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<mods:title id="2C91D1223FE37AC230218ACDED6B2B44">Adding the West-African riverine component: Revision of the Recent freshwater snails belonging to Pseudocleopatra Thiele, 1928 (Caenogastropoda, Cerithioidea, Paludomidae)</mods:title>
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<taxonomicName id="27AE4D7DF632C514FF6C65C377BECF2A" authority="Mandahl-Barth, 1973" authorityName="Mandahl-Barth" authorityYear="1973" box="[151,727,1788,1815]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF632C514FF6C65C377BECF2A" bold="true" box="[151,727,1788,1815]" pageId="11" pageNumber="312">
<emphasis id="D2DAEAECF632C514FF6C65C374A8CF2A" bold="true" box="[151,449,1788,1815]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra voltana</emphasis>
<bibRefCitation id="843F4B0FF632C514FE3365C277BECF2A" author="Mandahl-Barth, G." box="[456,727,1789,1815]" pageId="11" pageNumber="312" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
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<subSubSection id="A8B46575F632C514FF6C641E742BCF06" box="[151,322,1825,1851]" pageId="11" pageNumber="312" type="description">
<paragraph id="E01136FEF632C514FF6C641E742BCF06" blockId="11.[151,727,1788,1851]" box="[151,322,1825,1851]" pageId="11" pageNumber="312">
<figureCitation id="78952A7BF632C514FF6C641E7598CF06" box="[151,241,1825,1851]" captionStart="FIGURE 8" captionStartId="11.[151,250,926,951]" captionTargetBox="[481,1112,201,877]" captionTargetId="figure@11.[466,1121,184,877]" captionTargetPageId="11" captionText="FIGURE 8. Opercula of Pseudocleopatra Thiele, 1928 species. A. Lectotype of P. togoensis Thiele, 1928, ZMB 190048b. B. Paratype of Pseudocleopatra voltana Mandahl-Barth, 1973, Mogeyenga village, DBL 2591. C. Probable paratype of Pseudocleopatra dartevellei Mandahl-Barth, 1973, Matadi, DBL 2583. D. Pseudocleopatra broecki (Putzeys, 1899) n. comb., MCZ 171520. E. Lectotype of Pseudocleopatra broecki (Putzeys, 1899) n. comb., MRAC 47360. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3460148" httpUri="https://zenodo.org/record/3460148/files/figure.png" pageId="11" pageNumber="312">Figs 8B</figureCitation>
,
<figureCitation id="78952A7BF632C514FF06641E742BCF06" box="[253,322,1825,1851]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStartId-0="14.[151,250,1189,1214]" captionStartId-1="15.[151,250,1003,1028]" captionStartId-2="15.[151,250,1947,1972]" captionStartId-3="16.[151,250,814,839]" captionTargetBox-0="[202,1361,196,1152]" captionTargetBox-1="[253,1334,174,982]" captionTargetBox-2="[250,1337,1116,1928]" captionTargetBox-3="[486,1101,174,795]" captionTargetId-0="figure@14.[183,1405,174,1170]" captionTargetId-1="figure@15.[253,1334,174,984]" captionTargetId-2="figure@15.[250,1337,1114,1928]" captionTargetId-3="figure@16.[486,1101,174,795]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="15" captionTargetPageId-3="16" captionText-0="FIGURE 11. Shells of Pseudocleopatra voltana Mandahl-Barth, 1973. A. Holotype, DBL 2594-1. B. Paratype, from the type locality; DBL 2594-2. C. Paratype, from the type locality, DBL 2594-3. D. Paratype, Ghana, NE region, without exact locality, DBL 2593. E. Paratype, Mogeyenga village, DBL 2591. F. Paralectotype of Pseudocleopatra togoensis Thiele, 1928, ZMB 190049. Scale bar: 5 mm." captionText-1="FIGURE 12. Radula of Pseudocleopatra voltana Mandahl-Barth, 1973, paratype, Apaso village, DBL 2592. A. Radula ribbon, scale bar: 25 μm. B. Lateral and central teeth; scale bar: 5 μm. C. Cutting edge of lateral and central teeth; scale bar: 3 μm. D. Central, lateral and marginal teeth; scale bar: 5 μm." captionText-2="FIGURE 13. Radula of Pseudocleopatra voltana Mandahl-Barth, 1973, paratype, Daboya, DBL 2594. A. Radula ribbon, scale bar: 25 μm. B. Lateral and central teeth; scale bar: 20 μm. C. Cutting edge of lateral and central teeth; scale bar: 6 μm. D. Central, lateral and marginal teeth; scale bar: 8 μm." captionText-3="FIGURE 14. Known occurrence records of Pseudocleopatra voltana Mandahl-Barth, 1973 in the Volta River system. The asterisk indicates the type locality; open symbols indicate samples of dry material. The arrow indicates the approximate location of the Akosombo dam." figureDoi-0="http://doi.org/10.5281/zenodo.3460154" figureDoi-1="http://doi.org/10.5281/zenodo.3460156" figureDoi-2="http://doi.org/10.5281/zenodo.3460158" figureDoi-3="http://doi.org/10.5281/zenodo.3460162" httpUri-0="https://zenodo.org/record/3460154/files/figure.png" httpUri-1="https://zenodo.org/record/3460156/files/figure.png" httpUri-2="https://zenodo.org/record/3460158/files/figure.png" httpUri-3="https://zenodo.org/record/3460162/files/figure.png" pageId="11" pageNumber="312">1114</figureCitation>
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<paragraph id="E01136FEF632C514FF6C645A7680CF40" blockId="11.[151,1436,1892,2009]" box="[151,1001,1892,1917]" pageId="11" pageNumber="312">
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<taxonomicName id="27AE4D7DF632C514FF6C645A770BCF40" authority="Thiele, 1928: 394" authorityName="Thiele" authorityPageNumber="394" authorityYear="1928" box="[151,610,1892,1917]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="togoensis">
<emphasis id="D2DAEAECF632C514FF6C645A74C0CF40" box="[151,425,1893,1917]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra togoensis</emphasis>
<treatmentCitation id="610F10EFF632C514FE54645B770BCF40" author="Thiele, J." box="[431,610,1892,1917]" page="394" pageId="11" pageNumber="312" year="1928">
<bibRefCitation id="843F4B0FF632C514FE54645B770BCF40" author="Thiele, J." box="[431,610,1892,1917]" pageId="11" pageNumber="312" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele, 1928: 394</bibRefCitation>
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(in part);
<treatmentCitation id="610F10EFF632C514FD33645A76EBCF40" author="Brown, D. S." box="[712,898,1893,1917]" page="111" pageId="11" pageNumber="312" year="1980">
<bibRefCitation id="843F4B0FF632C514FD33645A76EBCF40" author="Brown, D. S." box="[712,898,1893,1917]" pageId="11" pageNumber="312" refId="ref13993" refString="Brown, D. S. (1980) Freshwater snails of Africa and their medical importance. Taylor &amp; Francis, London, 487 pp." type="book" year="1980">Brown (1980: 111</bibRefCitation>
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, fig. 63e).
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<taxonomicName id="27AE4D7DF632C514FF6C64BB778BCFA1" authority="Mandahl-Barth, 1973: 279 - 281" authorityName="Mandahl-Barth" authorityPageNumber="279" authorityYear="1973" box="[151,738,1923,1948]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF632C514FF6C64BB74FFCFA1" box="[151,406,1924,1948]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra voltana</emphasis>
<bibRefCitation id="843F4B0FF632C514FE6664BC778BCFA1" author="Mandahl-Barth, G." box="[413,738,1923,1948]" pageId="11" pageNumber="312" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973: 279281</bibRefCitation>
</taxonomicName>
, fig. 3a, h;
<bibRefCitation id="843F4B0FF632C514FCAC64BB7124CFA1" author="Brown, D. S." box="[855,1101,1924,1948]" pageId="11" pageNumber="312" refId="ref13993" refString="Brown, D. S. (1980) Freshwater snails of Africa and their medical importance. Taylor &amp; Francis, London, 487 pp." type="book" year="1980">Brown (1980: 111112)</bibRefCitation>
;
<bibRefCitation id="843F4B0FF632C514FBA264BB70F1CFA1" author="Brown, D. S. &amp; Kristensen, T. K." box="[1113,1432,1923,1948]" pageId="11" pageNumber="312" refId="ref14054" refString="Brown, D. S. &amp; Kristensen, T. K. (1993) A field guide to African freshwater snails. Vol. 1. West African species. Danish Bilharziasis Laboratory, Charlottenlund, 2 + 55 pp." type="book" year="1993">Brown &amp; Kristensen (1993: 28</bibRefCitation>
, 51);
<bibRefCitation id="843F4B0FF632C514FF03649C7480CF87" author="Brown, D. S." box="[248,489,1954,1978]" pageId="11" pageNumber="312" refId="ref14020" refString="Brown, D. S. (1994) Freshwater snails of Africa and their medical importance. Revised 2 nd edition. Taylor &amp; Francis, London, x + 608 pp." type="book" year="1994">Brown (1994: 128129</bibRefCitation>
, 537, 580, fig. 61f);
<bibRefCitation id="843F4B0FF632C514FD44649D76B5CF86" author="Glaubrecht, M." box="[703,988,1954,1979]" pageId="11" pageNumber="312" refId="ref14270" refString="Glaubrecht, M. (1996) Evolutionsokologie und Systematik am Beispiel von Suss- und Brackwasserschnecken (Mollusca: Caenogastropoda: Cerithioidea): Ontogenese-Strategien, palaontologische Befunde und historische Zoogeographie. Backhuys Publishers, Leiden, xvi + 499 pp., 25 pls." type="book" year="1996">Glaubrecht (1996: 144145</bibRefCitation>
, 147148, fig. 24b, c);
<bibRefCitation id="843F4B0FF632C514FB35649D7439CFE4" author="Kristensen, T. K. &amp; Stensgaard, A. - S." pageId="11" pageNumber="312" pagination="6007457" refId="ref15373" refString="Kristensen, T. K. &amp; Stensgaard, A. - S. (2010 a) Pseudocleopatra voltana. The IUCN Red List of Threatened Species, 2010, e. T 165376 A 6007457." type="journal article" year="2010">Kristensen &amp; Stensgaard (2010a</bibRefCitation>
: e.T165376A6007457).
</paragraph>
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<caption id="B4D16676F635C513FF6C67B17766CCD2" ID-DOI="http://doi.org/10.5281/zenodo.3460150" ID-Zenodo-Dep="3460150" httpUri="https://zenodo.org/record/3460150/files/figure.png" pageId="12" pageNumber="313" startId="12.[151,250,1166,1191]" targetBox="[151,1436,195,1088]" targetPageId="12">
<paragraph id="E01136FEF635C513FF6C67B17766CCD2" blockId="12.[151,1436,1166,1263]" pageId="12" pageNumber="313">
<emphasis id="D2DAEAECF635C513FF6C67B1747ACC9A" bold="true" box="[151,275,1166,1191]" pageId="12" pageNumber="313">FIGURE 9.</emphasis>
<taxonomicName id="27AE4D7DF635C513FEE167B1740BCC9A" box="[282,354,1166,1191]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="12" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
of
<taxonomicName id="27AE4D7DF635C513FE7E67B07648CC9A" authority="Thiele, 1928" authorityName="Thiele" authorityYear="1928" box="[389,801,1166,1191]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="12" pageNumber="313" phylum="Mollusca" rank="species" species="togoensis">
<emphasis id="D2DAEAECF635C513FE7E67B077F1CC9A" box="[389,664,1167,1191]" italics="true" pageId="12" pageNumber="313">Pseudocleopatra togoensis</emphasis>
<bibRefCitation id="843F4B0FF635C513FD6567B17648CC9A" author="Thiele, J." box="[670,801,1166,1191]" pageId="12" pageNumber="313" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele, 1928</bibRefCitation>
</taxonomicName>
, Ghana, NE region, DBL 214.
<emphasis id="D2DAEAECF635C513FBA667B1711FCC9A" bold="true" box="[1117,1142,1166,1191]" pageId="12" pageNumber="313">A.</emphasis>
<taxonomicName id="27AE4D7DF635C513FB8667B171ACCC9A" box="[1149,1221,1166,1191]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="12" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
ribbon; scale bar: 20 μm.
<emphasis id="D2DAEAECF635C513FF3F678C75B2CCF6" bold="true" box="[196,219,1203,1227]" pageId="12" pageNumber="313">B.</emphasis>
Lateral and central teeth; scale bar: 10 μm.
<emphasis id="D2DAEAECF635C513FD6B678D77C1CCF6" bold="true" box="[656,680,1202,1227]" pageId="12" pageNumber="313">C.</emphasis>
Cutting edge of lateral and central teeth; scale bar: 2 μm.
<emphasis id="D2DAEAECF635C513FB12678C7068CCF6" bold="true" box="[1257,1281,1203,1227]" pageId="12" pageNumber="313">D.</emphasis>
Central, lateral and marginal teeth; scale bar: 20 μm.
</paragraph>
</caption>
<caption id="B4D16676F635C513FF6C64457136CF8A" ID-DOI="http://doi.org/10.5281/zenodo.3460152" ID-Zenodo-Dep="3460152" httpUri="https://zenodo.org/record/3460152/files/figure.png" pageId="12" pageNumber="313" startId="12.[151,250,1914,1939]" targetBox="[485,1102,1278,1895]" targetPageId="12">
<paragraph id="E01136FEF635C513FF6C64457136CF8A" blockId="12.[151,1436,1914,1975]" pageId="12" pageNumber="313">
<emphasis id="D2DAEAECF635C513FF6C6445744BCFAE" bold="true" box="[151,290,1914,1939]" pageId="12" pageNumber="313">FIGURE 10.</emphasis>
Known occurrence records of
<taxonomicName id="27AE4D7DF635C513FD9F6445716BCFAE" authority="Thiele, 1928" authorityName="Thiele" authorityYear="1928" box="[612,1026,1914,1939]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="12" pageNumber="313" phylum="Mollusca" rank="species" species="togoensis">
<emphasis id="D2DAEAECF635C513FD9F64457611CFAF" box="[612,888,1914,1938]" italics="true" pageId="12" pageNumber="313">Pseudocleopatra togoensis</emphasis>
<bibRefCitation id="843F4B0FF635C513FC7B6445716BCFAE" author="Thiele, J." box="[896,1026,1914,1939]" pageId="12" pageNumber="313" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele, 1928</bibRefCitation>
</taxonomicName>
in the Volta River system. The asterisk indicates the type locality. The arrow indicates the approximate location of the Akosombo dam.
</paragraph>
</caption>
<caption id="B4D16676F634C512FF6C63A57771C8C7" ID-Table-UUID="B4D16676F634C512FF6C63A57771C8C7" httpUri="http://table.plazi.org/id/B4D16676F634C512FF6C63A57771C8C7" pageId="13" pageNumber="314" targetBox="[166,1298,261,437]" targetIsTable="true" targetPageId="13">
<paragraph id="E01136FEF634C512FF6C63A57771C8C7" blockId="13.[151,1436,154,250]" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FF6C63A57463C88F" bold="true" box="[151,266,154,178]" pageId="13" pageNumber="314">TABLE 1.</emphasis>
Pairwise correct classifications based on linear discriminant analyses of h/w and h/la ratios of
<taxonomicName id="27AE4D7DF634C512FB0563A575DFC8EB" authorityName="Thiele" authorityYear="1928" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="genus">
<emphasis id="D2DAEAECF634C512FB0563A575DFC8EB" italics="true" pageId="13" pageNumber="314">Pseudocleopatra</emphasis>
</taxonomicName>
species. Values for species from the same river system, i.e. Volta River or Congo River, in bold. For abbreviations of measured shell variables, see Fig. 2.
</paragraph>
</caption>
<paragraph id="E01136FEF634C512FF5D623A7462C988" pageId="13" pageNumber="314">
<table id="92AEC45EF6343AE0FF5D623A707BC988" box="[166,1298,261,437]" gridcols="5" gridrows="5" pageId="13" pageNumber="314">
<tr id="5E9E34BCF6343AE0FF5D623A707BC923" box="[166,1298,261,286]" gridrow="0" pageId="13" pageNumber="314">
<th id="1D4F5DC0F6343AE0FF5D623A7443C923" box="[166,298,261,286]" gridcol="0" gridrow="0" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FF5D623A759CC923" bold="true" box="[166,245,261,286]" pageId="13" pageNumber="314">Species</emphasis>
</th>
<th id="1D4F5DC0F6343AE0FE5C623A7748C923" box="[423,545,261,286]" gridcol="1" gridrow="0" pageId="13" pageNumber="314">
<taxonomicName id="27AE4D7DF634C512FE5C62397748C923" authorityName="Thiele" authorityYear="1928" box="[423,545,262,286]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="togoensis">
<emphasis id="D2DAEAECF634C512FE5C62397748C923" bold="true" box="[423,545,262,286]" italics="true" pageId="13" pageNumber="314">P. togoensis</emphasis>
</taxonomicName>
</th>
<th id="1D4F5DC0F6343AE0FD52623A7666C923" box="[681,783,261,286]" gridcol="2" gridrow="0" pageId="13" pageNumber="314">
<taxonomicName id="27AE4D7DF634C512FD5262397666C923" authorityName="Mandahl-Barth" authorityYear="1973" box="[681,783,261,286]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF634C512FD5262397666C923" bold="true" box="[681,783,261,286]" italics="true" pageId="13" pageNumber="314">P. voltana</emphasis>
</taxonomicName>
</th>
<th id="1D4F5DC0F6343AE0FC50623A7146C923" box="[939,1071,261,286]" gridcol="3" gridrow="0" pageId="13" pageNumber="314">
<taxonomicName id="27AE4D7DF634C512FC5062397146C923" authorityName="Mandahl-Barth" authorityYear="1973" box="[939,1071,261,286]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF634C512FC5062397146C923" bold="true" box="[939,1071,261,286]" italics="true" pageId="13" pageNumber="314">P. dartevellei</emphasis>
</taxonomicName>
</th>
<th id="1D4F5DC0F6343AE0FB57623A707BC923" box="[1196,1298,261,286]" gridcol="4" gridrow="0" pageId="13" pageNumber="314">
<taxonomicName id="27AE4D7DF634C512FB576239707BC923" authorityName="Neiber &amp; Kahl &amp; Wiggering &amp; Glaubrecht" authorityYear="2019" baseAuthorityName="Putzeys" baseAuthorityYear="1899" box="[1196,1298,261,286]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="broecki">
<emphasis id="D2DAEAECF634C512FB576239707BC923" bold="true" box="[1196,1298,261,286]" italics="true" pageId="13" pageNumber="314">P. broecki</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="5E9E34BCF6343AE0FF5D6214707BC979" box="[166,1298,299,324]" gridrow="1" pageId="13" pageNumber="314" rowspan-1="1">
<th id="1D4F5DC0F6343AE0FF5D62147443C979" box="[166,298,299,324]" gridcol="0" gridrow="1" pageId="13" pageNumber="314">
<taxonomicName id="27AE4D7DF634C512FF5D62137476C97E" authorityName="Thiele" authorityYear="1928" box="[166,287,299,323]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="togoensis">
<emphasis id="D2DAEAECF634C512FF5D62137476C97E" bold="true" box="[166,287,299,323]" italics="true" pageId="13" pageNumber="314">P. togoensis</emphasis>
</taxonomicName>
</th>
<td id="1D4F5DC0F6343AE0FD5262147666C979" box="[681,783,299,324]" gridcol="2" gridrow="1" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FD5262147792C979" bold="true" box="[681,763,299,324]" pageId="13" pageNumber="314">95.45%</emphasis>
</td>
<td id="1D4F5DC0F6343AE0FC5062147146C979" box="[939,1071,299,324]" gridcol="3" gridrow="1" pageId="13" pageNumber="314">76.79%</td>
<td id="1D4F5DC0F6343AE0FB576214707BC979" box="[1196,1298,299,324]" gridcol="4" gridrow="1" pageId="13" pageNumber="314">92.75%</td>
</tr>
<tr id="5E9E34BCF6343AE0FF5D626E707BC957" box="[166,1298,337,362]" gridrow="2" pageId="13" pageNumber="314" rowspan-1="1" rowspan-2="1">
<th id="1D4F5DC0F6343AE0FF5D626E7443C957" box="[166,298,337,362]" gridcol="0" gridrow="2" pageId="13" pageNumber="314">
<taxonomicName id="27AE4D7DF634C512FF5D626D7462C957" authorityName="Mandahl-Barth" authorityYear="1973" box="[166,267,337,362]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF634C512FF5D626D7462C957" bold="true" box="[166,267,337,362]" italics="true" pageId="13" pageNumber="314">P. voltana</emphasis>
</taxonomicName>
</th>
<td id="1D4F5DC0F6343AE0FC50626E7146C957" box="[939,1071,337,362]" gridcol="3" gridrow="2" pageId="13" pageNumber="314">97.95%</td>
<td id="1D4F5DC0F6343AE0FB57626E707BC957" box="[1196,1298,337,362]" gridcol="4" gridrow="2" pageId="13" pageNumber="314">99.73%</td>
</tr>
<tr id="5E9E34BCF6343AE0FF5D6249707BC9B2" box="[166,1298,374,399]" gridrow="3" pageId="13" pageNumber="314" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="1D4F5DC0F6343AE0FF5D62497443C9B2" box="[166,298,374,399]" gridcol="0" gridrow="3" pageId="13" pageNumber="314">
<taxonomicName id="27AE4D7DF634C512FF5D62487443C9B2" authorityName="Mandahl-Barth" authorityYear="1973" box="[166,298,374,399]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF634C512FF5D62487443C9B2" bold="true" box="[166,298,374,399]" italics="true" pageId="13" pageNumber="314">P. dartevellei</emphasis>
</taxonomicName>
</th>
<td id="1D4F5DC0F6343AE0FB576249707BC9B2" box="[1196,1298,374,399]" gridcol="4" gridrow="3" pageId="13" pageNumber="314">89.87%</td>
</tr>
<tr id="5E9E34BCF6343AE0FF5D62A3707BC988" box="[166,1298,412,437]" gridrow="4" pageId="13" pageNumber="314">
<th id="1D4F5DC0F6343AE0FF5D62A3707BC988" box="[166,1298,412,437]" colspan="5" colspanRight="4" gridcol="0" gridrow="4" pageId="13" pageNumber="314">
<taxonomicName id="27AE4D7DF634C512FF5D62A27462C988" authorityName="Neiber &amp; Kahl &amp; Wiggering &amp; Glaubrecht" authorityYear="2019" baseAuthorityName="Putzeys" baseAuthorityYear="1899" box="[166,267,412,437]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="broecki">P. broecki</taxonomicName>
</th>
</tr>
</table>
</paragraph>
<subSubSection id="A8B46575F634C512FF6C61347076CCED" pageId="13" pageNumber="314" type="materials_examined">
<paragraph id="E01136FEF634C512FF6C613474D0CA74" blockId="13.[151,1437,521,1953]" pageId="13" pageNumber="314">
<materialsCitation id="50C63CA3F634C512FF6C61347700CA19" ID-GBIF-Occurrence="2418059372" box="[151,617,521,549]" location="Daboya" municipality="River" pageId="13" pageNumber="314" specimenCount="1" stateProvince="Volta" typeStatus="holotype">
<emphasis id="D2DAEAECF634C512FF6C61347453CA19" bold="true" box="[151,314,521,549]" pageId="13" pageNumber="314">
<typeStatus id="3F15885CF634C512FF6C613475BBCA18" box="[151,210,523,549]" pageId="13" pageNumber="314">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="0075AC84F634C512FEB4613574F9CA19" box="[335,400,522,548]" pageId="13" pageNumber="314">River</collectingMunicipality>
<collectingRegion id="226AF81CF634C512FE60613574BFCA19" box="[411,470,522,548]" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta</collectingRegion>
at
<location id="E5716025F634C512FDF961357736CA19" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FDF961357736CA19" box="[514,607,522,548]" municipality="River" name="Daboya" pageId="13" pageNumber="314" stateProvince="Volta">Daboya</location>
</materialsCitation>
;
<materialsCitation id="50C63CA3F634C512FD82613574DCCA74" ID-GBIF-Occurrence="2418059384" country="Ghana" county="West Gonja District" latitude="9.535001" location="White" longLatPrecision="21" longitude="-1.38" municipality="Daboya" pageId="13" pageNumber="314" specimenCount="1" stateProvince="Northern" typeStatus="holotype">
<collectingCountry id="98B9766EF634C512FD82613577ACCA19" box="[633,709,522,548]" name="Ghana" pageId="13" pageNumber="314">Ghana</collectingCountry>
,
<collectingRegion id="226AF81CF634C512FD2F613576F0CA19" box="[724,921,522,548]" country="Ghana" name="Northern" pageId="13" pageNumber="314">Northern Region</collectingRegion>
,
<collectingCounty id="09704E72F634C512FC53613471FBCA19" box="[936,1170,522,549]" pageId="13" pageNumber="314">West Gonja District</collectingCounty>
,
<collectingMunicipality id="0075AC84F634C512FB5961357195CA19" box="[1186,1276,522,548]" pageId="13" pageNumber="314">Daboya</collectingMunicipality>
,
<location id="E5716025F634C512FAF061357026CA19" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FAF061357026CA19" box="[1291,1359,522,548]" country="Ghana" county="West Gonja District" latitude="9.535001" longLatPrecision="21" longitude="-1.38" municipality="Daboya" name="White" pageId="13" pageNumber="314" stateProvince="Northern">White</location>
<collectingRegion id="226AF81CF634C512FAA0613570F0CA19" box="[1371,1433,522,548]" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta</collectingRegion>
,
<geoCoordinate id="859A5039F634C512FF6C61117477CA74" box="[151,286,558,585]" degrees="9" direction="north" minutes="32" orientation="latitude" pageId="13" pageNumber="314" precision="15" seconds="06" value="9.535001">9°3206 N</geoCoordinate>
,
<geoCoordinate id="859A5039F634C512FED2611174DCCA74" box="[297,437,558,585]" degrees="1" direction="west" minutes="22" orientation="longitude" pageId="13" pageNumber="314" precision="15" seconds="48" value="-1.38">1°2248 W</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="E01136FEF634C512FF3C616C7103CBB0" blockId="13.[151,1437,521,1953]" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FF3C616C741CCA51" bold="true" box="[199,373,593,621]" pageId="13" pageNumber="314">Type material.</emphasis>
<materialsCitation id="50C63CA3F634C512FE87616D7068CA51" ID-GBIF-Occurrence="2423596352" box="[380,1281,593,621]" collectionCode="DBL, MRAC" collectorName="F. S. McCullough. On &amp; From &amp; All" country="Ghana" county="Krachi East District" latitude="9.535001" location="Lake" longLatPrecision="15" longitude="0.118" municipality="Apase" pageId="13" pageNumber="314" specimenCode="DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144" specimenCount="13" stateProvince="Volta Region" typeStatus="holotype">
<typeStatus id="3F15885CF634C512FE87616D748ECA51" box="[380,487,594,620]" pageId="13" pageNumber="314" type="holotype">Holotype</typeStatus>
,
<specimenCode id="B0089E85F634C512FE09616C7716CA51" box="[498,639,593,621]" collectionCode="DBL" pageId="13" pageNumber="314">DBL 2594-1</specimenCode>
and two
<typeStatus id="3F15885CF634C512FD1D616C763DCA50" box="[742,852,595,621]" pageId="13" pageNumber="314" type="paratype">paratypes</typeStatus>
,
<specimenCode id="B0089E85F634C512FCA4616C76B0CA51" box="[863,985,593,621]" collectionCode="DBL" pageId="13" pageNumber="314">DBL 2594</specimenCode>
-
<specimenCode id="B0089E85F634C512FC1B616E716DCA51" box="[992,1028,593,620]" collectionCode="DBL" pageId="13" pageNumber="314">2,3</specimenCode>
, leg.
<collectorName id="4D5B5328F634C512FBC4616D7068CA51" box="[1087,1281,594,620]" pageId="13" pageNumber="314">F.S. McCullough</collectorName>
</materialsCitation>
.
<materialsCitation id="50C63CA3F634C512FAF7616D7073CA89" ID-GBIF-Occurrence="2423596350" collectionCode="DBL, MRAC" collectorName="F. S. McCullough. On &amp; From &amp; All" country="Ghana" county="Krachi East District" latitude="9.535001" location="Lake" longLatPrecision="15" longitude="0.118" municipality="Apase" pageId="13" pageNumber="314" specimenCode="DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144" specimenCount="13" stateProvince="Volta Region" typeStatus="holotype">
On the original label all three specimens are indicated as “Type”.
<collectorName id="4D5B5328F634C512FCFF61487628CAAC" box="[772,833,631,657]" pageId="13" pageNumber="314">From</collectorName>
the original description it is clear, however, that the specimen depicted here in
<figureCitation id="78952A7BF634C512FE3C61A57740CA89" box="[455,553,666,692]" captionStart="FIGURE 11" captionStartId="14.[151,250,1189,1214]" captionTargetBox="[202,1361,196,1152]" captionTargetId="figure@14.[183,1405,174,1170]" captionTargetPageId="14" captionText="FIGURE 11. Shells of Pseudocleopatra voltana Mandahl-Barth, 1973. A. Holotype, DBL 2594-1. B. Paratype, from the type locality; DBL 2594-2. C. Paratype, from the type locality, DBL 2594-3. D. Paratype, Ghana, NE region, without exact locality, DBL 2593. E. Paratype, Mogeyenga village, DBL 2591. F. Paralectotype of Pseudocleopatra togoensis Thiele, 1928, ZMB 190049. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3460154" httpUri="https://zenodo.org/record/3460154/files/figure.png" pageId="13" pageNumber="314">Fig. 11A</figureCitation>
is the
<typeStatus id="3F15885CF634C512FD8361A577B3CA89" box="[632,730,666,692]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
originally selected by
<bibRefCitation id="843F4B0FF634C512FC2561A57073CA89" author="Mandahl-Barth, G." box="[990,1306,666,692]" pageId="13" pageNumber="314" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth (1973: 281)</bibRefCitation>
</materialsCitation>
.
<materialsCitation id="50C63CA3F634C512FADF61A57419CB1D" ID-GBIF-Occurrence="2423596349" collectionCode="DBL, MRAC" collectorName="F. S. McCullough. On &amp; From &amp; All" country="Ghana" county="Krachi East District" latitude="9.535001" location="Lake" longLatPrecision="15" longitude="0.118" municipality="Apase" pageId="13" pageNumber="314" specimenCode="DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144" specimenCount="13" stateProvince="Volta Region" typeStatus="paratype">
Additional
<typeStatus id="3F15885CF634C512FF6C6180746ECAE4" box="[151,263,703,729]" pageId="13" pageNumber="314" type="paratype">paratypes</typeStatus>
:
<collectorName id="4D5B5328F634C512FEF46181745ACAE5" box="[271,307,702,728]" pageId="13" pageNumber="314">All</collectorName>
from
<collectingCountry id="98B9766EF634C512FE8C618174ADCAE5" box="[375,452,702,728]" name="Ghana" pageId="13" pageNumber="314">Ghana</collectingCountry>
: “Mogeyenga village, NE-Region” (not localised),
<specimenCode id="B0089E85F634C512FBFB61807110CAE5" box="[1024,1145,701,729]" collectionCode="DBL" pageId="13" pageNumber="314">DBL 2591</specimenCode>
,
<specimenCount id="F6A8FD77F634C512FB7861807079CAE5" box="[1155,1296,702,729]" pageId="13" pageNumber="314" type="generic">7 specimens</specimenCount>
; “Apaso village” (=
<collectingRegion id="226AF81CF634C512FF0C61DD74E5CAC1" box="[247,396,738,764]" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta Region</collectingRegion>
,
<collectingCounty id="09704E72F634C512FE6D61DD771FCAC1" box="[406,630,738,765]" pageId="13" pageNumber="314">Krachi East District</collectingCounty>
,
<collectingMunicipality id="0075AC84F634C512FD8461DC77A1CAC0" box="[639,712,739,765]" pageId="13" pageNumber="314">Apase</collectingMunicipality>
,
<geoCoordinate id="859A5039F634C512FD2961DD7639CAC1" box="[722,848,738,764]" pageId="13" pageNumber="314">7°3133.6”</geoCoordinate>
,
<geoCoordinate id="859A5039F634C512FCA061DD7699CAC0" box="[859,1008,738,765]" degrees="0" direction="east" minutes="07" orientation="longitude" pageId="13" pageNumber="314" precision="1" seconds="04.8" value="0.118">0°0704.8” E</geoCoordinate>
, today inundated by
<location id="E5716025F634C512FB2C61DD7066CAC1" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FB2C61DD7066CAC1" box="[1239,1295,738,764]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="Lake" pageId="13" pageNumber="314" stateProvince="Volta Region">Lake</location>
<collectingRegion id="226AF81CF634C512FAEF61DD703ACAC0" box="[1300,1363,738,765]" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta</collectingRegion>
),
<specimenCode id="B0089E85F634C512FA9E61DC75BBCB1D" collectionCode="DBL" pageId="13" pageNumber="314">DBL 2592</specimenCode>
,
<specimenCount id="F6A8FD77F634C512FF1B60397419CB1D" box="[224,368,774,800]" pageId="13" pageNumber="314" type="generic">3 specimens</specimenCount>
</materialsCitation>
;
<materialsCitation id="50C63CA3F634C512FE85603970F1CB1D" ID-GBIF-Occurrence="2423596341" box="[382,1432,773,801]" collectionCode="DBL, MRAC" collectorName="F. S. McCullough. On &amp; From &amp; All" country="Ghana" county="Krachi East District" latitude="9.535001" location="Lake" longLatPrecision="15" longitude="0.118" municipality="Apase" pageId="13" pageNumber="314" specimenCode="DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144" specimenCount="13" stateProvince="Volta Region" typeStatus="holotype">
“Northeast region” (=
<location id="E5716025F634C512FD796039767ACB1D" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FD796039767ACB1D" box="[642,787,774,800]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="Northeastern" pageId="13" pageNumber="314" stateProvince="Volta Region">Northeastern</location>
<collectingCountry id="98B9766EF634C512FCE660397600CB1D" box="[797,873,774,800]" name="Ghana" pageId="13" pageNumber="314">Ghana</collectingCountry>
, without exact locality),
<specimenCode id="B0089E85F634C512FB726038706CCB1D" box="[1161,1285,773,801]" collectionCode="DBL" pageId="13" pageNumber="314">DBL 2593</specimenCode>
,
<specimenCount id="F6A8FD77F634C512FAE8603970F1CB1D" box="[1299,1432,774,800]" pageId="13" pageNumber="314" type="generic">1 specimen</specimenCount>
</materialsCitation>
;
<materialsCitation id="50C63CA3F634C512FF6C6015777BCB55" ID-GBIF-Occurrence="2423596345" collectionCode="DBL, MRAC" collectorName="F. S. McCullough. On &amp; From &amp; All" country="Ghana" county="Krachi East District" latitude="9.535001" location="Lake" longLatPrecision="15" longitude="0.118" municipality="Apase" pageId="13" pageNumber="314" specimenCode="DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144" specimenCount="13" stateProvince="Volta Region" typeStatus="holotype">
“White
<collectingRegion id="226AF81CF634C512FF0A60157445CB79" box="[241,300,810,836]" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta</collectingRegion>
at Daboya” (=
<collectingRegion id="226AF81CF634C512FE1A601577CACB79" box="[481,675,810,836]" country="Ghana" name="Northern" pageId="13" pageNumber="314">Northern Region</collectingRegion>
,
<location id="E5716025F634C512FD4B601476FCCB79" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FD4B601476FCCB79" box="[688,917,810,837]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="West Gonja District" pageId="13" pageNumber="314" stateProvince="Volta Region">West Gonja District</location>
,
<location id="E5716025F634C512FC5960157695CB79" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FC5960157695CB79" box="[930,1020,810,836]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="Daboya" pageId="13" pageNumber="314" stateProvince="Volta Region">Daboya</location>
,
<location id="E5716025F634C512FBF260157124CB79" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FBF260157124CB79" box="[1033,1101,810,836]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="White" pageId="13" pageNumber="314" stateProvince="Volta Region">White</location>
<collectingRegion id="226AF81CF634C512FBAD601571FDCB79" box="[1110,1172,810,836]" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta</collectingRegion>
,
<geoCoordinate id="859A5039F634C512FB5A60157041CB78" box="[1185,1320,810,837]" degrees="9" direction="north" minutes="32" orientation="latitude" pageId="13" pageNumber="314" precision="15" seconds="06" value="9.535001">9°3206” N</geoCoordinate>
,
<geoCoordinate id="859A5039F634C512FACE601575DCCB54" degrees="1" direction="west" minutes="22" orientation="longitude" pageId="13" pageNumber="314" precision="15" seconds="48" value="-1.38">1°2248” W</geoCoordinate>
),
<specimenCode id="B0089E85F634C512FF3260717413CB55" box="[201,378,846,872]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="13" pageNumber="314" type="Museum">MRAC 799144</specimenCode>
,
<specimenCount id="F6A8FD77F634C512FE7E6071777BCB55" box="[389,530,846,872]" pageId="13" pageNumber="314" type="generic">2 specimens</specimenCount>
</materialsCitation>
;
<materialsCitation id="50C63CA3F634C512FDE560717103CBB0" ID-GBIF-Occurrence="2423596342" collectionCode="DBL, MRAC" collectorName="F. S. McCullough. On &amp; From &amp; All" country="Ghana" county="Krachi East District" latitude="9.535001" location="Lake" longLatPrecision="15" longitude="0.118" municipality="Apase" pageId="13" pageNumber="314" specimenCode="DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144" specimenCount="13" stateProvince="Volta Region" typeStatus="holotype">
“Fluss
<collectingRegion id="226AF81CF634C512FD97607177CECB55" box="[620,679,846,872]" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta</collectingRegion>
bei Apaso (
<collectingCountry id="98B9766EF634C512FCD760717603CB55" box="[812,874,846,872]" name="Togo" pageId="13" pageNumber="314">Togo</collectingCountry>
)” (
<location id="E5716025F634C512FC7060717164CB55" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FC7060717164CB55" box="[907,1037,846,872]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="Volta River" pageId="13" pageNumber="314" stateProvince="Volta Region">Volta River</location>
near
<location id="E5716025F634C512FBB1607071FBCB54" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FBB1607071FBCB54" box="[1098,1170,847,873]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="Apaso" pageId="13" pageNumber="314" stateProvince="Volta Region">Apaso</location>
(
<collectingCountry id="98B9766EF634C512FB5A607171B4CB55" box="[1185,1245,846,872]" name="Togo" pageId="13" pageNumber="314">Togo</collectingCountry>
) =
<location id="E5716025F634C512FAF9607170F1CB55" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FAF9607170F1CB55" box="[1282,1432,846,872]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="Volta Region" pageId="13" pageNumber="314" stateProvince="Volta Region">Volta Region</location>
,
<location id="E5716025F634C512FF6C604D7410CBB1" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FF6C604D7410CBB1" box="[151,377,882,909]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="Krachi East District" pageId="13" pageNumber="314" stateProvince="Volta Region">Krachi East District</location>
,
<location id="E5716025F634C512FE79604C74A2CBB0" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FE79604C74A2CBB0" box="[386,459,883,909]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="Apase" pageId="13" pageNumber="314" stateProvince="Volta Region">Apase</location>
,
<geoCoordinate id="859A5039F634C512FE2D604D773DCBB1" box="[470,596,882,908]" pageId="13" pageNumber="314">7°3133.6”</geoCoordinate>
,
<geoCoordinate id="859A5039F634C512FD9B604D779FCBB0" box="[608,758,882,909]" degrees="0" direction="east" minutes="07" orientation="longitude" pageId="13" pageNumber="314" precision="1" seconds="04.8" value="0.118">0°0704.8” E</geoCoordinate>
, today inundated by
<location id="E5716025F634C512FC1B604D7137CBB0" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FC1B604D7137CBB0" box="[992,1118,882,909]" country="Ghana" county="Krachi East District" latitude="9.535001" longLatPrecision="15" longitude="0.118" municipality="Apase" name="Lake Volta" pageId="13" pageNumber="314" stateProvince="Volta Region">Lake Volta</location>
).
</materialsCitation>
</paragraph>
<paragraph id="E01136FEF634C512FF3C60AA7152CBC5" blockId="13.[151,1437,521,1953]" pageId="13" pageNumber="314">
<materialsCitation id="50C63CA3F634C512FF3C60AA715ECBC5" ID-GBIF-Occurrence="2418059360" collectionCode="ZMB" country="Togo" county="River Volta" location="Krachi East District" longitude="0.118" municipality="Apaso" pageId="13" pageNumber="314" specimenCode="ZMB 190049" specimenCount="1" stateProvince="Volta Region" typeStatus="syntype">
<emphasis id="D2DAEAECF634C512FF3C60AA7768CB8D" bold="true" box="[199,513,917,944]" pageId="13" pageNumber="314">Other material examined.</emphasis>
“Fluss Volta bei Apaso (
<collectingCountry id="98B9766EF634C512FCD960A97609CB8D" box="[802,864,918,944]" name="Togo" pageId="13" pageNumber="314">Togo</collectingCountry>
)” (
<collectingCounty id="09704E72F634C512FC7860A9716ECB8D" box="[899,1031,918,944]" pageId="13" pageNumber="314">River Volta</collectingCounty>
near
<collectingMunicipality id="0075AC84F634C512FBB260A871F8CB8C" box="[1097,1169,919,945]" pageId="13" pageNumber="314">Apaso</collectingMunicipality>
(
<collectingCountry id="98B9766EF634C512FB5A60A971B4CB8D" box="[1185,1245,918,944]" name="Togo" pageId="13" pageNumber="314">Togo</collectingCountry>
) =
<collectingCountry id="98B9766EF634C512FAF360A9703DCB8D" box="[1288,1364,918,944]" name="Ghana" pageId="13" pageNumber="314">Ghana</collectingCountry>
,
<collectingRegion id="226AF81CF634C512FA9960A97582CBE9" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta Region</collectingRegion>
,
<location id="E5716025F634C512FF03608574B4CBE9" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FF03608574B4CBE9" box="[248,477,954,981]" country="Togo" county="River Volta" longitude="0.118" municipality="Apaso" name="Krachi East District" pageId="13" pageNumber="314" stateProvince="Volta Region">Krachi East District</location>
,
<location id="E5716025F634C512FE1360847758CBE8" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FE1360847758CBE8" box="[488,561,955,981]" country="Togo" county="River Volta" longitude="0.118" municipality="Apaso" name="Apase" pageId="13" pageNumber="314" stateProvince="Volta Region">Apase</location>
,
<geoCoordinate id="859A5039F634C512FDC6608577D5CBE9" box="[573,700,954,980]" pageId="13" pageNumber="314">7°3133.6”</geoCoordinate>
,
<geoCoordinate id="859A5039F634C512FD3260857608CBE8" box="[713,865,954,981]" degrees="0" direction="east" minutes="07" orientation="longitude" pageId="13" pageNumber="314" precision="1" seconds="04.8" value="0.118">0°0704.8” E</geoCoordinate>
, today inundated by
<location id="E5716025F634C512FBAA608571E0CBE9" LSID="urn:lsid:plazi:treatment:680787E8F632C512FF6C65C377B6CF9D:E5716025F634C512FBAA608571E0CBE9" box="[1105,1161,954,980]" country="Togo" county="River Volta" longitude="0.118" municipality="Apaso" name="Lake" pageId="13" pageNumber="314" stateProvince="Volta Region">Lake</location>
<collectingRegion id="226AF81CF634C512FB6A608571B9CBE8" box="[1169,1232,954,981]" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta</collectingRegion>
),
<specimenCode id="B0089E85F634C512FB1E608470EBCBE9" box="[1253,1410,954,981]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="13" pageNumber="314" type="Museum">ZMB 190049</specimenCode>
,
<specimenCount id="F6A8FD77F634C512FA756085746DCBC5" pageId="13" pageNumber="314" type="generic">1 specimen</specimenCount>
. Note that this specimen is a former
<typeStatus id="3F15885CF634C512FD6560E0779CCBC4" box="[670,757,991,1017]" pageId="13" pageNumber="314" type="syntype">syntype</typeStatus>
of
<taxonomicName id="27AE4D7DF634C512FCE160E0715ECBC5" authority="Thiele, 1928" authorityName="Thiele" authorityYear="1928" box="[794,1079,990,1016]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="togoensis">
<emphasis id="D2DAEAECF634C512FCE160E076C9CBC5" box="[794,928,991,1016]" italics="true" pageId="13" pageNumber="314">P. togoensis</emphasis>
<bibRefCitation id="843F4B0FF634C512FC5D60E1715ECBC5" author="Thiele, J." box="[934,1079,990,1016]" pageId="13" pageNumber="314" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele, 1928</bibRefCitation>
</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph id="E01136FEF634C512FF3C673E7076CCED" blockId="13.[151,1437,521,1953]" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FF3C673E741ECC21" bold="true" box="[199,375,1025,1052]" pageId="13" pageNumber="314">
Shell (
<figureCitation id="78952A7BF634C512FEE8673E7400CC21" box="[275,361,1025,1052]" captionStart="FIGURE 11" captionStartId="14.[151,250,1189,1214]" captionTargetBox="[202,1361,196,1152]" captionTargetId="figure@14.[183,1405,174,1170]" captionTargetPageId="14" captionText="FIGURE 11. Shells of Pseudocleopatra voltana Mandahl-Barth, 1973. A. Holotype, DBL 2594-1. B. Paratype, from the type locality; DBL 2594-2. C. Paratype, from the type locality, DBL 2594-3. D. Paratype, Ghana, NE region, without exact locality, DBL 2593. E. Paratype, Mogeyenga village, DBL 2591. F. Paralectotype of Pseudocleopatra togoensis Thiele, 1928, ZMB 190049. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3460154" httpUri="https://zenodo.org/record/3460154/files/figure.png" pageId="13" pageNumber="314">Fig. 11</figureCitation>
).
</emphasis>
The shell of
<taxonomicName id="27AE4D7DF634C512FDE8673C77E1CC21" authorityName="Mandahl-Barth" authorityYear="1973" box="[531,648,1026,1053]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF634C512FDE8673C774DCC21" box="[531,548,1027,1052]" italics="true" pageId="13" pageNumber="314">P</emphasis>
.
<emphasis id="D2DAEAECF634C512FDCF673D77E1CC21" box="[564,648,1026,1052]" italics="true" pageId="13" pageNumber="314">voltana</emphasis>
</taxonomicName>
is reddish to dark brown or yellowish-ochre, sometimes with darker bands or a darker area near the columella, and has an ovate to elongate shape, with up to seven whorls and a pointed apex (if not eroded). The whorls are convex to almost flat, with 16 distinct spiral cords; in few specimens these ridges are only weakly developed. The aperture is holostome and ovate, higher than wide and slightly drawn out at the upper angle; outer margin sinuous in lateral view, columella slightly curved to almost straight and the basal margin evenly rounded (in the most mature specimens slightly produced). Umbilicus slit-like or closed.
</paragraph>
</subSubSection>
<subSubSection id="A8B46575F634C512FF3C67E476D5CEF5" pageId="13" pageNumber="314" type="description">
<paragraph id="E01136FEF634C512FF3C67E47446CD00" blockId="13.[151,1437,521,1953]" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FF3C67E4748DCCC9" bold="true" box="[199,484,1241,1269]" pageId="13" pageNumber="314">
Measurements (
<emphasis id="D2DAEAECF634C512FE7367E474F1CCC8" bold="true" box="[392,408,1243,1269]" italics="true" pageId="13" pageNumber="314">n</emphasis>
= 16).
</emphasis>
h: 8.52 ±
<quantity id="27569B1BF634C512FDA267E677D7CCC8" box="[601,702,1241,1269]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="13" pageNumber="314" unit="mm" value="1.53">1.53 mm</quantity>
(range: 6.012.0 mm); w: 4.83 ±
<quantity id="27569B1BF634C512FBBB67E571CCCCC8" box="[1088,1189,1242,1269]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="314" unit="mm" value="0.6">0.60 mm</quantity>
(range:
<quantity id="27569B1BF634C512FAFF67E670F9CCC8" box="[1284,1424,1241,1269]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.8" metricValueMin="3.7" pageId="13" pageNumber="314" unit="mm" value="4.75" valueMax="5.8" valueMin="3.7">3.75.8 mm</quantity>
); bw: 6.83 ±
<quantity id="27569B1BF634C512FEE067C174E9CD24" box="[283,384,1278,1305]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="13" pageNumber="314" unit="mm" value="0.89">0.89 mm</quantity>
(range:
<quantity id="27569B1BF634C512FE1B67C17704CD24" box="[480,621,1278,1305]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="8.3" metricValueMin="4.8" pageId="13" pageNumber="314" unit="mm" value="6.55" valueMax="8.3" valueMin="4.8">4.88.3 mm</quantity>
); la: 4.11 ±
<quantity id="27569B1BF634C512FD0D67C17635CD24" box="[758,860,1278,1305]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="13" pageNumber="314" unit="mm" value="0.43">0.43 mm</quantity>
(range:
<quantity id="27569B1BF634C512FC4767C17121CD24" box="[956,1096,1278,1305]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.8" metricValueMin="3.2" pageId="13" pageNumber="314" unit="mm" value="4.0" valueMax="4.8" valueMin="3.2">3.24.8 mm</quantity>
); wa: 2.72 ±
<quantity id="27569B1BF634C512FB2467C1702CCD24" box="[1247,1349,1278,1305]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="314" unit="mm" value="0.27">0.27 mm</quantity>
(range:
<quantity id="27569B1BF634C512FF6C661D744BCD00" box="[151,290,1314,1341]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.2" metricValueMin="2.2" pageId="13" pageNumber="314" unit="mm" value="2.7" valueMax="3.2" valueMin="2.2">2.23.2 mm</quantity>
).
</paragraph>
<paragraph id="E01136FEF634C512FF3C667A71D0CD5C" blockId="13.[151,1437,521,1953]" box="[199,1209,1349,1377]" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FF3C667A7417CD5D" bold="true" box="[199,382,1349,1377]" pageId="13" pageNumber="314">
Ratios (
<emphasis id="D2DAEAECF634C512FEDF6678745DCD5C" bold="true" box="[292,308,1351,1377]" italics="true" pageId="13" pageNumber="314">n</emphasis>
= 16).
</emphasis>
h/w: 1.75 ± 0.16 (range: 1.512.17); h/la: 2.06 ± 0.20 (range: 1.862.60).
</paragraph>
<paragraph id="E01136FEF634C512FF3C6656760DCD95" blockId="13.[151,1437,521,1953]" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FF3C665674AFCDB9" bold="true" box="[199,454,1385,1412]" pageId="13" pageNumber="314">
Operculum (
<figureCitation id="78952A7BF634C512FEA5665674DECDB9" box="[350,439,1385,1412]" captionStart="FIGURE 8" captionStartId="11.[151,250,926,951]" captionTargetBox="[481,1112,201,877]" captionTargetId="figure@11.[466,1121,184,877]" captionTargetPageId="11" captionText="FIGURE 8. Opercula of Pseudocleopatra Thiele, 1928 species. A. Lectotype of P. togoensis Thiele, 1928, ZMB 190048b. B. Paratype of Pseudocleopatra voltana Mandahl-Barth, 1973, Mogeyenga village, DBL 2591. C. Probable paratype of Pseudocleopatra dartevellei Mandahl-Barth, 1973, Matadi, DBL 2583. D. Pseudocleopatra broecki (Putzeys, 1899) n. comb., MCZ 171520. E. Lectotype of Pseudocleopatra broecki (Putzeys, 1899) n. comb., MRAC 47360. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3460148" httpUri="https://zenodo.org/record/3460148/files/figure.png" pageId="13" pageNumber="314">Fig. 8B</figureCitation>
).
</emphasis>
The operculum is corneous, with a paucispiral, rather large nucleus that can reach up to ⅓ of the maximum diameter and possesses a concentric margin.
</paragraph>
<paragraph id="E01136FEF634C512FF3C668C76D5CEF5" blockId="13.[151,1437,521,1953]" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FF3C668C7745CDF1" bold="true" box="[199,556,1457,1485]" pageId="13" pageNumber="314">
<taxonomicName id="27AE4D7DF634C512FF3C668C7477CDF0" box="[199,286,1459,1485]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="13" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
(
<figureCitation id="78952A7BF634C512FED4668E74DDCDF1" box="[303,436,1457,1484]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStartId-0="15.[151,250,1003,1028]" captionStartId-1="15.[151,250,1947,1972]" captionTargetBox-0="[253,1334,174,982]" captionTargetBox-1="[250,1337,1116,1928]" captionTargetId-0="figure@15.[253,1334,174,984]" captionTargetId-1="figure@15.[250,1337,1114,1928]" captionTargetPageId-0="15" captionTargetPageId-1="15" captionText-0="FIGURE 12. Radula of Pseudocleopatra voltana Mandahl-Barth, 1973, paratype, Apaso village, DBL 2592. A. Radula ribbon, scale bar: 25 μm. B. Lateral and central teeth; scale bar: 5 μm. C. Cutting edge of lateral and central teeth; scale bar: 3 μm. D. Central, lateral and marginal teeth; scale bar: 5 μm." captionText-1="FIGURE 13. Radula of Pseudocleopatra voltana Mandahl-Barth, 1973, paratype, Daboya, DBL 2594. A. Radula ribbon, scale bar: 25 μm. B. Lateral and central teeth; scale bar: 20 μm. C. Cutting edge of lateral and central teeth; scale bar: 6 μm. D. Central, lateral and marginal teeth; scale bar: 8 μm." figureDoi-0="http://doi.org/10.5281/zenodo.3460156" figureDoi-1="http://doi.org/10.5281/zenodo.3460158" httpUri-0="https://zenodo.org/record/3460156/files/figure.png" httpUri-1="https://zenodo.org/record/3460158/files/figure.png" pageId="13" pageNumber="314">Figs 1213</figureCitation>
,
<tableCitation id="AD2C0345F634C512FE39668C7777CDF1" box="[450,542,1457,1485]" captionStart="TABLE 2" captionStartId="17.[151,239,694,718]" captionTargetPageId="17" captionText="TABLE 2. Number of denticles on the different radula teeth of examined samples of Pseudocleopatra Thiele, 1928 and Cleopatra Troschel, 1857 species. Data for Cleopatra bulimoides (Olivier, 1804) from Glaubrecht (1996) and data for P." httpUri="http://table.plazi.org/id/B4D16676F628C50EFF6C618977F2CB2A" pageId="13" pageNumber="314" tableUuid="B4D16676F628C50EFF6C618977F2CB2A">Table 2</tableCitation>
).
</emphasis>
Due to the lack of alcohol material, already prepared radula slides from the collection of the DBL were examined. Two slides of specimens from two different localities were examined (DBL 2592 and DBL 2594). The rachidian tooth is of squarish shape, with a straight to slightly V-shaped upper edge. The number of denticles is variable, with a prominent mesocone and 45 flanking denticles on either side, or mesocone not prominent, i.e. about the same size as the flanking denticles. The lateral teeth have relatively short but straight lateral extensions. The cutting edge bears 1215 small denticles hardly differing in size and lacking a prominent mesocone. Inner and outer marginal teeth subequal in size and slightly curved. Both, the inner and outer marginal teeth bear 1012 denticles each that are of approximately the same size.
</paragraph>
</subSubSection>
<subSubSection id="A8B46575F634C512FF3C65EE77F1CF2D" pageId="13" pageNumber="314" type="distribution">
<paragraph id="E01136FEF634C512FF3C65EE77F1CF2D" blockId="13.[151,1437,521,1953]" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FF3C65EE74A0CED1" bold="true" box="[199,457,1745,1772]" pageId="13" pageNumber="314">
Distribution (
<figureCitation id="78952A7BF634C512FE9C65EE74D2CED1" box="[359,443,1745,1772]" captionStart="FIGURE 14" captionStartId="16.[151,250,814,839]" captionTargetBox="[486,1101,174,795]" captionTargetId="figure@16.[486,1101,174,795]" captionTargetPageId="16" captionText="FIGURE 14. Known occurrence records of Pseudocleopatra voltana Mandahl-Barth, 1973 in the Volta River system. The asterisk indicates the type locality; open symbols indicate samples of dry material. The arrow indicates the approximate location of the Akosombo dam." figureDoi="http://doi.org/10.5281/zenodo.3460162" httpUri="https://zenodo.org/record/3460162/files/figure.png" pageId="13" pageNumber="314">Fig. 14</figureCitation>
).
</emphasis>
<taxonomicName id="27AE4D7DF634C512FE3465ED778FCED1" authorityName="Mandahl-Barth" authorityYear="1973" box="[463,742,1746,1772]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF634C512FE3465ED778FCED1" box="[463,742,1746,1772]" italics="true" pageId="13" pageNumber="314">Pseudocleopatra voltana</emphasis>
</taxonomicName>
is known from the southern part of present day Lake
<collectingRegion id="226AF81CF634C512FAC865ED7007CED1" box="[1331,1390,1746,1772]" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta</collectingRegion>
and several localities along the White
<collectingRegion id="226AF81CF634C512FDE965C97724CF2D" box="[530,589,1782,1808]" country="Ghana" name="Volta" pageId="13" pageNumber="314">Volta</collectingRegion>
River.
</paragraph>
</subSubSection>
<subSubSection id="A8B46575F634C512FF3C642477B6CF9D" pageId="13" pageNumber="314" type="discussion">
<paragraph id="E01136FEF634C512FF3C642477B6CF9D" blockId="13.[151,1437,521,1953]" pageId="13" pageNumber="314">
<emphasis id="D2DAEAECF634C512FF3C64247452CF08" bold="true" box="[199,315,1819,1845]" pageId="13" pageNumber="314">Remarks.</emphasis>
<taxonomicName id="27AE4D7DF634C512FEB864257734CF09" authorityName="Mandahl-Barth" authorityYear="1973" box="[323,605,1818,1844]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF634C512FEB864257734CF09" box="[323,605,1818,1844]" italics="true" pageId="13" pageNumber="314">Pseudocleopatra voltana</emphasis>
</taxonomicName>
has a more elongate shell than the remaining three
<taxonomicName id="27AE4D7DF634C512FB536425700FCF09" authorityName="Thiele" authorityYear="1928" box="[1192,1382,1818,1844]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="13" pageNumber="314" phylum="Mollusca" rank="genus">
<emphasis id="D2DAEAECF634C512FB536425700FCF09" box="[1192,1382,1818,1844]" italics="true" pageId="13" pageNumber="314">Pseudocleopatra</emphasis>
</taxonomicName>
species. When the spiral sculpture is well-developed, the species is very distinctive and cannot be confused with any of its congeners. Only in few specimens the spiral sculpture was weak on the last whorl. Nothing is known about the ecology of the species and new records do not exist.
</paragraph>
</subSubSection>
</treatment>
</document>