1260,1286]" pageId="22" pageNumber="283">
(
<figureCitation id="7AE6F9D9FFFA7249FF6B58B3FF00FAB8" box="[159,246,1260,1286]" captionStart="FIGURE 10" captionStartId="23.[151,250,1867,1892]" captionTargetBox="[159,1428,273,1842]" captionTargetId="figure@23.[159,1428,273,1842]" captionTargetPageId="23" captionText="FIGURE 10. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Specimen in situ. B—Fixed specimen. C—Oscular membrane. D—Cross-section of the choanosome. E—Chonanosome showing a canal (black arrow: tetractine; white arrow: triactine). The asterisk indicates larvae. F—Subatrial (arrows) and atrial skeletons. Abbreviation: at = atrium; cx = cortex." figureDoi="http://doi.org/10.5281/zenodo.3698797" httpUri="https://zenodo.org/record/3698797/files/figure.png" pageId="22" pageNumber="283">Figs 10</figureCitation>
,
<figureCitation id="7AE6F9D9FFFA7249FEF658B3FEE9FAB8" box="[258,287,1260,1286]" captionStart="FIGURE 11" captionStartId="24.[151,250,1256,1281]" captionTargetBox="[159,1428,181,1230]" captionTargetId="figure@24.[159,1428,181,1230]" captionTargetPageId="24" captionText="FIGURE 11. Leucandra tahuatae sp. nov. (UFRJPOR 6454). SEM. A, B—Tangential section of the cortex. C, D—Tangential section of the atrium." figureDoi="http://doi.org/10.5281/zenodo.3698799" httpUri="https://zenodo.org/record/3698799/files/figure.png" pageId="22" pageNumber="283">11</figureCitation>
,
<figureCitation id="7AE6F9D9FFFA7249FEDF58B3FEBFFAB8" box="[299,329,1260,1286]" captionStart="FIGURE 12" captionStartId="25.[151,250,1256,1281]" captionTargetBox="[159,1428,181,1230]" captionTargetId="figure@25.[159,1428,181,1230]" captionTargetPageId="25" captionText="FIGURE 12. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Cortical triactines. B—Choanosomal large triactines. C— Choanosomal small triactines. D—Choanosomal small tetractines. E—Subatrial triactines. F—Atrial tetractine." figureDoi="http://doi.org/10.5281/zenodo.3698801" httpUri="https://zenodo.org/record/3698801/files/figure.png" pageId="22" pageNumber="283">12</figureCitation>
,
<tableCitation id="AF5FD0E7FFFA7249FEA058B3FE40FAB8" box="[340,438,1260,1286]" captionStart="TABLE 10" captionStartId="26.[151,239,298,322]" captionTargetPageId="26" captionText="TABLE 10. Spicule measurements of the holotype of Leucandra tahuatae sp. nov. (UFRJPOR 6454)." pageId="22" pageNumber="283">Table 10</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="AAC7B6D7FFFA7249FF63596BFD78FAF1" box="[151,654,1332,1359]" pageId="22" pageNumber="283" type="etymology">
<paragraph id="E262E55CFFFA7249FF63596BFD78FAF1" blockId="22.[151,1437,1332,2043]" box="[151,654,1332,1359]" pageId="22" pageNumber="283">
<emphasis id="D0A9394EFFFA7249FF63596BFEE8FAF0" bold="true" box="[151,286,1332,1358]" pageId="22" pageNumber="283">Etymology.</emphasis>
From the
<typeStatus id="3D665BFEFFFA7249FE66596BFE34FAF0" box="[402,450,1332,1358]" pageId="22" pageNumber="283">type</typeStatus>
locality (Tahuata)
</paragraph>
</subSubSection>
<subSubSection id="AAC7B6D7FFFA7249FF335907FAECFA04" pageId="22" pageNumber="283" type="materials_examined">
<paragraph id="E262E55CFFFA7249FF335907FBF6FACC" blockId="22.[151,1437,1332,2043]" box="[199,1024,1367,1394]" pageId="22" pageNumber="283">
<materialsCitation id="52B5EF01FFFA7249FF335907FC0BFACC" ID-GBIF-Occurrence="2574624281" box="[199,1021,1367,1394]" country="French Polynesia" county="Marquesas Island" location="Matatehoke" municipality="Tahuata" pageId="22" pageNumber="283" specimenCount="1" typeStatus="holotype">
<emphasis id="D0A9394EFFFA7249FF335907FE90FACC" bold="true" box="[199,358,1367,1394]" pageId="22" pageNumber="283">
<typeStatus id="3D665BFEFFFA7249FF335907FEF4FACC" box="[199,258,1368,1394]" pageId="22" pageNumber="283">Type</typeStatus>
locality.
</emphasis>
<location id="E702B387FFFA7249FE985907FE05FACC" LSID="urn:lsid:plazi:treatment:6A74544AFFFA7245FF635898FF27FF45:E702B387FFFA7249FE985907FE05FACC" box="[364,499,1368,1394]" country="French Polynesia" county="Marquesas Island" municipality="Tahuata" name="Matatehoke" pageId="22" pageNumber="283">Matatehoke</location>
,
<collectingMunicipality id="02067F26FFFA7249FE0A5907FDAFFACC" box="[510,601,1368,1394]" pageId="22" pageNumber="283">Tahuata</collectingMunicipality>
,
<collectingCounty id="0B039DD0FFFA7249FD905907FCDAFACC" box="[612,812,1368,1394]" pageId="22" pageNumber="283">Marquesas Island</collectingCounty>
,
<collectingCountry id="9ACAA5CCFFFA7249FCC35907FC0BFACC" box="[823,1021,1368,1394]" name="French Polynesia" pageId="22" pageNumber="283">French Polynesia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="E262E55CFFFA7249FF335924FAECFA04" blockId="22.[151,1437,1332,2043]" pageId="22" pageNumber="283">
<emphasis id="D0A9394EFFFA7249FF335924FE45FA28" bold="true" box="[199,435,1403,1430]" pageId="22" pageNumber="283">Material examined.</emphasis>
<materialsCitation id="52B5EF01FFFA7249FE495923FAECFA04" ID-GBIF-Occurrence="2574624262" collectingDate="2009-09-10" collectingDateMax="2025" collectingDateMin="2009-09-10" collectionCode="MT" collectorName="C. Debitus" county="Station" elevation="33" latitude="-9.89315" location="Matatehoke" longLatPrecision="1" longitude="-149.55367" municipality="Marquesas Island" pageId="22" pageNumber="283" specimenCode="MT01" specimenCount="1" typeStatus="holotype">
<emphasis id="D0A9394EFFFA7249FE495923FCF7FA28" bold="true" box="[445,769,1403,1430]" pageId="22" pageNumber="283">
<typeStatus id="3D665BFEFFFA7249FE495923FDD8FA28" box="[445,558,1404,1430]" pageId="22" pageNumber="283" type="holotype">Holotype</typeStatus>
: UFRJPOR 6454
</emphasis>
=
<emphasis id="D0A9394EFFFA7249FCD15923FBD0FA28" bold="true" box="[805,1062,1404,1430]" pageId="22" pageNumber="283">
MNHN-IP-
<date id="9663C39CFFFA7249FC5A5923FBFEFA28" box="[942,1032,1404,1430]" pageId="22" pageNumber="283" value="2018" valueMax="2025">
<collectingDate id="86273A74FFFA7249FC5A5923FBFEFA28" box="[942,1032,1404,1430]" pageId="22" pageNumber="283" value="2018" valueMax="2025">2018-25</collectingDate>
</date>
</emphasis>
<location id="E702B387FFFA7249FBD15923FB5AFA28" LSID="urn:lsid:plazi:treatment:6A74544AFFFA7245FF635898FF27FF45:E702B387FFFA7249FBD15923FB5AFA28" box="[1061,1196,1404,1430]" county="Station" latitude="-9.89315" longLatPrecision="1" longitude="-149.55367" municipality="Marquesas Island" name="Matatehoke" pageId="22" pageNumber="283">Matatehoke</location>
,
<location id="E702B387FFFA7249FB4E5923FAE3FA28" LSID="urn:lsid:plazi:treatment:6A74544AFFFA7245FF635898FF27FF45:E702B387FFFA7249FB4E5923FAE3FA28" box="[1210,1301,1404,1430]" county="Station" latitude="-9.89315" longLatPrecision="1" longitude="-149.55367" municipality="Marquesas Island" name="Tahuata" pageId="22" pageNumber="283">Tahuata</location>
,
<collectingMunicipality id="02067F26FFFA7249FAD75923FF28FA04" pageId="22" pageNumber="283">Marquesas Island</collectingMunicipality>
,
<collectingCounty id="0B039DD0FFFA7249FF1E59FFFECFFA04" box="[234,313,1440,1466]" pageId="22" pageNumber="283">Station</collectingCounty>
<specimenCode id="B27B4D27FFFA7249FEB459FFFE70FA04" box="[320,390,1440,1466]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="22" pageNumber="283">MT01</specimenCode>
(
<geoCoordinate id="87E9839BFFFA7249FE6259FFFDC7FA04" box="[406,561,1439,1466]" degrees="9" direction="south" minutes="53.589" orientation="latitude" pageId="22" pageNumber="283" precision="1" value="-9.89315">9° 53.589 S</geoCoordinate>
<geoCoordinate id="87E9839BFFFA7249FDC559FFFD12FA04" box="[561,740,1440,1466]" degrees="149" direction="west" minutes="33.220" orientation="longitude" pageId="22" pageNumber="283" precision="1" value="-149.55367">149° 33.220 W</geoCoordinate>
), depth:
<quantity id="252548B9FFFA7249FCB159FFFC76FA04" box="[837,896,1440,1466]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="22" pageNumber="283" unit="m" value="33.0">
<elevation id="69F0026FFFFA7249FCB159FFFC76FA04" box="[837,896,1440,1466]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="22" pageNumber="283" unit="m" value="33.0">33 m</elevation>
</quantity>
, coll.
<collectorName id="4F28808AFFFA7249FC3059FFFBC8FA04" box="[964,1086,1440,1466]" pageId="22" pageNumber="283">C. Debitus</collectorName>
,
<date id="9663C39CFFFA7249FBBE59FFFB38FA04" box="[1098,1230,1440,1466]" pageId="22" pageNumber="283" value="2009-09-10">
<collectingDate id="86273A74FFFA7249FBBE59FFFB38FA04" box="[1098,1230,1440,1466]" pageId="22" pageNumber="283" value="2009-09-10">10/IX/2009</collectingDate>
</date>
, P177.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="AAC7B6D7FFFA7247FF33599CFC2CF930" lastPageId="24" lastPageNumber="285" pageId="22" pageNumber="283" type="diagnosis">
<paragraph id="E262E55CFFFA7249FF33599CFE3AF9BC" blockId="22.[151,1437,1332,2043]" pageId="22" pageNumber="283">
<emphasis id="D0A9394EFFFA7249FF33599CFEB4FA60" bold="true" box="[199,322,1475,1502]" pageId="22" pageNumber="283">Diagnosis.</emphasis>
Sponge white, tubular, with cortical triactines, large and small choanosomal triactines, and choanosomal and atrial tetractines.
</paragraph>
<paragraph id="E262E55CFFFA7249FF335A54FCF9F998" blockId="22.[151,1437,1332,2043]" box="[199,783,1547,1574]" pageId="22" pageNumber="283">
<emphasis id="D0A9394EFFFA7249FF335A54FED6F998" bold="true" box="[199,288,1547,1574]" pageId="22" pageNumber="283">Colour.</emphasis>
White alive and beige in ethanol (
<figureCitation id="7AE6F9D9FFFA7249FD555A53FCF4F998" box="[673,770,1548,1574]" captionStart="FIGURE 10" captionStartId="23.[151,250,1867,1892]" captionTargetBox="[159,1428,273,1842]" captionTargetId="figure@23.[159,1428,273,1842]" captionTargetPageId="23" captionText="FIGURE 10. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Specimen in situ. B—Fixed specimen. C—Oscular membrane. D—Cross-section of the choanosome. E—Chonanosome showing a canal (black arrow: tetractine; white arrow: triactine). The asterisk indicates larvae. F—Subatrial (arrows) and atrial skeletons. Abbreviation: at = atrium; cx = cortex." figureDoi="http://doi.org/10.5281/zenodo.3698797" httpUri="https://zenodo.org/record/3698797/files/figure.png" pageId="22" pageNumber="283">Fig 10A</figureCitation>
).
</paragraph>
<paragraph id="E262E55CFFFA7249FF335A6FFC2DF9D0" blockId="22.[151,1437,1332,2043]" pageId="22" pageNumber="283">
<emphasis id="D0A9394EFFFA7249FF335A6FFDF2F9F4" bold="true" box="[199,516,1584,1610]" pageId="22" pageNumber="283">Morphology and anatomy.</emphasis>
Sponge tubular with apical osculum (
<figureCitation id="7AE6F9D9FFFA7249FC5A5A6FFBCEF9F4" box="[942,1080,1584,1610]" captionStart="FIGURE 10" captionStartId="23.[151,250,1867,1892]" captionTargetBox="[159,1428,273,1842]" captionTargetId="figure@23.[159,1428,273,1842]" captionTargetPageId="23" captionText="FIGURE 10. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Specimen in situ. B—Fixed specimen. C—Oscular membrane. D—Cross-section of the choanosome. E—Chonanosome showing a canal (black arrow: tetractine; white arrow: triactine). The asterisk indicates larvae. F—Subatrial (arrows) and atrial skeletons. Abbreviation: at = atrium; cx = cortex." figureDoi="http://doi.org/10.5281/zenodo.3698797" httpUri="https://zenodo.org/record/3698797/files/figure.png" pageId="22" pageNumber="283">Figs 10A, B</figureCitation>
) surrounded by membrane (
<figureCitation id="7AE6F9D9FFFA7249FA815A6FFF3CF9D0" captionStart="FIGURE 10" captionStartId="23.[151,250,1867,1892]" captionTargetBox="[159,1428,273,1842]" captionTargetId="figure@23.[159,1428,273,1842]" captionTargetPageId="23" captionText="FIGURE 10. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Specimen in situ. B—Fixed specimen. C—Oscular membrane. D—Cross-section of the choanosome. E—Chonanosome showing a canal (black arrow: tetractine; white arrow: triactine). The asterisk indicates larvae. F—Subatrial (arrows) and atrial skeletons. Abbreviation: at = atrium; cx = cortex." figureDoi="http://doi.org/10.5281/zenodo.3698797" httpUri="https://zenodo.org/record/3698797/files/figure.png" pageId="22" pageNumber="283">Fig 10C</figureCitation>
). Surface smooth. Large central atrium. Aquiferous system leuconoid.
</paragraph>
<paragraph id="E262E55CFFFA7249FF335A27FCDCF89C" blockId="22.[151,1437,1332,2043]" pageId="22" pageNumber="283">
The cortical skeleton is composed of tangential triactines (
<figureCitation id="7AE6F9D9FFFA7249FC9F5A27FC0AF92C" box="[875,1020,1656,1682]" captionStart="FIGURE 11" captionStartId="24.[151,250,1256,1281]" captionTargetBox="[159,1428,181,1230]" captionTargetId="figure@24.[159,1428,181,1230]" captionTargetPageId="24" captionText="FIGURE 11. Leucandra tahuatae sp. nov. (UFRJPOR 6454). SEM. A, B—Tangential section of the cortex. C, D—Tangential section of the atrium." figureDoi="http://doi.org/10.5281/zenodo.3698799" httpUri="https://zenodo.org/record/3698799/files/figure.png" pageId="22" pageNumber="283">Figs 11A, B</figureCitation>
). In the choanosome there are large triactines (larger than those of the cortex) and small choanosomal triactines and tetractines. Some of these small triactines and the small tetractines surround the choanosomal canals (
<figureCitation id="7AE6F9D9FFFA7249FC545A9FFC09F964" box="[928,1023,1728,1754]" captionStart="FIGURE 10" captionStartId="23.[151,250,1867,1892]" captionTargetBox="[159,1428,273,1842]" captionTargetId="figure@23.[159,1428,273,1842]" captionTargetPageId="23" captionText="FIGURE 10. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Specimen in situ. B—Fixed specimen. C—Oscular membrane. D—Cross-section of the choanosome. E—Chonanosome showing a canal (black arrow: tetractine; white arrow: triactine). The asterisk indicates larvae. F—Subatrial (arrows) and atrial skeletons. Abbreviation: at = atrium; cx = cortex." figureDoi="http://doi.org/10.5281/zenodo.3698797" httpUri="https://zenodo.org/record/3698797/files/figure.png" pageId="22" pageNumber="283">Fig 10E</figureCitation>
). The subatrial skeleton is composed of triactines that point their unpaired actine to the cortex (
<figureCitation id="7AE6F9D9FFFA7249FCE15ABBFC87F940" box="[789,881,1764,1790]" captionStart="FIGURE 10" captionStartId="23.[151,250,1867,1892]" captionTargetBox="[159,1428,273,1842]" captionTargetId="figure@23.[159,1428,273,1842]" captionTargetPageId="23" captionText="FIGURE 10. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Specimen in situ. B—Fixed specimen. C—Oscular membrane. D—Cross-section of the choanosome. E—Chonanosome showing a canal (black arrow: tetractine; white arrow: triactine). The asterisk indicates larvae. F—Subatrial (arrows) and atrial skeletons. Abbreviation: at = atrium; cx = cortex." figureDoi="http://doi.org/10.5281/zenodo.3698797" httpUri="https://zenodo.org/record/3698797/files/figure.png" pageId="22" pageNumber="283">Fig 10F</figureCitation>
). The atrial skeleton has tangential tetractines that point their apical actine into the atrium (
<figureCitation id="7AE6F9D9FFFA7249FDAE5B57FD48F89C" box="[602,702,1800,1826]" captionStart="FIGURE 10" captionStartId="23.[151,250,1867,1892]" captionTargetBox="[159,1428,273,1842]" captionTargetId="figure@23.[159,1428,273,1842]" captionTargetPageId="23" captionText="FIGURE 10. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Specimen in situ. B—Fixed specimen. C—Oscular membrane. D—Cross-section of the choanosome. E—Chonanosome showing a canal (black arrow: tetractine; white arrow: triactine). The asterisk indicates larvae. F—Subatrial (arrows) and atrial skeletons. Abbreviation: at = atrium; cx = cortex." figureDoi="http://doi.org/10.5281/zenodo.3698797" httpUri="https://zenodo.org/record/3698797/files/figure.png" pageId="22" pageNumber="283">Figs 10F</figureCitation>
,
<figureCitation id="7AE6F9D9FFFA7249FD3D5B57FCEBF89C" box="[713,797,1800,1826]" captionStart="FIGURE 11" captionStartId="24.[151,250,1256,1281]" captionTargetBox="[159,1428,181,1230]" captionTargetId="figure@24.[159,1428,181,1230]" captionTargetPageId="24" captionText="FIGURE 11. Leucandra tahuatae sp. nov. (UFRJPOR 6454). SEM. A, B—Tangential section of the cortex. C, D—Tangential section of the atrium." figureDoi="http://doi.org/10.5281/zenodo.3698799" httpUri="https://zenodo.org/record/3698799/files/figure.png" pageId="22" pageNumber="283">11C, D</figureCitation>
).
</paragraph>
<paragraph id="E262E55CFFFA7249FF335B74FE52F8F8" blockId="22.[151,1437,1332,2043]" box="[199,420,1835,1862]" pageId="22" pageNumber="283">
<emphasis id="D0A9394EFFFA7249FF335B74FEDFF8F8" bold="true" box="[199,297,1835,1862]" pageId="22" pageNumber="283">Spicules</emphasis>
(
<tableCitation id="AF5FD0E7FFFA7249FECC5B73FE6DF8F8" box="[312,411,1836,1862]" captionStart="TABLE 10" captionStartId="26.[151,239,298,322]" captionTargetPageId="26" captionText="TABLE 10. Spicule measurements of the holotype of Leucandra tahuatae sp. nov. (UFRJPOR 6454)." pageId="22" pageNumber="283">Table 10</tableCitation>
)
</paragraph>
<paragraph id="E262E55CFFFA7249FF335B0FFA6EF831" blockId="22.[151,1437,1332,2043]" pageId="22" pageNumber="283">
<emphasis id="D0A9394EFFFA7249FF335B0FFE62F8D4" box="[199,404,1872,1898]" italics="true" pageId="22" pageNumber="283">Cortical triactines</emphasis>
. Subregular. Actines are slightly conical with sharp tips. Sometimes they are undulated. The unpaired actine is shorter than the paired ones (
<figureCitation id="7AE6F9D9FFFA7249FD575B2BFCF2F830" box="[675,772,1908,1934]" captionStart="FIGURE 12" captionStartId="25.[151,250,1256,1281]" captionTargetBox="[159,1428,181,1230]" captionTargetId="figure@25.[159,1428,181,1230]" captionTargetPageId="25" captionText="FIGURE 12. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Cortical triactines. B—Choanosomal large triactines. C— Choanosomal small triactines. D—Choanosomal small tetractines. E—Subatrial triactines. F—Atrial tetractine." figureDoi="http://doi.org/10.5281/zenodo.3698801" httpUri="https://zenodo.org/record/3698801/files/figure.png" pageId="22" pageNumber="283">Fig 12A</figureCitation>
). Size: 449.0/
<quantity id="252548B9FFFA7249FC565B2BFBF6F830" box="[930,1024,1908,1934]" metricMagnitude="-2" metricUnit="m" metricValue="3.26" pageId="22" pageNumber="283" unit="mm" value="32.6">32.6 µm</quantity>
(paired), 369.0/
<quantity id="252548B9FFFA7249FB4D5B2BFAE0F830" box="[1209,1302,1908,1934]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="22" pageNumber="283" unit="mm" value="28.3">28.3 µm</quantity>
(unpaired).
</paragraph>
<paragraph id="E262E55CFFFA7249FF335BC7FEB6F845" blockId="22.[151,1437,1332,2043]" pageId="22" pageNumber="283">
<emphasis id="D0A9394EFFFA7249FF335BC7FDEAF80C" box="[199,540,1944,1970]" italics="true" pageId="22" pageNumber="283">Choanosomal large triactines</emphasis>
. Subregular. Actines are slightly conical with sharp tips. Sometimes they are undulated. The unpaired actine is shorter than the paired ones (
<figureCitation id="7AE6F9D9FFFA7249FCA75BE3FC45F868" box="[851,947,1980,2006]" captionStart="FIGURE 12" captionStartId="25.[151,250,1256,1281]" captionTargetBox="[159,1428,181,1230]" captionTargetId="figure@25.[159,1428,181,1230]" captionTargetPageId="25" captionText="FIGURE 12. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Cortical triactines. B—Choanosomal large triactines. C— Choanosomal small triactines. D—Choanosomal small tetractines. E—Subatrial triactines. F—Atrial tetractine." figureDoi="http://doi.org/10.5281/zenodo.3698801" httpUri="https://zenodo.org/record/3698801/files/figure.png" pageId="22" pageNumber="283">Fig 12B</figureCitation>
). Size: 796.7/ 53.0 µm (paired), 537.8/
<quantity id="252548B9FFFA7249FA9F5BE4FF4BF844" metricMagnitude="-2" metricUnit="m" metricValue="5.029999999999999" pageId="22" pageNumber="283" unit="mm" value="50.3">50.3 µm</quantity>
(unpaired).
</paragraph>
<paragraph id="E262E55CFFFB7248FF335CC6FC6FFF42" blockId="23.[151,1436,153,252]" pageId="23" pageNumber="284">
<emphasis id="D0A9394EFFFB7248FF335CC6FDEEFF0D" box="[199,536,153,179]" italics="true" pageId="23" pageNumber="284">Choanosomal small triactines</emphasis>
. Strongly sagittal, with curved paired actines when they are surrounding the canals. Actines are slightly conical with sharp tips. One of the paired actines is frequently shorter than the other actines (
<figureCitation id="7AE6F9D9FFFB7248FF6B5CBEFF09FF45" box="[159,255,225,251]" captionStart="FIGURE 12" captionStartId="25.[151,250,1256,1281]" captionTargetBox="[159,1428,181,1230]" captionTargetId="figure@25.[159,1428,181,1230]" captionTargetPageId="25" captionText="FIGURE 12. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Cortical triactines. B—Choanosomal large triactines. C— Choanosomal small triactines. D—Choanosomal small tetractines. E—Subatrial triactines. F—Atrial tetractine." figureDoi="http://doi.org/10.5281/zenodo.3698801" httpUri="https://zenodo.org/record/3698801/files/figure.png" pageId="23" pageNumber="284">Fig 12C</figureCitation>
). Size: 151.4/ 11.0 µm (paired), 172.1/
<quantity id="252548B9FFFB7248FD4C5CBEFCE0FF45" box="[696,790,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="23" pageNumber="284" unit="mm" value="12.6">12.6 µm</quantity>
(unpaired).
</paragraph>
<caption id="B6A2B5D4FFFB7248FF635B14FACAF815" ID-DOI="http://doi.org/10.5281/zenodo.3698797" ID-Zenodo-Dep="3698797" httpUri="https://zenodo.org/record/3698797/files/figure.png" pageId="23" pageNumber="284" startId="23.[151,250,1867,1892]" targetBox="[159,1428,273,1842]" targetPageId="23">
<paragraph id="E262E55CFFFB7248FF635B14FACAF815" blockId="23.[151,1436,1867,1964]" pageId="23" pageNumber="284">
<emphasis id="D0A9394EFFFB7248FF635B14FED7F8DA" bold="true" box="[151,289,1867,1892]" pageId="23" pageNumber="284">FIGURE 10.</emphasis>
<taxonomicName id="25DD9EDFFFFB7248FEDD5B13FE0EF8DA" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[297,504,1868,1892]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="23" pageNumber="284" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFFB7248FEDD5B13FE0EF8DA" box="[297,504,1868,1892]" italics="true" pageId="23" pageNumber="284">Leucandra tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFFB7248FDF45B13FDA4F8DA" bold="true" box="[512,594,1868,1892]" pageId="23" pageNumber="284">
<taxonomicNameLabel id="CB9A8435FFFB7248FDF45B13FDA4F8DA" box="[512,594,1868,1892]" pageId="23" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UFRJPOR 6454). A—Specimen
<emphasis id="D0A9394EFFFB7248FC435B12FC0EF8DA" box="[951,1016,1869,1892]" italics="true" pageId="23" pageNumber="284">in situ</emphasis>
. B—Fixed specimen. C—Oscular membrane. D—Cross-section of the choanosome. E—Chonanosome showing a canal (black arrow: tetractine; white arrow: triactine). The asterisk indicates larvae. F—Subatrial (arrows) and atrial skeletons. Abbreviation: at = atrium; cx = cortex.
</paragraph>
</caption>
<caption id="B6A2B5D4FFF47247FF6358B7FE99FA9B" ID-DOI="http://doi.org/10.5281/zenodo.3698799" ID-Zenodo-Dep="3698799" httpUri="https://zenodo.org/record/3698799/files/figure.png" pageId="24" pageNumber="285" startId="24.[151,250,1256,1281]" targetBox="[159,1428,181,1230]" targetPageId="24">
<paragraph id="E262E55CFFF47247FF6358B7FE99FA9B" blockId="24.[151,1436,1256,1317]" pageId="24" pageNumber="285">
<emphasis id="D0A9394EFFF47247FF6358B7FEE9FABF" bold="true" box="[151,287,1256,1281]" pageId="24" pageNumber="285">FIGURE 11.</emphasis>
<taxonomicName id="25DD9EDFFFF47247FED158B6FE04FABF" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[293,498,1257,1281]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="24" pageNumber="285" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF47247FED158B6FE04FABF" box="[293,498,1257,1281]" italics="true" pageId="24" pageNumber="285">Leucandra tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF47247FE0C58B6FDBFFABF" bold="true" box="[504,585,1257,1281]" pageId="24" pageNumber="285">
<taxonomicNameLabel id="CB9A8435FFF47247FE0C58B6FDBFFABF" box="[504,585,1257,1281]" pageId="24" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UFRJPOR 6454). SEM. A, B—Tangential section of the cortex. C, D—Tangential section of the atrium.
</paragraph>
</caption>
<paragraph id="E262E55CFFF47247FF33590CFAE6FA64" blockId="24.[151,1437,1363,2037]" pageId="24" pageNumber="285">
<emphasis id="D0A9394EFFF47247FF33590CFDD0FAD3" box="[199,550,1363,1389]" italics="true" pageId="24" pageNumber="285">Choanosomal small tetractines</emphasis>
. Strongly sagittal with curved paired actines, as they are find surrounding the canals. Actines are slightly conical with sharp tips. The unpaired actine can present the same length of the paired ones or be shorter. The paired actines are undulated. The apical actine is much shorter and thinner. It is conical, sharp an smooth (
<figureCitation id="7AE6F9D9FFF47247FEEE59E0FE8AFA67" box="[282,380,1471,1497]" captionStart="FIGURE 12" captionStartId="25.[151,250,1256,1281]" captionTargetBox="[159,1428,181,1230]" captionTargetId="figure@25.[159,1428,181,1230]" captionTargetPageId="25" captionText="FIGURE 12. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Cortical triactines. B—Choanosomal large triactines. C— Choanosomal small triactines. D—Choanosomal small tetractines. E—Subatrial triactines. F—Atrial tetractine." figureDoi="http://doi.org/10.5281/zenodo.3698801" httpUri="https://zenodo.org/record/3698801/files/figure.png" pageId="24" pageNumber="285">Fig 12D</figureCitation>
). Size: 158.1/
<quantity id="252548B9FFF47247FDE859E0FD8FFA64" box="[540,633,1471,1498]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="24" pageNumber="285" unit="mm" value="11.7">11.7 µm</quantity>
(paired), 184.1/ 13.0 µm (unpaired), 50.9/
<quantity id="252548B9FFF47247FBA859E0FB5AFA64" box="[1116,1196,1471,1498]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="24" pageNumber="285" unit="mm" value="8.9">8.9 µm</quantity>
(apical).
</paragraph>
<paragraph id="E262E55CFFF47247FF3359BCFAECF99C" blockId="24.[151,1437,1363,2037]" pageId="24" pageNumber="285">
<emphasis id="D0A9394EFFF47247FF3359BCFE57FA43" box="[199,417,1507,1533]" italics="true" pageId="24" pageNumber="285">Subatrial triactines</emphasis>
. Strongly sagittal (T-shaped). Actines are slightly conical with sharp tips. The unpaired actine is longer than the paired ones (
<figureCitation id="7AE6F9D9FFF47247FDD45A58FD89F99F" box="[544,639,1543,1569]" captionStart="FIGURE 12" captionStartId="25.[151,250,1256,1281]" captionTargetBox="[159,1428,181,1230]" captionTargetId="figure@25.[159,1428,181,1230]" captionTargetPageId="25" captionText="FIGURE 12. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Cortical triactines. B—Choanosomal large triactines. C— Choanosomal small triactines. D—Choanosomal small tetractines. E—Subatrial triactines. F—Atrial tetractine." figureDoi="http://doi.org/10.5281/zenodo.3698801" httpUri="https://zenodo.org/record/3698801/files/figure.png" pageId="24" pageNumber="285">Fig 12E</figureCitation>
). Size: 163.6/
<quantity id="252548B9FFF47247FCEB5A58FC8BF99C" box="[799,893,1543,1570]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="24" pageNumber="285" unit="mm" value="13.2">13.2 µm</quantity>
(paired), 261.3/
<quantity id="252548B9FFF47247FBCC5A58FB60F99C" box="[1080,1174,1543,1570]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="24" pageNumber="285" unit="mm" value="16.3">16.3 µm</quantity>
(unpaired).
</paragraph>
<paragraph id="E262E55CFFF47247FF335A74FC2CF930" blockId="24.[151,1437,1363,2037]" pageId="24" pageNumber="285">
<emphasis id="D0A9394EFFF47247FF335A74FE70F9FB" box="[199,390,1579,1605]" italics="true" pageId="24" pageNumber="285">Atrial tetractines</emphasis>
. Sagittal. Actines are slightly conical with sharp tips. The basal actines show the same length and they are straight. The apical actine is shorter than the basal ones, conical, sharp and smooth (
<figureCitation id="7AE6F9D9FFF47247FB045A10FAB9F9D7" box="[1264,1359,1615,1641]" captionStart="FIGURE 12" captionStartId="25.[151,250,1256,1281]" captionTargetBox="[159,1428,181,1230]" captionTargetId="figure@25.[159,1428,181,1230]" captionTargetPageId="25" captionText="FIGURE 12. Leucandra tahuatae sp. nov. (UFRJPOR 6454). A—Cortical triactines. B—Choanosomal large triactines. C— Choanosomal small triactines. D—Choanosomal small tetractines. E—Subatrial triactines. F—Atrial tetractine." figureDoi="http://doi.org/10.5281/zenodo.3698801" httpUri="https://zenodo.org/record/3698801/files/figure.png" pageId="24" pageNumber="285">Fig 12F</figureCitation>
). Size: 214.8/
<quantity id="252548B9FFF47247FF115A2CFEB5F930" box="[229,323,1651,1678]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="24" pageNumber="285" unit="mm" value="14.9">14.9 µm</quantity>
(paired), 215.5/
<quantity id="252548B9FFF47247FE0A5A2CFDAAF930" box="[510,604,1651,1678]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="24" pageNumber="285" unit="mm" value="14.8">14.8 µm</quantity>
(unpaired), 50.1/
<quantity id="252548B9FFF47247FCD25A2CFC80F930" box="[806,886,1651,1678]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="24" pageNumber="285" unit="mm" value="8.1">8.1 µm</quantity>
(apical).
</paragraph>
</subSubSection>
<subSubSection id="AAC7B6D7FFF47247FF335AC8FBA5F968" pageId="24" pageNumber="285" type="biology_ecology">
<paragraph id="E262E55CFFF47247FF335AC8FCA1F90C" blockId="24.[151,1437,1363,2037]" box="[199,855,1687,1714]" pageId="24" pageNumber="285">
<emphasis id="D0A9394EFFF47247FF335AC8FEDAF90F" bold="true" box="[199,300,1687,1713]" pageId="24" pageNumber="285">Ecology.</emphasis>
There was algae inside the atrium of this sponge.
</paragraph>
<paragraph id="E262E55CFFF47247FF335AE5FBA5F968" blockId="24.[151,1437,1363,2037]" box="[199,1107,1722,1750]" pageId="24" pageNumber="285">
<emphasis id="D0A9394EFFF47247FF335AE5FDF3F96B" bold="true" box="[199,517,1722,1749]" pageId="24" pageNumber="285">Geographical distribution.</emphasis>
Marquesas Island,
<collectingCountry id="9ACAA5CCFFF47247FD2A5AE4FC56F96B" box="[734,928,1723,1749]" name="French Polynesia" pageId="24" pageNumber="285">French Polynesia</collectingCountry>
(present work).
</paragraph>
</subSubSection>
<subSubSection id="AAC7B6D7FFF47245FF335A80FF27FF45" lastPageId="26" lastPageNumber="287" pageId="24" pageNumber="285" type="discussion">
<paragraph id="E262E55CFFF47247FF335A80FF21F84B" blockId="24.[151,1437,1363,2037]" pageId="24" pageNumber="285">
<emphasis id="D0A9394EFFF47247FF335A80FECDF947" bold="true" box="[199,315,1759,1785]" pageId="24" pageNumber="285">Remarks.</emphasis>
Most leucandras present diactines in their skeleton but
<taxonomicName id="25DD9EDFFFF47247FC5A5ABFFBD8F947" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[942,1070,1759,1785]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="24" pageNumber="285" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF47247FC5A5ABFFBD8F947" box="[942,1070,1759,1785]" italics="true" pageId="24" pageNumber="285">L. tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF47247FBC25A80FB66F947" bold="true" box="[1078,1168,1759,1785]" pageId="24" pageNumber="285">
<taxonomicNameLabel id="CB9A8435FFF47247FBC25A80FB66F947" box="[1078,1168,1759,1785]" pageId="24" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is part of a small group of this genus without diactines. Another characteristic that differentiates the new species from most leucandras is the presence of a subatrial skeleton. However, sometimes it is very difficult to be sure if a species does not have subatrial skeleton or if its author just did not mention it. Hence, to compare our new species other species of
<taxonomicName id="25DD9EDFFFF47247FAE85B14FA63F8DB" authorityName="Haeckel" authorityYear="1872" box="[1308,1429,1867,1893]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="24" pageNumber="285" phylum="Porifera" rank="genus">
<emphasis id="D0A9394EFFF47247FAE85B14FA63F8DB" box="[1308,1429,1867,1893]" italics="true" pageId="24" pageNumber="285">Leucandra</emphasis>
</taxonomicName>
, we considered only those with skeleton composed of cortical triactines, choanosomal large and small triactines, choanosomal tetractines, and atrial tetractines. We found only three most similar species to
<taxonomicName id="25DD9EDFFFF47247FB5B5BCBFAC6F813" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[1199,1328,1939,1965]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="24" pageNumber="285" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF47247FB5B5BCBFAC6F813" box="[1199,1328,1939,1965]" italics="true" pageId="24" pageNumber="285">L. tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF47247FACD5BCCFA62F813" bold="true" box="[1337,1428,1939,1965]" pageId="24" pageNumber="285">
<taxonomicNameLabel id="CB9A8435FFF47247FACD5BCCFA62F813" box="[1337,1428,1939,1965]" pageId="24" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
:
<taxonomicName id="25DD9EDFFFF47247FF635BE7FE4FF86F" authority="(Burton, 1934)" baseAuthorityName="Burton" baseAuthorityYear="1934" box="[151,441,1975,2001]" class="Calcarea" family="Clathrinidae" genus="Clathrina" kingdom="Animalia" order="Clathrinida" pageId="24" pageNumber="285" phylum="Porifera" rank="species" species="ramosa">
<emphasis id="D0A9394EFFF47247FF635BE7FEFFF86F" box="[151,265,1976,2001]" italics="true" pageId="24" pageNumber="285">L. ramosa</emphasis>
(
<bibRefCitation id="864C98ADFFF47247FEEC5BE8FE44F86F" author="Burton, M." box="[280,434,1975,2001]" pageId="24" pageNumber="285" pagination="513 - 621" refId="ref17171" refString="Burton, M. (1934) Sponges. Great Barrier reef Expedition, 1928 - 29, 4 (14), 513 - 621." type="journal article" year="1934">Burton, 1934</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="25DD9EDFFFF47247FE315BE7FC06F86F" authority="Van Soest &amp; De Voogd, 2018" authorityName="Van Soest &amp; De Voogd" authorityYear="2018" box="[453,1008,1975,2002]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="24" pageNumber="285" phylum="Porifera" rank="species" species="mozambiquensis">
<emphasis id="D0A9394EFFF47247FE315BE7FD6AF86F" box="[453,668,1975,2001]" italics="true" pageId="24" pageNumber="285">L. mozambiquensis</emphasis>
<bibRefCitation id="864C98ADFFF47247FD575BE7FC06F86F" author="Van Soest, R. W. N. &amp; De Voogd, N. J." box="[675,1008,1975,2002]" pageId="24" pageNumber="285" pagination="1 - 160" refId="ref20893" refString="Van Soest, R. W. N. &amp; De Voogd, N. J. (2018) Calcareous sponges of the Western Indian Ocean and Red Sea. Zootaxa, 4426 (1), 1 - 160. https: // doi. org / 10.11646 / zootaxa. 4426.1.1" type="journal article" year="2018">Van Soest &amp; De Voogd, 2018</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="25DD9EDFFFF47247FBDF5BE7FF25F84B" authority="Van Soest &amp; De Voogd, 2018" authorityName="Van Soest &amp; De Voogd" authorityYear="2018" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="24" pageNumber="285" phylum="Porifera" rank="species" species="pilula">
<emphasis id="D0A9394EFFF47247FBDF5BE7FB7CF86F" box="[1067,1162,1975,2001]" italics="true" pageId="24" pageNumber="285">L. pilula</emphasis>
<bibRefCitation id="864C98ADFFF47247FB655BE7FF25F84B" author="Van Soest, R. W. N. &amp; De Voogd, N. J." pageId="24" pageNumber="285" pagination="1 - 160" refId="ref20893" refString="Van Soest, R. W. N. &amp; De Voogd, N. J. (2018) Calcareous sponges of the Western Indian Ocean and Red Sea. Zootaxa, 4426 (1), 1 - 160. https: // doi. org / 10.11646 / zootaxa. 4426.1.1" type="journal article" year="2018">Van Soest &amp; De Voogd, 2018</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="B6A2B5D4FFF57246FF6358B7FAF4FA9B" ID-DOI="http://doi.org/10.5281/zenodo.3698801" ID-Zenodo-Dep="3698801" httpUri="https://zenodo.org/record/3698801/files/figure.png" pageId="25" pageNumber="286" startId="25.[151,250,1256,1281]" targetBox="[159,1428,181,1230]" targetPageId="25">
<paragraph id="E262E55CFFF57246FF6358B7FAF4FA9B" blockId="25.[151,1436,1256,1317]" pageId="25" pageNumber="286">
<emphasis id="D0A9394EFFF57246FF6358B7FED7FABF" bold="true" box="[151,289,1256,1281]" pageId="25" pageNumber="286">FIGURE 12.</emphasis>
<taxonomicName id="25DD9EDFFFF57246FEDD58B6FE0EFABF" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[297,504,1257,1281]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF57246FEDD58B6FE0EFABF" box="[297,504,1257,1281]" italics="true" pageId="25" pageNumber="286">Leucandra tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF57246FDF458B6FDA4FABF" bold="true" box="[512,594,1257,1281]" pageId="25" pageNumber="286">
<taxonomicNameLabel id="CB9A8435FFF57246FDF458B6FDA4FABF" box="[512,594,1257,1281]" pageId="25" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UFRJPOR 6454). A—Cortical triactines. B—Choanosomal large triactines. C— Choanosomal small triactines. D—Choanosomal small tetractines. E—Subatrial triactines. F—Atrial tetractine.
</paragraph>
</caption>
<paragraph id="E262E55CFFF57246FF33590CFB74F9F8" blockId="25.[151,1437,1363,2038]" pageId="25" pageNumber="286">
<taxonomicName id="25DD9EDFFFF57246FF33590CFE6AFAD3" baseAuthorityName="Burton" baseAuthorityYear="1934" box="[199,412,1363,1389]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="ramosa">
<emphasis id="D0A9394EFFF57246FF33590CFE6AFAD3" box="[199,412,1363,1389]" italics="true" pageId="25" pageNumber="286">Leucandra ramosa</emphasis>
</taxonomicName>
was originally described from
<collectingCountry id="9ACAA5CCFFF57246FD09590CFC91FAD3" box="[765,871,1363,1389]" name="Australia" pageId="25" pageNumber="286">Australia</collectingCountry>
. It can be differentiated from
<taxonomicName id="25DD9EDFFFF57246FB4D590BFACCFAD3" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[1209,1338,1363,1389]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF57246FB4D590BFACCFAD3" box="[1209,1338,1363,1389]" italics="true" pageId="25" pageNumber="286">L. tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF57246FAB6590CFA6AFAD3" bold="true" box="[1346,1436,1363,1389]" pageId="25" pageNumber="286">
<taxonomicNameLabel id="CB9A8435FFF57246FAB6590CFA6AFAD3" box="[1346,1436,1363,1389]" pageId="25" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by its greyish-brown colour in ethanol (the new species is white) and by the size of some spicule categories. Cortical triactines: up to 210.0/ 11.0 (
<taxonomicName id="25DD9EDFFFF57246FE0B59C3FD87FA0B" baseAuthorityName="Burton" baseAuthorityYear="1934" box="[511,625,1436,1461]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="ramosa">
<emphasis id="D0A9394EFFF57246FE0B59C3FD87FA0B" box="[511,625,1436,1461]" italics="true" pageId="25" pageNumber="286">L. ramosa</emphasis>
</taxonomicName>
); 260.0˗
<emphasis id="D0A9394EFFF57246FD2559C4FCE6FA0B" box="[721,784,1435,1461]" italics="true" pageId="25" pageNumber="286">449.0</emphasis>
˗700.0/ 25.0˗
<emphasis id="D0A9394EFFF57246FC5559C4FC24FA0B" box="[929,978,1435,1461]" italics="true" pageId="25" pageNumber="286">32.6</emphasis>
˗40.0—paired actine, 180.0˗
<emphasis id="D0A9394EFFF57246FAF959C4FABAFA0B" box="[1293,1356,1435,1461]" italics="true" pageId="25" pageNumber="286">369.0</emphasis>
˗550.0/ 10.0˗
<emphasis id="D0A9394EFFF57246FF2659E0FEF5FA67" box="[210,259,1471,1497]" italics="true" pageId="25" pageNumber="286">28.3</emphasis>
˗35.0—unpaired actine (
<taxonomicName id="25DD9EDFFFF57246FDE6599FFD66FA67" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[530,656,1471,1497]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF57246FDE6599FFD66FA67" box="[530,656,1471,1497]" italics="true" pageId="25" pageNumber="286">L. tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF57246FD6159E0FD1BFA67" bold="true" box="[661,749,1471,1497]" pageId="25" pageNumber="286">
<taxonomicNameLabel id="CB9A8435FFF57246FD6159E0FD1BFA67" box="[661,749,1471,1497]" pageId="25" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Choanosomal large triactines: up to 960.0/ 64.0 (
<taxonomicName id="25DD9EDFFFF57246FAEF599FFA7DFA67" baseAuthorityName="Burton" baseAuthorityYear="1934" box="[1307,1419,1472,1497]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="ramosa">
<emphasis id="D0A9394EFFF57246FAEF599FFA7DFA67" box="[1307,1419,1472,1497]" italics="true" pageId="25" pageNumber="286">L. ramosa</emphasis>
</taxonomicName>
); 508.1˗
<emphasis id="D0A9394EFFF57246FF1459BCFEE9FA43" box="[224,287,1507,1533]" italics="true" pageId="25" pageNumber="286">796.7</emphasis>
˗1016.1/ 37.8˗
<emphasis id="D0A9394EFFF57246FE4959BCFE18FA43" box="[445,494,1507,1533]" italics="true" pageId="25" pageNumber="286">53.0</emphasis>
˗64.9—paired actine, 378.4˗
<emphasis id="D0A9394EFFF57246FCD359BCFC90FA43" box="[807,870,1507,1533]" italics="true" pageId="25" pageNumber="286">537.8</emphasis>
˗756.7/ 37.8˗
<emphasis id="D0A9394EFFF57246FC0259BCFBD1FA43" box="[1014,1063,1507,1533]" italics="true" pageId="25" pageNumber="286">50.3</emphasis>
˗54.1—unpaired actine (
<taxonomicName id="25DD9EDFFFF57246FAC359BBFF4FF99F" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF57246FAC359BBFF4FF99F" italics="true" pageId="25" pageNumber="286">L. tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF57246FF375A58FED6F99F" bold="true" box="[195,288,1543,1569]" pageId="25" pageNumber="286">
<taxonomicNameLabel id="CB9A8435FFF57246FF375A58FED6F99F" box="[195,288,1543,1569]" pageId="25" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Choanosomal small triactines: up to 240.0/ 12.0 (
<taxonomicName id="25DD9EDFFFF57246FC8D5A57FC18F99F" baseAuthorityName="Burton" baseAuthorityYear="1934" box="[889,1006,1544,1569]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="ramosa">
<emphasis id="D0A9394EFFF57246FC8D5A57FC18F99F" box="[889,1006,1544,1569]" italics="true" pageId="25" pageNumber="286">L. ramosa</emphasis>
</taxonomicName>
); 108.0˗
<emphasis id="D0A9394EFFF57246FBA55A58FB66F99F" box="[1105,1168,1543,1569]" italics="true" pageId="25" pageNumber="286">151.4</emphasis>
˗202.5/ 9.5˗
<emphasis id="D0A9394EFFF57246FAE35A58FAB0F99F" box="[1303,1350,1543,1569]" italics="true" pageId="25" pageNumber="286">11.0</emphasis>
˗13.5— paired actine, 129.6˗
<emphasis id="D0A9394EFFF57246FE895A74FE4AF9FB" box="[381,444,1579,1605]" italics="true" pageId="25" pageNumber="286">172.1</emphasis>
˗221.4/ 10.8˗
<emphasis id="D0A9394EFFF57246FDBA5A74FD89F9FB" box="[590,639,1579,1605]" italics="true" pageId="25" pageNumber="286">12.6</emphasis>
˗13.5—unpaired actine (
<taxonomicName id="25DD9EDFFFF57246FC655A73FBE7F9FB" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[913,1041,1579,1605]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF57246FC655A73FBE7F9FB" box="[913,1041,1579,1605]" italics="true" pageId="25" pageNumber="286">L. tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF57246FBEC5A74FB87F9FB" bold="true" box="[1048,1137,1579,1605]" pageId="25" pageNumber="286">
<taxonomicNameLabel id="CB9A8435FFF57246FBEC5A74FB87F9FB" box="[1048,1137,1579,1605]" pageId="25" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<paragraph id="E262E55CFFF57246FF335A10FF39F968" blockId="25.[151,1437,1363,2038]" pageId="25" pageNumber="286">
<taxonomicName id="25DD9EDFFFF57246FF335A10FDF6F9D7" authorityName="Van Soest &amp; De Voogd" authorityYear="2018" box="[199,512,1615,1641]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="mozambiquensis">
<emphasis id="D0A9394EFFF57246FF335A10FDF6F9D7" box="[199,512,1615,1641]" italics="true" pageId="25" pageNumber="286">Leucandra mozambiquensis</emphasis>
</taxonomicName>
, recently described from
<collectingCountry id="9ACAA5CCFFF57246FCEF5A10FC59F9D7" box="[795,943,1615,1641]" name="Mozambique" pageId="25" pageNumber="286">Mozambique</collectingCountry>
Channel, can be differentiated from the new species by its external morphology, as it is an “irregular cup-shaped hollow mass” (
<bibRefCitation id="864C98ADFFF57246FBBE5A2CFA63F933" author="Van Soest, R. W. N. &amp; De Voogd, N. J." box="[1098,1429,1651,1678]" pageId="25" pageNumber="286" pagination="1 - 160" refId="ref20893" refString="Van Soest, R. W. N. &amp; De Voogd, N. J. (2018) Calcareous sponges of the Western Indian Ocean and Red Sea. Zootaxa, 4426 (1), 1 - 160. https: // doi. org / 10.11646 / zootaxa. 4426.1.1" type="journal article" year="2018">Van Soest &amp; De Voogd 2018</bibRefCitation>
) while the new species is tubular, and mainly by the presence of tetractines instead of triactines in the subatrial skeleton.
</paragraph>
<paragraph id="E262E55CFFF57245FF335A80FF27FF45" blockId="25.[151,1437,1363,2038]" lastBlockId="26.[151,1436,152,251]" lastPageId="26" lastPageNumber="287" pageId="25" pageNumber="286">
<taxonomicName id="25DD9EDFFFF57246FF335A80FE78F947" authorityName="Van Soest &amp; De Voogd" authorityYear="2018" box="[199,398,1759,1785]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="pilula">
<emphasis id="D0A9394EFFF57246FF335A80FE78F947" box="[199,398,1759,1785]" italics="true" pageId="25" pageNumber="286">Leucandra pilula</emphasis>
</taxonomicName>
, described from
<collectingCountry id="9ACAA5CCFFF57246FDA95A80FD2EF947" box="[605,728,1759,1785]" name="Seychelles" pageId="25" pageNumber="286">Seychelles</collectingCountry>
, is globose, while
<taxonomicName id="25DD9EDFFFF57246FC495ABFFBB4F947" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[957,1090,1759,1785]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF57246FC495ABFFBB4F947" box="[957,1090,1759,1785]" italics="true" pageId="25" pageNumber="286">L. tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF57246FBBB5A80FB58F947" bold="true" box="[1103,1198,1759,1785]" pageId="25" pageNumber="286">
<taxonomicNameLabel id="CB9A8435FFF57246FBBB5A80FB58F947" box="[1103,1198,1759,1785]" pageId="25" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is tubular. The former has oxhorn-shaped cortical triactines, while ours has equiangular cortical triactines. Besides, some of their spicule categories have sizes. Cortical triactines: 216.0˗
<emphasis id="D0A9394EFFF57246FC615B78FC22F8FF" box="[917,980,1831,1857]" italics="true" pageId="25" pageNumber="286">281.0</emphasis>
˗372.0/ 16.0˗
<emphasis id="D0A9394EFFF57246FB865B78FB55F8FF" box="[1138,1187,1831,1857]" italics="true" pageId="25" pageNumber="286">21.6</emphasis>
˗28.0—paired actine, 178.0˗
<emphasis id="D0A9394EFFF57246FF145B14FEE9F8DB" box="[224,287,1867,1893]" italics="true" pageId="25" pageNumber="286">245.0</emphasis>
˗326.0/ 15.0˗
<emphasis id="D0A9394EFFF57246FE5B5B14FE16F8DB" box="[431,480,1867,1893]" italics="true" pageId="25" pageNumber="286">22.7</emphasis>
˗31.0—unpaired actine (
<taxonomicName id="25DD9EDFFFF57246FD055B13FCB9F8DB" authorityName="Van Soest &amp; De Voogd" authorityYear="2018" box="[753,847,1867,1893]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="pilula">
<emphasis id="D0A9394EFFF57246FD055B13FCB9F8DB" box="[753,847,1867,1893]" italics="true" pageId="25" pageNumber="286">L. pilula</emphasis>
</taxonomicName>
); 260.0˗
<emphasis id="D0A9394EFFF57246FC5B5B14FC18F8DB" box="[943,1006,1867,1893]" italics="true" pageId="25" pageNumber="286">449.0</emphasis>
˗700.0/ 25.0˗
<emphasis id="D0A9394EFFF57246FB8B5B14FB46F8DB" box="[1151,1200,1867,1893]" italics="true" pageId="25" pageNumber="286">32.6</emphasis>
˗40.0—paired actine, 180.0˗
<emphasis id="D0A9394EFFF57246FF145B30FEE9F837" box="[224,287,1903,1929]" italics="true" pageId="25" pageNumber="286">369.0</emphasis>
˗550.0/ 10.0˗
<emphasis id="D0A9394EFFF57246FE5B5B30FE16F837" box="[431,480,1903,1929]" italics="true" pageId="25" pageNumber="286">28.3</emphasis>
˗35.0—unpaired actine (
<taxonomicName id="25DD9EDFFFF57246FD055B2FFC86F837" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[753,880,1903,1929]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF57246FD055B2FFC86F837" box="[753,880,1903,1929]" italics="true" pageId="25" pageNumber="286">L. tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF57246FC825B30FC39F837" bold="true" box="[886,975,1903,1929]" pageId="25" pageNumber="286">
<taxonomicNameLabel id="CB9A8435FFF57246FC825B30FC39F837" box="[886,975,1903,1929]" pageId="25" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Subatrial triactines: 100.0˗
<emphasis id="D0A9394EFFF57246FAF95B30FABAF837" box="[1293,1356,1903,1929]" italics="true" pageId="25" pageNumber="286">186.0</emphasis>
˗303.0/ 11.0˗
<emphasis id="D0A9394EFFF57246FF245BCCFEF7F813" box="[208,257,1939,1965]" italics="true" pageId="25" pageNumber="286">16.6</emphasis>
˗29.0—paired actine, 94.0˗
<emphasis id="D0A9394EFFF57246FDC35BCCFD80F813" box="[567,630,1939,1965]" italics="true" pageId="25" pageNumber="286">172.0</emphasis>
˗254.0/ 9.0˗
<emphasis id="D0A9394EFFF57246FD095BCCFCD8F813" box="[765,814,1939,1965]" italics="true" pageId="25" pageNumber="286">17.8</emphasis>
˗26.0—unpaired actine (
<taxonomicName id="25DD9EDFFFF57246FBBC5BCBFB5CF813" authorityName="Van Soest &amp; De Voogd" authorityYear="2018" box="[1096,1194,1939,1965]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="pilula">
<emphasis id="D0A9394EFFF57246FBBC5BCBFB5CF813" box="[1096,1194,1939,1965]" italics="true" pageId="25" pageNumber="286">L. pilula</emphasis>
</taxonomicName>
); 116.1˗
<emphasis id="D0A9394EFFF57246FAF95BCCFABAF813" box="[1293,1356,1939,1965]" italics="true" pageId="25" pageNumber="286">163.8</emphasis>
˗243.0/ 10.2˗
<emphasis id="D0A9394EFFF57246FF265BE8FEF5F86F" box="[210,259,1975,2001]" italics="true" pageId="25" pageNumber="286">13.2</emphasis>
˗16.2—paired actine, 164.7˗
<emphasis id="D0A9394EFFF57246FDB75BE8FD74F86F" box="[579,642,1975,2001]" italics="true" pageId="25" pageNumber="286">261.3</emphasis>
˗335.1/ 10.8˗
<emphasis id="D0A9394EFFF57246FCE15BE8FCB0F86F" box="[789,838,1975,2001]" italics="true" pageId="25" pageNumber="286">16.3</emphasis>
˗21.6—unpaired actine (
<taxonomicName id="25DD9EDFFFF57246FBA95BE7FB29F86F" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[1117,1247,1975,2001]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="25" pageNumber="286" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF57246FBA95BE7FB29F86F" box="[1117,1247,1975,2001]" italics="true" pageId="25" pageNumber="286">L. tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF57246FB1C5BE8FAB5F86F" bold="true" box="[1256,1347,1975,2001]" pageId="25" pageNumber="286">
<taxonomicNameLabel id="CB9A8435FFF57246FB1C5BE8FAB5F86F" box="[1256,1347,1975,2001]" pageId="25" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Atrial tetractines: 136.0˗
<emphasis id="D0A9394EFFF57246FE9C5B84FE51F84B" box="[360,423,2011,2037]" italics="true" pageId="25" pageNumber="286">239.0</emphasis>
˗380.0/ 9.0˗
<emphasis id="D0A9394EFFF57246FDC55B84FD94F84B" box="[561,610,2011,2037]" italics="true" pageId="25" pageNumber="286">17.1</emphasis>
˗32.0—paired actine, 101.0˗
<emphasis id="D0A9394EFFF57246FC585B84FC1DF84B" box="[940,1003,2011,2037]" italics="true" pageId="25" pageNumber="286">186.0</emphasis>
˗271.0/ 14.0˗
<emphasis id="D0A9394EFFF57246FB775B84FB42F84B" box="[1155,1204,2011,2037]" italics="true" pageId="25" pageNumber="286">19.4</emphasis>
˗32.0—unpaired ac- tine, 45.0˗
<emphasis id="D0A9394EFFF67245FEF95CC6FEC8FF0D" box="[269,318,153,179]" italics="true" pageId="26" pageNumber="287">84.0</emphasis>
˗130.0/ 4.0˗
<emphasis id="D0A9394EFFF67245FE315CC6FE1EFF0D" box="[453,488,153,179]" italics="true" pageId="26" pageNumber="287">8.4</emphasis>
˗11.0—apical actine (
<taxonomicName id="25DD9EDFFFF67245FD155CC5FCB5FF0D" authorityName="Van Soest &amp; De Voogd" authorityYear="2018" box="[737,835,153,179]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="26" pageNumber="287" phylum="Porifera" rank="species" species="pilula">
<emphasis id="D0A9394EFFF67245FD155CC5FCB5FF0D" box="[737,835,153,179]" italics="true" pageId="26" pageNumber="287">L. pilula</emphasis>
</taxonomicName>
); 155.0˗
<emphasis id="D0A9394EFFF67245FC535CC6FC10FF0D" box="[935,998,153,179]" italics="true" pageId="26" pageNumber="287">214.8</emphasis>
˗325.0/ 13.5˗
<emphasis id="D0A9394EFFF67245FB8F5CC6FB5AFF0D" box="[1147,1196,153,179]" italics="true" pageId="26" pageNumber="287">14.9</emphasis>
˗15.0—paired actine, 150.0˗
<emphasis id="D0A9394EFFF67245FF145CE2FEE9FF69" box="[224,287,189,215]" italics="true" pageId="26" pageNumber="287">215.5</emphasis>
˗300.0/ 12.5˗
<emphasis id="D0A9394EFFF67245FE455CE2FE14FF69" box="[433,482,189,215]" italics="true" pageId="26" pageNumber="287">14.8</emphasis>
˗15.0—unpaired actine, 30.0˗
<emphasis id="D0A9394EFFF67245FCC45CE2FC97FF69" box="[816,865,189,215]" italics="true" pageId="26" pageNumber="287">50.1</emphasis>
˗67.5/ 6.3˗
<emphasis id="D0A9394EFFF67245FC2C5CE2FC0DFF69" box="[984,1019,189,215]" italics="true" pageId="26" pageNumber="287">8.1</emphasis>
˗10.0—apical actine (
<taxonomicName id="25DD9EDFFFF67245FB055CE1FA84FF69" authority="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus, 2020" authorityName="Klautau &amp; Lopes &amp; Guarabyra &amp; Folcher &amp; Ekins &amp; Debitus" authorityYear="2020" box="[1265,1394,189,215]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="26" pageNumber="287" phylum="Porifera" rank="species" species="tahuatae" status="sp. nov.">
<emphasis id="D0A9394EFFF67245FB055CE1FA84FF69" box="[1265,1394,189,215]" italics="true" pageId="26" pageNumber="287">L. tahuatae</emphasis>
</taxonomicName>
<emphasis id="D0A9394EFFF67245FA8F5CE2FF3EFF45" bold="true" pageId="26" pageNumber="287">
<taxonomicNameLabel id="CB9A8435FFF67245FA8F5CE2FF3EFF45" pageId="26" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
)
</paragraph>
</subSubSection>
</treatment>
</document>