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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1086.77180" ID-GBIF-Dataset="b8abb16d-caa1-4f58-bf4d-61c14200dddd" ID-PMC="PMC8863764" ID-Pensoft-Pub="1313-2970-1086-1" ID-Pensoft-UUID="BD013F15E39C5B5FBE666837836810E1" ID-PubMed="35210911" ID-ZooBank="0AFF7CFB9400477F81160120C393C4FD" ModsDocID="1313-2970-1086-1" checkinTime="1644966841060" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Ablett, Jonathan D., Szabo, Marton &amp; Neubert, Eike" docDate="2022" docId="684CF16C1B55500CAE16A19ADD467913" docLanguage="en" docName="ZooKeys 1086: 1-31" docOrigin="ZooKeys 1086" docPubDate="2022-02-15" docSource="http://dx.doi.org/10.3897/zookeys.1086.77180" docTitle="Helix (Trachia) delibrata subsp. var. var. khasiensis Nevill 1877" docType="treatment" docVersion="4" id="BD013F15E39C5B5FBE666837836810E1" lastPageNumber="1" masterDocId="BD013F15E39C5B5FBE666837836810E1" masterDocTitle="Revision of the &quot; Chloritis delibrata (Benson, 1836) &quot; group (Gastropoda, Stylommatophora, Camaenidae)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="1" updateTime="1668151475586" updateUser="ExternalLinkService">
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<mods:title>Revision of the &quot; Chloritis delibrata (Benson, 1836) &quot; group (Gastropoda, Stylommatophora, Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto Street 15, Budapest, H- 1022, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ablett, Jonathan D.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7277-1934</mods:nameIdentifier>
<mods:affiliation>Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Szabo, Marton</mods:namePart>
<mods:affiliation>Hungarian Natural History Museum, Department of Paleontology and Geology, Ludovika ter 2, Budapest 1083, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Neubert, Eike</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0277-2894</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of the Burgergemeinde Bern, Bernastr. 15, CH- 3005 Berne, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, 3012 Bern, Switzerland</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-02-15</mods:number>
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<mods:detail type="volume">
<mods:number>1086</mods:number>
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<mods:start>1</mods:start>
<mods:end>31</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1086.77180</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1086.77180</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1086-1</mods:identifier>
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<treatment ID-GBIF-Taxon="193377860" LSID="urn:lsid:plazi:treatment:684CF16C1B55500CAE16A19ADD467913" httpUri="http://treatment.plazi.org/id/684CF16C1B55500CAE16A19ADD467913" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="684CF16C-1B55-500C-AE16-A19ADD467913" authority="Nevill, 1877, taxon inquirendum" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Trachia) delibrata subsp. var. var. khasiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="delibrata" subGenus="Trachia" subSpecies="var." variety="khasiensis">Helix (Trachia) delibrata var. khasiensis Nevill, 1877, taxon inquirendum</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" authorityYear="2022" baseAuthorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix delibrata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="delibrata">Helix delibrata</taxonomicName>
(?) var. Hanley &amp; Theobald, 1870: pl. 14, fig. 9.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Trachia) delibrata subsp. Var. var. khasiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="delibrata" subGenus="Trachia" subSpecies="Var." variety="khasiensis">Helix (Trachia) delibrata Var. khasiensis</taxonomicName>
Nevill, 1877: 21.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="G. Nevill" baseAuthorityYear="1877" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis (Trichochloritis) delibrata subsp. var. var. khasiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="delibrata" subGenus="Trichochloritis" subSpecies="var." variety="khasiensis">Chloritis (Trichochloritis) delibrata var. khasiensis</taxonomicName>
Gude, 1914: 173.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="G. Nevill" baseAuthorityYear="1877" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis delibrata subsp. var. var. khasiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="delibrata" subSpecies="var." variety="khasiensis">Chloritis delibrata var. khasiensis</taxonomicName>
Richardson, 1985: 93.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="G. Nevill" baseAuthorityYear="1877" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis delibrata subsp. khasiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="delibrata" subSpecies="khasiensis">Chloritis delibrata khasiensis</taxonomicName>
<bibRefCitation author="Ramakrishna," journalOrPublisher="Zoological Survey of India, Kolkata" pageId="0" pageNumber="1" refId="B34" refString="Ramakrishna, , Mitra, SC, Dey, A, 2010. Annotated checklist of Indian land molluscs. Zoological Survey of India, Kolkata" title="Annotated checklist of Indian land molluscs." year="2010">Ramakrishna et al., 2010</bibRefCitation>
: 326.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">&quot;Khasi Hills&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="types">
<paragraph pageId="0" pageNumber="1">Types.</paragraph>
<paragraph pageId="0" pageNumber="1">Not found in the NHM or in the ZSI (Sheikh Sajan, pers. comm. April 2020).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Nevill, G" journalOrPublisher="The Journal of the Asiatic Society of Bengal, Part II" pageId="0" pageNumber="1" pagination="14 - 41" refId="B24" refString="Nevill, G, 1877. List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species. The Journal of the Asiatic Society of Bengal, Part II 46 (1): 14 - 41" title="List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species." volume="46" year="1877">Nevill (1877)</bibRefCitation>
named the form illustrated in Hanley and
<normalizedToken originalValue="Theobalds">Theobald's</normalizedToken>
(1870) Conchologia Indica (pl. 14, fig. 9) as var.
<taxonomicName baseAuthorityName="G. Nevill" baseAuthorityYear="1877" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis var. khasiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="delibrata" variety="khasiensis">Chloritis khasiensis</taxonomicName>
, and mentioned that it has more raised and rounded whorls and a less open umbilicus than the typical
<taxonomicName lsidName="H. delibrata" pageId="0" pageNumber="1" rank="species" species="delibrata">
<emphasis italics="true" pageId="0" pageNumber="1">H. delibrata</emphasis>
</taxonomicName>
.
<bibRefCitation author="Nevill, G" journalOrPublisher="The Journal of the Asiatic Society of Bengal, Part II" pageId="0" pageNumber="1" pagination="14 - 41" refId="B24" refString="Nevill, G, 1877. List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species. The Journal of the Asiatic Society of Bengal, Part II 46 (1): 14 - 41" title="List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species." volume="46" year="1877">
<normalizedToken originalValue="Nevills">Nevill's</normalizedToken>
(1877)
</bibRefCitation>
description was as follows: &quot;The raised and rounded whorls, less open umbilicus, and contracted aperture will distinguish the form; it has sometimes a single brown band, but is often without it; it is tolerably abundant in the Naga and Khasi Hills. (...) Var.
<taxonomicName baseAuthorityName="G. Nevill" baseAuthorityYear="1877" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis var. khasiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="delibrata" variety="khasiensis">Chloritis khasiensis</taxonomicName>
, from Khasi Hills, axis 8.5, diam. 19.5 (apert. 9, diam 10.5) mm&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely" authorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Bouchetcamaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bouchetcamaena fusca" order="Testacea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fusca">
<emphasis italics="true" pageId="0" pageNumber="1">Bouchetcamaena fusca</emphasis>
</taxonomicName>
sp. nov. agrees with some parts of
<normalizedToken originalValue="Nevills">Nevill's</normalizedToken>
description (more raised spire and narrower umbilicus than in other similar species), and the Naga and Khasi Hills also match, but the specimens we described above as
<taxonomicName lsidName="B. fusca" pageId="0" pageNumber="1" rank="species" species="fusca">
<emphasis italics="true" pageId="0" pageNumber="1">B. fusca</emphasis>
</taxonomicName>
sp. nov. possessed no band and are all smaller than the size mentioned by Nevill. Furthermore, the shell illustrated by Hanley and Theobald (1870) possesses a subsutural furrow (a furrow running between the middle of the body whorl and the suture), which is absent in all
<taxonomicName lsidName="B. fusca" pageId="0" pageNumber="1" rank="species" species="fusca">
<emphasis italics="true" pageId="0" pageNumber="1">B. fusca</emphasis>
</taxonomicName>
sp. nov. shells we examined. Thus, we decided to describe these specimens as a new species and consider the name
<taxonomicName authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Trachia) delibrata subsp. var. var. khasiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="delibrata" subGenus="Trachia" subSpecies="var." variety="khasiensis">Helix (Trachia) delibrata var. khasiensis</taxonomicName>
as a taxon inquirendum.
</paragraph>
</subSubSection>
</treatment>
</document>