494 lines
62 KiB
XML
494 lines
62 KiB
XML
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<mods:title id="AA1BED78AAE4522F1B55B7A5956A8773">Adding the West-African riverine component: Revision of the Recent freshwater snails belonging to Pseudocleopatra Thiele, 1928 (Caenogastropoda, Cerithioidea, Paludomidae)</mods:title>
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<taxonomicName id="27AE4D7DF630C516FF6C606D771FCB50" authority="Thiele, 1928" authorityName="Thiele" authorityYear="1928" box="[151,630,850,877]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="9" pageNumber="310" phylum="Mollusca" rank="species" species="togoensis">
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<emphasis id="D2DAEAECF630C516FF6C606D74BECB50" bold="true" box="[151,471,850,877]" italics="true" pageId="9" pageNumber="310">Pseudocleopatra togoensis</emphasis>
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<bibRefCitation id="843F4B0FF630C516FE25606C771FCB50" author="Thiele, J." box="[478,630,851,877]" pageId="9" pageNumber="310" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele, 1928</bibRefCitation>
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<figureCitation id="78952A7BF630C516FF6C604875B7CBAC" box="[151,222,887,913]" captionStart="FIGURE 7" captionStartId="10.[151,250,1507,1532]" captionTargetBox="[182,1406,227,1362]" captionTargetId="figure@10.[151,1436,191,1404]" captionTargetPageId="10" captionText="FIGURE 7. Shells of Pseudocleopatra togoensis Thiele, 1928. A. Lectotype, ZMB 190048a. B. Oti River at Bdakune village, DBL 2601. C. NE-Ghana, DBL 214. D. White Volta River, DBL 12761. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3460146" httpUri="https://zenodo.org/record/3460146/files/figure.png" pageId="9" pageNumber="310">Figs 7</figureCitation>
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,
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<figureCitation id="78952A7BF630C516FF1160487466CBAC" box="[234,271,887,913]" captionStart="FIGURE 8" captionStartId="11.[151,250,926,951]" captionTargetBox="[481,1112,201,877]" captionTargetId="figure@11.[466,1121,184,877]" captionTargetPageId="11" captionText="FIGURE 8. Opercula of Pseudocleopatra Thiele, 1928 species. A. Lectotype of P. togoensis Thiele, 1928, ZMB 190048b. B. Paratype of Pseudocleopatra voltana Mandahl-Barth, 1973, Mogeyenga village, DBL 2591. C. Probable paratype of Pseudocleopatra dartevellei Mandahl-Barth, 1973, Matadi, DBL 2583. D. Pseudocleopatra broecki (Putzeys, 1899) n. comb., MCZ 171520. E. Lectotype of Pseudocleopatra broecki (Putzeys, 1899) n. comb., MRAC 47360. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3460148" httpUri="https://zenodo.org/record/3460148/files/figure.png" pageId="9" pageNumber="310">8A</figureCitation>
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,
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<figureCitation id="78952A7BF630C516FEE06048743ACBAC" box="[283,339,887,913]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="12.[151,250,1166,1191]" captionStartId-1="12.[151,250,1914,1939]" captionTargetBox-0="[151,1436,195,1088]" captionTargetBox-1="[485,1102,1278,1895]" captionTargetId-0="figure@12.[151,1436,195,1088]" captionTargetId-1="figure@12.[485,1102,1278,1895]" captionTargetPageId-0="12" captionTargetPageId-1="12" captionText-0="FIGURE 9. Radula of Pseudocleopatra togoensis Thiele, 1928, Ghana, NE region, DBL 214. A. Radula ribbon; scale bar: 20 μm. B. Lateral and central teeth; scale bar: 10 μm. C. Cutting edge of lateral and central teeth; scale bar: 2 μm. D. Central, lateral and marginal teeth; scale bar: 20 μm." captionText-1="FIGURE 10. Known occurrence records of Pseudocleopatra togoensis Thiele, 1928 in the Volta River system. The asterisk indicates the type locality. The arrow indicates the approximate location of the Akosombo dam." figureDoi-0="http://doi.org/10.5281/zenodo.3460150" figureDoi-1="http://doi.org/10.5281/zenodo.3460152" httpUri-0="https://zenodo.org/record/3460150/files/figure.png" httpUri-1="https://zenodo.org/record/3460152/files/figure.png" pageId="9" pageNumber="310">9–10</figureCitation>
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<taxonomicName id="27AE4D7DF630C516FF6C608477C9CBEE" authority="Thiele, 1928: 394 - 395" authorityName="Thiele" authorityPageNumber="394" authorityYear="1928" box="[151,672,954,979]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="9" pageNumber="310" phylum="Mollusca" rank="species" species="togoensis">
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<emphasis id="D2DAEAECF630C516FF6C608474C2CBEE" box="[151,427,955,979]" italics="true" pageId="9" pageNumber="310">Pseudocleopatra togoensis</emphasis>
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<bibRefCitation id="843F4B0FF630C516FE48608577C9CBEE" author="Thiele, J." box="[435,672,954,979]" pageId="9" pageNumber="310" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele, 1928: 394–395</bibRefCitation>
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, 400, 402, textfig. 58, pl. 8, figs 18, 18a (in part);
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<bibRefCitation id="843F4B0FF630C516FB4C6085744ACBCF" author="Mandahl-Barth, G." pageId="9" pageNumber="310" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth (1973: 279–281</bibRefCitation>
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, fig.
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); Mandahl-Barth (1974: 355–356, fig. 2a);
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<bibRefCitation id="843F4B0FF630C516FCD160E57175CBCF" author="Brown, D. S." box="[810,1052,986,1010]" pageId="9" pageNumber="310" refId="ref13993" refString="Brown, D. S. (1980) Freshwater snails of Africa and their medical importance. Taylor & Francis, London, 487 pp." type="book" year="1980">Brown (1980: 111–112)</bibRefCitation>
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(in part);
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, 51);
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, 128, 537, 579, figs
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, 61f);
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<bibRefCitation id="843F4B0FF630C516FD0E60C776B9CC2D" author="Glaubrecht, M." box="[757,976,1016,1041]" pageId="9" pageNumber="310" refId="ref14270" refString="Glaubrecht, M. (1996) Evolutionsokologie und Systematik am Beispiel von Suss- und Brackwasserschnecken (Mollusca: Caenogastropoda: Cerithioidea): Ontogenese-Strategien, palaontologische Befunde und historische Zoogeographie. Backhuys Publishers, Leiden, xvi + 499 pp., 25 pls." type="book" year="1996">Glaubrecht (1996: 27</bibRefCitation>
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, 144–145, 147–148, fig. 24a, d);
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<bibRefCitation id="843F4B0FF630C516FAE460C674A0CC12" author="Van Damme, D. & Pickford, M." pageId="9" pageNumber="310" pagination="1 - 83" refId="ref17026" refString="Van Damme, D. & Pickford, M. (2003) The late Cenozoic Thiaridae (Mollusca, Gastropoda, Cerithioidea) of the Albertine Rift Valley (Uganda-Congo) and their bearing on the origin and evolution of the Tanganyikan thalassoid malacofauna. Hydrobiologia, 498, 1 - 83. https: // doi. org / 10.1023 / A: 1026298512117" type="journal article" year="2003">Van Damme & Pickford (2003: 54–55</bibRefCitation>
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, fig. 41b);
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<bibRefCitation id="843F4B0FF630C516FDC3672976EFCC12" author="Kristensen, T. K. & Stensgaard, A. - S." box="[568,902,1046,1071]" pageId="9" pageNumber="310" pagination="6017727" refId="ref15411" refString="Kristensen, T. K. & Stensgaard, A. - S. (2010 b) Pseudocleopatra togoensis. The IUCN Red List of Threatened Species, 2010, e. T 165404 A 6017727." type="journal article" year="2010">Kristensen & Stensgaard (2010b</bibRefCitation>
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: e.T165404A6017727).
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<taxonomicName id="27AE4D7DF630C516FF6C6709742ACC73" authorityName="Thiele" authorityYear="1928" box="[151,323,1078,1102]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="9" pageNumber="310" phylum="Mollusca" rank="genus">Pseudocleopatra</taxonomicName>
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voltana—
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<treatmentCitation id="610F10EFF630C516FE576709770CCC73" author="Brown, D. S." box="[428,613,1078,1102]" page="111" pageId="9" pageNumber="310" year="1980">
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<bibRefCitation id="843F4B0FF630C516FE576709770CCC73" author="Brown, D. S." box="[428,613,1078,1102]" pageId="9" pageNumber="310" refId="ref13993" refString="Brown, D. S. (1980) Freshwater snails of Africa and their medical importance. Taylor & Francis, London, 487 pp." type="book" year="1980">Brown (1980: 111</bibRefCitation>
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, fig. 63f).
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locality.
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</emphasis>
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“
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<collectingMunicipality id="0075AC84F630C516FEB1674374A2CCAB" box="[330,459,1148,1174]" pageId="9" pageNumber="310">Fluss Volta</collectingMunicipality>
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bei
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(
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)”;
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River in
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;
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<collectingRegion id="226AF81CF630C516FC4167437138CCAB" box="[954,1105,1148,1174]" country="Ghana" name="Volta" pageId="9" pageNumber="310">Volta Region</collectingRegion>
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,
|
||
<materialsCitation id="50C63CA3F630C516FBA56743764DCC87" ID-GBIF-Occurrence="2418059380" country="Ghana" location="Krachi East District" longitude="0.118" pageId="9" pageNumber="310" specimenCount="1" stateProvince="Volta" typeStatus="holotype">
|
||
<location id="E5716025F630C516FBA56743702DCCAB" LSID="urn:lsid:plazi:treatment:680787E8F630C514FF6C606D7031CE96:E5716025F630C516FBA56743702DCCAB" box="[1118,1348,1148,1175]" country="Ghana" longitude="0.118" name="Krachi East District" pageId="9" pageNumber="310" stateProvince="Volta">Krachi East District</location>
|
||
, Apase,
|
||
<geoCoordinate id="859A5039F630C516FF6C679F7473CC87" box="[151,282,1184,1210]" pageId="9" pageNumber="310">7°31’33.6’’</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="859A5039F630C516FEDE679F74D7CC86" box="[293,446,1184,1211]" degrees="0" direction="east" minutes="07" orientation="longitude" pageId="9" pageNumber="310" precision="1" seconds="04.8" value="0.118">0°07’04.8’’ E</geoCoordinate>
|
||
; today inundated by Lake
|
||
<collectingRegion id="226AF81CF630C516FD1D679F764DCC87" box="[742,804,1184,1210]" country="Ghana" name="Volta" pageId="9" pageNumber="310">Volta</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E01136FEF630C516FF3C67FB7478CD52" blockId="9.[151,1437,1147,2039]" pageId="9" pageNumber="310">
|
||
<emphasis id="D2DAEAECF630C516FF3C67FB741ECCE3" bold="true" box="[199,375,1219,1246]" pageId="9" pageNumber="310">Type material.</emphasis>
|
||
<typeStatus id="3F15885CF630C516FE7B67FA7499CCE2" box="[384,496,1221,1247]" pageId="9" pageNumber="310" type="lectotype">Lectotype</typeStatus>
|
||
(herein designated to preserve stability of nomenclature;
|
||
<figureCitation id="78952A7BF630C516FB7E67FB7189CCE3" box="[1157,1248,1220,1246]" captionStart="FIGURE 7" captionStartId="10.[151,250,1507,1532]" captionTargetBox="[182,1406,227,1362]" captionTargetId="figure@10.[151,1436,191,1404]" captionTargetPageId="10" captionText="FIGURE 7. Shells of Pseudocleopatra togoensis Thiele, 1928. A. Lectotype, ZMB 190048a. B. Oti River at Bdakune village, DBL 2601. C. NE-Ghana, DBL 214. D. White Volta River, DBL 12761. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3460146" httpUri="https://zenodo.org/record/3460146/files/figure.png" pageId="9" pageNumber="310">Fig. 7A</figureCitation>
|
||
),
|
||
<collectionCode id="86BFAE3BF630C516FB0D67FA705ACCE2" box="[1270,1331,1221,1247]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="9" pageNumber="310" type="Museum">ZMB</collectionCode>
|
||
190048a (one dry shell),
|
||
<collectionCode id="86BFAE3BF630C516FEAA67D674E7CD3E" box="[337,398,1257,1283]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="9" pageNumber="310" type="Museum">ZMB</collectionCode>
|
||
190048b (one operculum) and
|
||
<collectionCode id="86BFAE3BF630C516FD0767D67650CD3E" box="[764,825,1257,1283]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="9" pageNumber="310" type="Museum">ZMB</collectionCode>
|
||
190048c (one
|
||
<taxonomicName id="27AE4D7DF630C516FC1367D771D5CD3E" box="[1000,1212,1256,1283]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="9" pageNumber="304" phylum="Bryophyta" rank="species" species="preparation">radula preparation</taxonomicName>
|
||
), leg. W. Kört. The original type series contained two
|
||
<typeStatus id="3F15885CF630C516FDD8663277E1CD1A" box="[547,648,1293,1319]" pageId="9" pageNumber="310" type="syntype">syntypes</typeStatus>
|
||
, the specimen here designated as the
|
||
<typeStatus id="3F15885CF630C516FBC1663371C8CD1B" box="[1082,1185,1292,1318]" pageId="9" pageNumber="310" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="27AE4D7DF630C516FB316632703CCD1B" authorityName="Thiele" authorityYear="1928" box="[1226,1365,1293,1319]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="9" pageNumber="310" phylum="Mollusca" rank="species" species="togoensis">
|
||
<emphasis id="D2DAEAECF630C516FB31663271B2CD1B" box="[1226,1243,1293,1318]" italics="true" pageId="9" pageNumber="310">P</emphasis>
|
||
.
|
||
<emphasis id="D2DAEAECF630C516FB106632703CCD1B" box="[1259,1365,1293,1318]" italics="true" pageId="9" pageNumber="310">togoensis</emphasis>
|
||
</taxonomicName>
|
||
and a second specimen (
|
||
<collectionCode id="86BFAE3BF630C516FE9D660F74CDCD77" box="[358,420,1328,1354]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="9" pageNumber="310" type="Museum">ZMB</collectionCode>
|
||
190049; one shell and preserved body;
|
||
<figureCitation id="78952A7BF630C516FC92660F76A2CD77" box="[873,971,1328,1354]" captionStart="FIGURE 11" captionStartId="14.[151,250,1189,1214]" captionTargetBox="[202,1361,196,1152]" captionTargetId="figure@14.[183,1405,174,1170]" captionTargetPageId="14" captionText="FIGURE 11. Shells of Pseudocleopatra voltana Mandahl-Barth, 1973. A. Holotype, DBL 2594-1. B. Paratype, from the type locality; DBL 2594-2. C. Paratype, from the type locality, DBL 2594-3. D. Paratype, Ghana, NE region, without exact locality, DBL 2593. E. Paratype, Mogeyenga village, DBL 2591. F. Paralectotype of Pseudocleopatra togoensis Thiele, 1928, ZMB 190049. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3460154" httpUri="https://zenodo.org/record/3460154/files/figure.png" pageId="9" pageNumber="310">Fig. 11F</figureCitation>
|
||
). The latter, however, actually belongs to
|
||
<taxonomicName id="27AE4D7DF630C516FF6C666A7463CD53" authorityName="Mandahl-Barth" authorityYear="1973" box="[151,266,1364,1391]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="9" pageNumber="310" phylum="Mollusca" rank="species" species="voltana">
|
||
<emphasis id="D2DAEAECF630C516FF6C666A75C1CD53" box="[151,168,1365,1390]" italics="true" pageId="9" pageNumber="310">P</emphasis>
|
||
.
|
||
<emphasis id="D2DAEAECF630C516FF4D666B7463CD53" box="[182,266,1364,1390]" italics="true" pageId="9" pageNumber="310">voltana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E01136FEF630C516FF3C6648747DCE57" blockId="9.[151,1437,1147,2039]" pageId="9" pageNumber="310">
|
||
<emphasis id="D2DAEAECF630C516FF3C66487495CDAF" bold="true" box="[199,508,1399,1426]" pageId="9" pageNumber="310">Other material examined.</emphasis>
|
||
<materialsCitation id="50C63CA3F630C516FDF866477069CD8B" ID-GBIF-Occurrence="2418059376" collectionCode="DBL" collectorName="Lake" country="Ghana" latitude="7.61" location="Otiso" longLatPrecision="21" longitude="0.08194445" municipality="Krachi East District" pageId="9" pageNumber="310" specimenCode="DBL 2603" specimenCount="25" stateProvince="Volta">
|
||
<collectingCountry id="98B9766EF630C516FDF866477739CDAF" box="[515,592,1400,1426]" name="Ghana" pageId="9" pageNumber="310">Ghana</collectingCountry>
|
||
: “Otiso village, river Oti drift, among gravel” (=
|
||
<collectingRegion id="226AF81CF630C516FB8366477067CDAF" box="[1144,1294,1400,1426]" country="Ghana" name="Volta" pageId="9" pageNumber="310">Volta Region</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="0075AC84F630C516FAE266477587CD8B" pageId="9" pageNumber="310">Krachi East District</collectingMunicipality>
|
||
,
|
||
<location id="E5716025F630C516FF0266A37451CD8B" LSID="urn:lsid:plazi:treatment:680787E8F630C514FF6C606D7031CE96:E5716025F630C516FF0266A37451CD8B" box="[249,312,1436,1462]" country="Ghana" latitude="7.61" longLatPrecision="21" longitude="0.08194445" municipality="Krachi East District" name="Otiso" pageId="9" pageNumber="310" stateProvince="Volta">Otiso</location>
|
||
,
|
||
<geoCoordinate id="859A5039F630C516FEB866A374A0CD8A" box="[323,457,1436,1463]" degrees="7" direction="north" minutes="36" orientation="latitude" pageId="9" pageNumber="310" precision="15" seconds="36" value="7.61">7°36’36’’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="859A5039F630C516FE2F66A4773FCD8A" box="[468,598,1435,1463]" degrees="0" direction="east" minutes="04" orientation="longitude" pageId="9" pageNumber="310" precision="15" seconds="55" value="0.08194445">0°04’55’’ E</geoCoordinate>
|
||
, today inundated by
|
||
<collectorName id="4D5B5328F630C516FCC566A3761FCD8B" box="[830,886,1436,1462]" pageId="9" pageNumber="310">Lake</collectorName>
|
||
<collectingRegion id="226AF81CF630C516FC8066A376D3CD8A" box="[891,954,1436,1463]" country="Ghana" name="Volta" pageId="9" pageNumber="310">Volta</collectingRegion>
|
||
),
|
||
<specimenCode id="B0089E85F630C516FC3666A2712CCD8B" box="[973,1093,1436,1463]" collectionCode="DBL" pageId="9" pageNumber="310">DBL 2603</specimenCode>
|
||
,>
|
||
<specimenCount id="F6A8FD77F630C516FB9D66A47069CD8B" box="[1126,1280,1435,1462]" pageId="9" pageNumber="310" type="generic">25 specimens</specimenCount>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="50C63CA3F630C516FAF066A374CACDC3" ID-GBIF-Occurrence="2418059369" collectionCode="DBL" country="Ghana" county="Central Gonja District" latitude="9.140833" location="White" longLatPrecision="21" longitude="-1.158611" municipality="Yapei" pageId="9" pageNumber="310" specimenCode="DBL 127" specimenCount="9" stateProvince="Volta">
|
||
“White
|
||
<collectingRegion id="226AF81CF630C516FA9966A370F4CD8B" box="[1378,1437,1436,1462]" country="Ghana" name="Volta" pageId="9" pageNumber="310">Volta</collectingRegion>
|
||
River at Yapei Ferry” (=
|
||
<collectingRegion id="226AF81CF630C516FE4866FF771DCDE7" box="[435,628,1472,1498]" country="Ghana" name="Northern" pageId="9" pageNumber="310">Northern Region</collectingRegion>
|
||
,
|
||
<collectingCounty id="09704E72F630C516FD7B66FF7617CDE7" box="[640,894,1472,1498]" pageId="9" pageNumber="310">Central Gonja District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="0075AC84F630C516FC7266FF76A5CDE7" box="[905,972,1472,1498]" pageId="9" pageNumber="310">Yapei</collectingMunicipality>
|
||
,
|
||
<location id="E5716025F630C516FC2C66FF7172CDE7" LSID="urn:lsid:plazi:treatment:680787E8F630C514FF6C606D7031CE96:E5716025F630C516FC2C66FF7172CDE7" box="[983,1051,1472,1498]" country="Ghana" county="Central Gonja District" latitude="9.140833" longLatPrecision="21" longitude="-1.158611" municipality="Yapei" name="White" pageId="9" pageNumber="310" stateProvince="Volta">White</location>
|
||
<collectingRegion id="226AF81CF630C516FBD866FF7108CDE7" box="[1059,1121,1472,1498]" country="Ghana" name="Volta" pageId="9" pageNumber="310">Volta</collectingRegion>
|
||
,
|
||
<geoCoordinate id="859A5039F630C516FB9666FF719CCDE6" box="[1133,1269,1472,1499]" degrees="9" direction="north" minutes="08" orientation="latitude" pageId="9" pageNumber="310" precision="15" seconds="27" value="9.140833">9°08’27’’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="859A5039F630C516FAFA66FF70E6CDE6" box="[1281,1423,1472,1499]" degrees="1" direction="west" minutes="09" orientation="longitude" pageId="9" pageNumber="310" precision="15" seconds="31" value="-1.158611">1°09’31’’ W</geoCoordinate>
|
||
),
|
||
<specimenCode id="B0089E85F630C516FF6C66DA746DCDC3" box="[151,260,1508,1535]" collectionCode="DBL" pageId="9" pageNumber="310">DBL 127</specimenCode>
|
||
,
|
||
<specimenCount id="F6A8FD77F630C516FEE866DB74CACDC3" box="[275,419,1508,1534]" pageId="9" pageNumber="310" type="generic">9 specimens</specimenCount>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="50C63CA3F630C516FE4A66DB70F1CDC3" ID-GBIF-Occurrence="2418059382" box="[433,1432,1507,1535]" collectionCode="DBL" country="Ghana" county="Central Gonja District" isTailEnumeration="true" latitude="9.140833" location="Northeastern region" longLatPrecision="21" longitude="-1.158611" municipality="Mogeyenga village" pageId="9" pageNumber="310" specimenCode="DBL 2604" specimenCount="25" stateProvince="Volta">
|
||
“
|
||
<collectingMunicipality id="0075AC84F630C516FE4066DB77F2CDC3" box="[443,667,1508,1534]" pageId="9" pageNumber="310">Mogeyenga village</collectingMunicipality>
|
||
,
|
||
<location id="E5716025F630C516FD5266DB76E7CDC3" LSID="urn:lsid:plazi:treatment:680787E8F630C514FF6C606D7031CE96:E5716025F630C516FD5266DB76E7CDC3" box="[681,910,1508,1534]" municipality="Mogeyenga village" name="Northeastern region" pageId="9" pageNumber="310">Northeastern region</location>
|
||
” (not localised),
|
||
<specimenCode id="B0089E85F630C516FBAD66DA71BBCDC3" box="[1110,1234,1508,1535]" collectionCode="DBL" pageId="9" pageNumber="310">DBL 2604</specimenCode>
|
||
,>
|
||
<specimenCount id="F6A8FD77F630C516FB0166DC70F1CDC3" box="[1274,1432,1507,1534]" pageId="9" pageNumber="310" type="generic">25 specimens</specimenCount>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="50C63CA3F630C516FF6C653771D6CE1F" ID-GBIF-Occurrence="2418059363" box="[151,1215,1543,1571]" collectionCode="DBL" country="Ghana" location="Northeastern" pageId="9" pageNumber="310" specimenCode="DBL 214" specimenCount="25">
|
||
“Northeast region” (=
|
||
<location id="E5716025F630C516FE6D6537774ECE1F" LSID="urn:lsid:plazi:treatment:680787E8F630C514FF6C606D7031CE96:E5716025F630C516FE6D6537774ECE1F" box="[406,551,1544,1570]" country="Ghana" name="Northeastern" pageId="9" pageNumber="310">Northeastern</location>
|
||
<collectingCountry id="98B9766EF630C516FDD565377713CE1F" box="[558,634,1544,1570]" name="Ghana" pageId="9" pageNumber="310">Ghana</collectingCountry>
|
||
, without exact locality),
|
||
<specimenCode id="B0089E85F630C516FC6865367696CE1F" box="[915,1023,1544,1571]" collectionCode="DBL" pageId="9" pageNumber="310">DBL 214</specimenCode>
|
||
,>
|
||
<specimenCount id="F6A8FD77F630C516FBD8653871D6CE1F" box="[1059,1215,1543,1570]" pageId="9" pageNumber="310" type="generic">25 specimens</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="50C63CA3F630C516FB3165367466CE57" ID-GBIF-Occurrence="2418059383" collectionCode="DBL" country="Togo" latitude="10.35" location="Oti River" longLatPrecision="1298" longitude="0.48333332" municipality="Sansanne-Mango area" pageId="9" pageNumber="310" specimenCode="DBL 2601" specimenCount="4" stateProvince="Savanes">
|
||
<collectingCountry id="98B9766EF630C516FB3165367061CE1E" box="[1226,1288,1545,1571]" name="Togo" pageId="9" pageNumber="310">Togo</collectingCountry>
|
||
: “Oti river at Bdakune village, Sunson area” (=
|
||
<collectingRegion id="226AF81CF630C516FDE265137711CE7B" box="[537,632,1580,1606]" country="Togo" name="Savanes" pageId="9" pageNumber="310">Savanes</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="0075AC84F630C516FD7F65137617CE7A" box="[644,894,1580,1607]" pageId="9" pageNumber="310">Sansanné-Mango area</collectingMunicipality>
|
||
,
|
||
<location id="E5716025F630C516FC716513769FCE7B" LSID="urn:lsid:plazi:treatment:680787E8F630C514FF6C606D7031CE96:E5716025F630C516FC716513769FCE7B" box="[906,1014,1580,1606]" country="Togo" latitude="10.35" longLatPrecision="1298" longitude="0.48333332" municipality="Sansanne-Mango area" name="Oti River" pageId="9" pageNumber="310" stateProvince="Savanes">Oti River</location>
|
||
, ca.
|
||
<geoCoordinate id="859A5039F630C516FBD3651371F8CE7A" box="[1064,1169,1580,1607]" degrees="10" direction="north" minutes="21" orientation="latitude" pageId="9" pageNumber="310" precision="925" value="10.35">10°21’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="859A5039F630C516FB666513719CCE7A" box="[1181,1269,1580,1607]" degrees="0" direction="east" minutes="29" orientation="longitude" pageId="9" pageNumber="310" precision="925" value="0.48333332">0°29’ E</geoCoordinate>
|
||
),
|
||
<specimenCode id="B0089E85F630C516FAF2651270EACE7B" box="[1289,1411,1580,1607]" collectionCode="DBL" pageId="9" pageNumber="310">DBL 2601</specimenCode>
|
||
,
|
||
<specimenCount id="F6A8FD77F630C516FA7565137466CE57" pageId="9" pageNumber="310" type="generic">4 specimens</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E01136FEF630C516FF3C654C7704CF5A" blockId="9.[151,1437,1147,2039]" pageId="9" pageNumber="310">
|
||
<emphasis id="D2DAEAECF630C516FF3C654C7403CEB3" bold="true" box="[199,362,1651,1678]" pageId="9" pageNumber="310">
|
||
Shell (
|
||
<figureCitation id="78952A7BF630C516FEE8654C7432CEB3" box="[275,347,1651,1678]" captionStart="FIGURE 7" captionStartId="10.[151,250,1507,1532]" captionTargetBox="[182,1406,227,1362]" captionTargetId="figure@10.[151,1436,191,1404]" captionTargetPageId="10" captionText="FIGURE 7. Shells of Pseudocleopatra togoensis Thiele, 1928. A. Lectotype, ZMB 190048a. B. Oti River at Bdakune village, DBL 2601. C. NE-Ghana, DBL 214. D. White Volta River, DBL 12761. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3460146" httpUri="https://zenodo.org/record/3460146/files/figure.png" pageId="9" pageNumber="310">Fig. 7</figureCitation>
|
||
).
|
||
</emphasis>
|
||
The shell of
|
||
<taxonomicName id="27AE4D7DF630C516FDFE654A77F9CEB3" authorityName="Thiele" authorityYear="1928" box="[517,656,1653,1679]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="9" pageNumber="310" phylum="Mollusca" rank="species" species="togoensis">
|
||
<emphasis id="D2DAEAECF630C516FDFE654A777FCEB3" box="[517,534,1653,1678]" italics="true" pageId="9" pageNumber="310">P</emphasis>
|
||
.
|
||
<emphasis id="D2DAEAECF630C516FDDD654A77F9CEB3" box="[550,656,1653,1678]" italics="true" pageId="9" pageNumber="310">togoensis</emphasis>
|
||
</taxonomicName>
|
||
has an elongate to depressed ovoid shape with 3–4 slightly to wellrounded whorls with sinuous growth lines and sometimes 2–3 faint spiral ridges. The colour of the shells varies from a golden yellowish brown with slightly darker brown, sinuous radial stripes to brown or olive green to dark brown without radial bands. The spire height is about ¼ to ⅓ of the total shell height; the first whorls are often eroded. The aperture is holostome and ovate, higher than wide and slightly drawn out at the upper angle; outer margin sinuous in lateral view, columella curved and the basal margin evenly rounded (in the most mature specimens slightly produced). An umbilicus present, but narrow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8B46575F630C515FF3C644F74B4CFED" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310" type="description">
|
||
<paragraph id="E01136FEF630C516FF3C644F7446CFEE" blockId="9.[151,1437,1147,2039]" pageId="9" pageNumber="310">
|
||
<emphasis id="D2DAEAECF630C516FF3C644F748ACFB7" bold="true" box="[199,483,1903,1930]" pageId="9" pageNumber="310">
|
||
Measurements (
|
||
<emphasis id="D2DAEAECF630C516FE73644F74F1CFB7" bold="true" box="[392,408,1904,1930]" italics="true" pageId="9" pageNumber="310">n</emphasis>
|
||
= 88).
|
||
</emphasis>
|
||
h: 9.50 ±
|
||
<quantity id="27569B1BF630C516FDA2645077D7CFB6" box="[601,702,1903,1931]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="310" unit="mm" value="1.25">1.25 mm</quantity>
|
||
(range:
|
||
<quantity id="27569B1BF630C516FCE6645076DECFB6" box="[797,951,1903,1931]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="13.6" metricValueMin="5.6" pageId="9" pageNumber="310" unit="mm" value="9.6" valueMax="13.6" valueMin="5.6">5.6–13.6 mm</quantity>
|
||
); w: 6.61 ±
|
||
<quantity id="27569B1BF630C516FBBB644F71CCCFB6" box="[1088,1189,1904,1931]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="9" pageNumber="310" unit="mm" value="0.96">0.96 mm</quantity>
|
||
(range:
|
||
<quantity id="27569B1BF630C516FAFF644F70F9CFB6" box="[1284,1424,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="6.45" metricValueMax="8.8" metricValueMin="4.1" pageId="9" pageNumber="310" unit="mm" value="6.45" valueMax="8.8" valueMin="4.1">4.1–8.8 mm</quantity>
|
||
); bw: 8.08 ±
|
||
<quantity id="27569B1BF630C516FEE264AB7417CF92" box="[281,382,1940,1967]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="9" pageNumber="310" unit="mm" value="1.21">1.21 mm</quantity>
|
||
(range:
|
||
<quantity id="27569B1BF630C516FE2764AB771FCF92" box="[476,630,1940,1967]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="10.9" metricValueMin="4.3" pageId="9" pageNumber="310" unit="mm" value="7.6" valueMax="10.9" valueMin="4.3">4.3–10.9 mm</quantity>
|
||
); la: 5.45 ±
|
||
<quantity id="27569B1BF630C516FD0564AB760BCF92" box="[766,866,1940,1967]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="9" pageNumber="310" unit="mm" value="0.72">0.72 mm</quantity>
|
||
(range:
|
||
<quantity id="27569B1BF630C516FC3A64AB7125CF92" box="[961,1100,1940,1967]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="7.1" metricValueMin="3.2" pageId="9" pageNumber="310" unit="mm" value="5.15" valueMax="7.1" valueMin="3.2">3.2–7.1 mm</quantity>
|
||
); wa: 3.63 ±
|
||
<quantity id="27569B1BF630C516FB1A64AC702FCF92" box="[1249,1350,1939,1967]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="310" unit="mm" value="0.54">0.54 mm</quantity>
|
||
(range: 2.0–
|
||
<quantity id="27569B1BF630C516FF336487744BCFEE" box="[200,290,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="310" unit="mm" value="4.9">4.9 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E01136FEF630C516FF3C64E471DECFCB" blockId="9.[151,1437,1147,2039]" box="[199,1207,2011,2039]" pageId="9" pageNumber="310">
|
||
<emphasis id="D2DAEAECF630C516FF3C64E47414CFCB" bold="true" box="[199,381,2011,2038]" pageId="9" pageNumber="310">
|
||
Ratios (
|
||
<emphasis id="D2DAEAECF630C516FEDF64E3745DCFCB" bold="true" box="[292,308,2012,2038]" italics="true" pageId="9" pageNumber="310">n</emphasis>
|
||
= 88).
|
||
</emphasis>
|
||
h/w: 1.45 ± 0.11 (range: 1.16–1.69); h/la: 1.75 ± 0.12 (range: 1.49–2.05).
|
||
</paragraph>
|
||
<caption id="B4D16676F633C515FF6C66DC714ACE22" ID-DOI="http://doi.org/10.5281/zenodo.3460146" ID-Zenodo-Dep="3460146" httpUri="https://zenodo.org/record/3460146/files/figure.png" pageId="10" pageNumber="311" startId="10.[151,250,1507,1532]" targetBox="[182,1406,227,1362]" targetPageId="10">
|
||
<paragraph id="E01136FEF633C515FF6C66DC714ACE22" blockId="10.[151,1436,1507,1568]" pageId="10" pageNumber="311">
|
||
<emphasis id="D2DAEAECF633C515FF6C66DC747ACDC1" bold="true" box="[151,275,1507,1532]" pageId="10" pageNumber="311">FIGURE 7.</emphasis>
|
||
Shells of
|
||
<taxonomicName id="27AE4D7DF633C515FE8166DB767FCDC1" authority="Thiele, 1928" authorityName="Thiele" authorityYear="1928" box="[378,790,1507,1532]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="10" pageNumber="311" phylum="Mollusca" rank="species" species="togoensis">
|
||
<emphasis id="D2DAEAECF633C515FE8166DB77E4CDC1" box="[378,653,1508,1532]" italics="true" pageId="10" pageNumber="311">Pseudocleopatra togoensis</emphasis>
|
||
<bibRefCitation id="843F4B0FF633C515FD6866DC767FCDC1" author="Thiele, J." box="[659,790,1507,1532]" pageId="10" pageNumber="311" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="D2DAEAECF633C515FCDB66DC7650CDC1" bold="true" box="[800,825,1507,1532]" pageId="10" pageNumber="311">A.</emphasis>
|
||
Lectotype, ZMB 190048a.
|
||
<emphasis id="D2DAEAECF633C515FBA866DB7102CDC1" bold="true" box="[1107,1131,1508,1532]" pageId="10" pageNumber="311">B.</emphasis>
|
||
Oti River at Bdakune village, DBL 2601.
|
||
<emphasis id="D2DAEAECF633C515FEF46538744ECE1D" bold="true" box="[271,295,1543,1568]" pageId="10" pageNumber="311">C.</emphasis>
|
||
NE-Ghana, DBL 214.
|
||
<emphasis id="D2DAEAECF633C515FDEA65377743CE1D" bold="true" box="[529,554,1544,1568]" pageId="10" pageNumber="311">D.</emphasis>
|
||
White Volta River, DBL 12761. Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E01136FEF633C515FF3C654E7736CE8D" blockId="10.[151,1437,1649,2001]" pageId="10" pageNumber="311">
|
||
<emphasis id="D2DAEAECF633C515FF3C654E74AECEB1" bold="true" box="[199,455,1649,1676]" pageId="10" pageNumber="311">
|
||
Operculum (
|
||
<figureCitation id="78952A7BF633C515FEA5654E74D0CEB1" box="[350,441,1649,1676]" captionStart="FIGURE 8" captionStartId="11.[151,250,926,951]" captionTargetBox="[481,1112,201,877]" captionTargetId="figure@11.[466,1121,184,877]" captionTargetPageId="11" captionText="FIGURE 8. Opercula of Pseudocleopatra Thiele, 1928 species. A. Lectotype of P. togoensis Thiele, 1928, ZMB 190048b. B. Paratype of Pseudocleopatra voltana Mandahl-Barth, 1973, Mogeyenga village, DBL 2591. C. Probable paratype of Pseudocleopatra dartevellei Mandahl-Barth, 1973, Matadi, DBL 2583. D. Pseudocleopatra broecki (Putzeys, 1899) n. comb., MCZ 171520. E. Lectotype of Pseudocleopatra broecki (Putzeys, 1899) n. comb., MRAC 47360. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3460148" httpUri="https://zenodo.org/record/3460148/files/figure.png" pageId="10" pageNumber="311">Fig. 8A</figureCitation>
|
||
).
|
||
</emphasis>
|
||
The operculum is corneous, with a concentric, paucispiral, rather large nucleus that can reach up to ⅔ of the maximum diameter.
|
||
</paragraph>
|
||
<paragraph id="E01136FEF633C515FF3C658474B4CFED" blockId="10.[151,1437,1649,2001]" pageId="10" pageNumber="311">
|
||
<emphasis id="D2DAEAECF633C515FF3C65847499CEE9" bold="true" box="[199,496,1721,1749]" pageId="10" pageNumber="311">
|
||
<taxonomicName id="27AE4D7DF633C515FF3C65847477CEE8" box="[199,286,1723,1749]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="10" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
|
||
(
|
||
<figureCitation id="78952A7BF633C515FECB65867411CEE9" box="[304,376,1721,1748]" captionStart="FIGURE 9" captionStartId="12.[151,250,1166,1191]" captionTargetBox="[151,1436,195,1088]" captionTargetId="figure@12.[151,1436,195,1088]" captionTargetPageId="12" captionText="FIGURE 9. Radula of Pseudocleopatra togoensis Thiele, 1928, Ghana, NE region, DBL 214. A. Radula ribbon; scale bar: 20 μm. B. Lateral and central teeth; scale bar: 10 μm. C. Cutting edge of lateral and central teeth; scale bar: 2 μm. D. Central, lateral and marginal teeth; scale bar: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3460150" httpUri="https://zenodo.org/record/3460150/files/figure.png" pageId="10" pageNumber="311">Fig. 9</figureCitation>
|
||
,
|
||
<tableCitation id="AD2C0345F633C515FE7C6584748BCEE9" box="[391,482,1721,1749]" captionStart="TABLE 2" captionStartId="17.[151,239,694,718]" captionTargetPageId="17" captionText="TABLE 2. Number of denticles on the different radula teeth of examined samples of Pseudocleopatra Thiele, 1928 and Cleopatra Troschel, 1857 species. Data for Cleopatra bulimoides (Olivier, 1804) from Glaubrecht (1996) and data for P." httpUri="http://table.plazi.org/id/B4D16676F628C50EFF6C618977F2CB2A" pageId="10" pageNumber="311" tableUuid="B4D16676F628C50EFF6C618977F2CB2A">Table 2</tableCitation>
|
||
).
|
||
</emphasis>
|
||
The rachidian tooth is of squarish shape; its cutting edge is tapering to a slightly V-shaped tip bearing 5–6/1/5–6 denticles. The mesocone is quite prominent, being ca. ⅓ longer as well as two to three times wider than the lateral denticles. Its length may vary slightly, however, due to abrasion. The lateral teeth have relatively short but straight lateral extensions. The cutting edge bears 2–4/1/4–7 denticles with the mesocone being the largest and widest. It is two times wider and 1/3 longer than the flanking denticles and of rounded shape. Towards the outside of the lateral teeth the lengths of flanking denticles are decreasing. Inner and outer marginal teeth subequal in size and slightly curved. Both the inner and outer marginals bear 6–7 denticles each that are of approximately the same size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="B4D16676F632C514FF6C60A1718CCC1F" ID-DOI="http://doi.org/10.5281/zenodo.3460148" ID-Zenodo-Dep="3460148" httpUri="https://zenodo.org/record/3460148/files/figure.png" pageId="11" pageNumber="312" startId="11.[151,250,926,951]" targetBox="[481,1112,201,877]" targetPageId="11">
|
||
<paragraph id="E01136FEF632C514FF6C60A1718CCC1F" blockId="11.[151,1436,926,1059]" pageId="11" pageNumber="312">
|
||
<emphasis id="D2DAEAECF632C514FF6C60A1747ACB8A" bold="true" box="[151,275,926,951]" pageId="11" pageNumber="312">FIGURE 8.</emphasis>
|
||
Opercula of
|
||
<taxonomicName id="27AE4D7DF632C514FE6060A177A7CB8B" authority="Thiele, 1928" authorityName="Thiele" authorityYear="1928" box="[411,718,926,951]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="genus">
|
||
<emphasis id="D2DAEAECF632C514FE6060A1772ECB8B" box="[411,583,926,950]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra</emphasis>
|
||
<bibRefCitation id="843F4B0FF632C514FDB660A177A7CB8B" author="Thiele, J." box="[589,718,926,951]" pageId="11" pageNumber="312" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
species.
|
||
<emphasis id="D2DAEAECF632C514FCD060A1762DCB8A" bold="true" box="[811,836,926,951]" pageId="11" pageNumber="312">A.</emphasis>
|
||
Lectotype of
|
||
<taxonomicName id="27AE4D7DF632C514FC2860A071BFCB8B" authority="Thiele, 1928" authorityName="Thiele" authorityYear="1928" box="[979,1238,926,951]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="togoensis">
|
||
<emphasis id="D2DAEAECF632C514FC2860A07124CB8B" box="[979,1101,927,950]" italics="true" pageId="11" pageNumber="312">P. togoensis</emphasis>
|
||
<bibRefCitation id="843F4B0FF632C514FBA860A171BFCB8B" author="Thiele, J." box="[1107,1238,926,951]" pageId="11" pageNumber="312" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
, ZMB 190048b.
|
||
<emphasis id="D2DAEAECF632C514FA7E60A170F5CB8B" bold="true" box="[1413,1436,926,950]" pageId="11" pageNumber="312">B.</emphasis>
|
||
Paratype of
|
||
<taxonomicName id="27AE4D7DF632C514FEE860FD779ECBE7" authority="Mandahl-Barth, 1973" authorityName="Mandahl-Barth" authorityYear="1973" box="[275,759,962,987]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="voltana">
|
||
<emphasis id="D2DAEAECF632C514FEE860FD777BCBE7" box="[275,530,962,986]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra voltana</emphasis>
|
||
<bibRefCitation id="843F4B0FF632C514FDE160FD779ECBE7" author="Mandahl-Barth, G." box="[538,759,962,987]" pageId="11" pageNumber="312" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
|
||
</taxonomicName>
|
||
, Mogeyenga village, DBL 2591.
|
||
<emphasis id="D2DAEAECF632C514FBB660FD710FCBE6" bold="true" box="[1101,1126,962,987]" pageId="11" pageNumber="312">C.</emphasis>
|
||
Probable paratype of
|
||
<taxonomicName id="27AE4D7DF632C514FAB260FD773BCBC3" authority="Mandahl-Barth, 1973" authorityName="Mandahl-Barth" authorityYear="1973" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="dartevellei">
|
||
<emphasis id="D2DAEAECF632C514FAB260FD7405CBC3" italics="true" pageId="11" pageNumber="312">Pseudocleopatra dartevellei</emphasis>
|
||
<bibRefCitation id="843F4B0FF632C514FE8F60D9773BCBC3" author="Mandahl-Barth, G." box="[372,594,998,1023]" pageId="11" pageNumber="312" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
|
||
</taxonomicName>
|
||
, Matadi, DBL 2583.
|
||
<emphasis id="D2DAEAECF632C514FCD560D9762ECBC3" bold="true" box="[814,839,998,1022]" pageId="11" pageNumber="312">D.</emphasis>
|
||
<taxonomicName id="27AE4D7DF632C514FCB460D9719FCBC3" authority="(Putzeys, 1899)" authorityName="Neiber & Kahl & Wiggering & Glaubrecht" authorityYear="2019" baseAuthorityName="Putzeys" baseAuthorityYear="1899" box="[847,1270,998,1022]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="broecki" status="comb. nov.">
|
||
<emphasis id="D2DAEAECF632C514FCB460D97124CBC3" box="[847,1101,998,1022]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra broecki</emphasis>
|
||
(
|
||
<bibRefCitation id="843F4B0FF632C514FBA760D97186CBC3" author="Putzeys, S." box="[1116,1263,998,1022]" pageId="11" pageNumber="312" pagination="55 - 60" refId="ref16286" refString="Putzeys, S. (1899) Diagnoses de quelques coquilles et d'un sous-genre nouveau provenant de l'Etat independant du Congo. Annales de la Societe Royale malacologique de Belgique. Bulletins des Seances, 34, 55 - 60." type="journal article" year="1899">Putzeys, 1899</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C9E95797F632C514FB0560D87033CBC2" box="[1278,1370,998,1023]" pageId="11" pageNumber="312" rank="species">n. comb.</taxonomicNameLabel>
|
||
, MCZ 171520.
|
||
<emphasis id="D2DAEAECF632C514FF0B6735746ECC1F" bold="true" box="[240,263,1034,1058]" pageId="11" pageNumber="312">E.</emphasis>
|
||
Lectotype of
|
||
<taxonomicName id="27AE4D7DF632C514FE6F6735765ECC1F" authority="(Putzeys, 1899)" authorityName="Neiber & Kahl & Wiggering & Glaubrecht" authorityYear="2019" baseAuthorityName="Putzeys" baseAuthorityYear="1899" box="[404,823,1034,1058]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="broecki" status="comb. nov.">
|
||
<emphasis id="D2DAEAECF632C514FE6F673577F8CC1F" box="[404,657,1034,1058]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra broecki</emphasis>
|
||
(
|
||
<bibRefCitation id="843F4B0FF632C514FD6467357659CC1F" author="Putzeys, S." box="[671,816,1034,1058]" pageId="11" pageNumber="312" pagination="55 - 60" refId="ref16286" refString="Putzeys, S. (1899) Diagnoses de quelques coquilles et d'un sous-genre nouveau provenant de l'Etat independant du Congo. Annales de la Societe Royale malacologique de Belgique. Bulletins des Seances, 34, 55 - 60." type="journal article" year="1899">Putzeys, 1899</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C9E95797F632C514FCC6673476F1CC1E" box="[829,920,1034,1059]" pageId="11" pageNumber="312" rank="species">n. comb.</taxonomicNameLabel>
|
||
, MRAC 47360. Scale bar: 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A8B46575F632C514FF3C674B74E9CCC1" pageId="11" pageNumber="312" type="distribution">
|
||
<paragraph id="E01136FEF632C514FF3C674B74E9CCC1" blockId="11.[151,1437,1140,1707]" pageId="11" pageNumber="312">
|
||
<emphasis id="D2DAEAECF632C514FF3C674B74A7CCB2" bold="true" box="[199,462,1140,1167]" pageId="11" pageNumber="312">
|
||
Distribution (
|
||
<figureCitation id="78952A7BF632C514FE91674B74A9CCB2" box="[362,448,1140,1167]" captionStart="FIGURE 10" captionStartId="12.[151,250,1914,1939]" captionTargetBox="[485,1102,1278,1895]" captionTargetId="figure@12.[485,1102,1278,1895]" captionTargetPageId="12" captionText="FIGURE 10. Known occurrence records of Pseudocleopatra togoensis Thiele, 1928 in the Volta River system. The asterisk indicates the type locality. The arrow indicates the approximate location of the Akosombo dam." figureDoi="http://doi.org/10.5281/zenodo.3460152" httpUri="https://zenodo.org/record/3460152/files/figure.png" pageId="11" pageNumber="312">Fig. 10</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<taxonomicName id="27AE4D7DF632C514FE2D674A766ECCB3" authorityName="Thiele" authorityYear="1928" box="[470,775,1141,1167]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="togoensis">
|
||
<emphasis id="D2DAEAECF632C514FE2D674A766ECCB3" box="[470,775,1141,1167]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra togoensis</emphasis>
|
||
</taxonomicName>
|
||
is restricted to the
|
||
<collectingRegion id="226AF81CF632C514FC18674A7177CCB2" box="[995,1054,1141,1167]" country="Ghana" name="Volta" pageId="11" pageNumber="312">Volta</collectingRegion>
|
||
River system in
|
||
<collectingCountry id="98B9766EF632C514FB1A674A7043CCB2" box="[1249,1322,1141,1167]" name="Ghana" pageId="11" pageNumber="312">Ghana</collectingCountry>
|
||
and
|
||
<collectingCountry id="98B9766EF632C514FA98674A70F5CCB2" box="[1379,1436,1141,1167]" name="Togo" pageId="11" pageNumber="312">Togo</collectingCountry>
|
||
where it is reported from widely separated localities, some of which are located in the area today inundated by the artificial Lake Volta, as a consequence of the completion of the Akosombo Dam in 1965 (see
|
||
<bibRefCitation id="843F4B0FF632C514FB4467827036CCEA" author="Gocking, R. S." box="[1215,1375,1212,1239]" pageId="11" pageNumber="312" refId="ref14713" refString="Gocking, R. S. (2005) The History of Ghana. Greenwood Press, Westport, London, xxxiii + 331 pp., 11 pls." type="book" year="2005">Gocking 2005</bibRefCitation>
|
||
for a historical overview).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8B46575F632C514FF3C663A7031CE96" pageId="11" pageNumber="312" type="discussion">
|
||
<paragraph id="E01136FEF632C514FF3C663A7629CE02" blockId="11.[151,1437,1140,1707]" pageId="11" pageNumber="312">
|
||
<emphasis id="D2DAEAECF632C514FF3C663A7452CD22" bold="true" box="[199,315,1285,1311]" pageId="11" pageNumber="312">Remarks.</emphasis>
|
||
<taxonomicName id="27AE4D7DF632C514FEB9663A7719CD23" authorityName="Thiele" authorityYear="1928" box="[322,624,1285,1311]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="togoensis">
|
||
<emphasis id="D2DAEAECF632C514FEB9663A7719CD23" box="[322,624,1285,1311]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra togoensis</emphasis>
|
||
</taxonomicName>
|
||
is the type species of
|
||
<taxonomicName id="27AE4D7DF632C514FC9F663A714BCD22" authorityName="Thiele" authorityYear="1928" box="[868,1058,1285,1311]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="genus">
|
||
<emphasis id="D2DAEAECF632C514FC9F663A714BCD22" box="[868,1058,1285,1311]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra</emphasis>
|
||
</taxonomicName>
|
||
. In his original description
|
||
<bibRefCitation id="843F4B0FF632C514FAAE663A7472CD7E" author="Thiele, J." pageId="11" pageNumber="312" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele (1928: 394)</bibRefCitation>
|
||
described two different shells from the
|
||
<collectingRegion id="226AF81CF632C514FD276616767ECD7E" box="[732,791,1321,1347]" country="Ghana" name="Volta" pageId="11" pageNumber="312">Volta</collectingRegion>
|
||
River near the village Apase that were collected by Willi Kört. Only the specimen designated as
|
||
<typeStatus id="3F15885CF632C514FD98667277A3CD5A" box="[611,714,1357,1383]" pageId="11" pageNumber="312" type="lectotype">lectotype</typeStatus>
|
||
above represents the species; the other former
|
||
<typeStatus id="3F15885CF632C514FB0E66727025CD5A" box="[1269,1356,1357,1383]" pageId="11" pageNumber="312" type="syntype">syntype</typeStatus>
|
||
(ZMB 190049) actually belongs to
|
||
<taxonomicName id="27AE4D7DF632C514FE1B664E773BCDB6" authorityName="Mandahl-Barth" authorityYear="1973" box="[480,594,1393,1419]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="voltana">
|
||
<emphasis id="D2DAEAECF632C514FE1B664E773BCDB6" box="[480,594,1393,1419]" italics="true" pageId="11" pageNumber="312">P. voltana</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<taxonomicName id="27AE4D7DF632C514FD6C664E760FCDB6" box="[663,870,1393,1419]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="11" pageNumber="304" phylum="Bryophyta" rank="species" species="preparation">radula preparation</taxonomicName>
|
||
of the
|
||
<typeStatus id="3F15885CF632C514FC47664E714ACDB6" box="[956,1059,1393,1419]" pageId="11" pageNumber="312" type="lectotype">lectotype</typeStatus>
|
||
(ZMB 190048b) by Thiele from 1928 was also examined. However, a radula ribbon could not be detected. It may have dissolved during its long storage time. The absence of denticles on the marginal teeth—as noted by
|
||
<bibRefCitation id="843F4B0FF632C514FC2066867119CDE9" author="Thiele, J." box="[987,1136,1465,1492]" pageId="11" pageNumber="312" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele (1928)</bibRefCitation>
|
||
—could not be confirmed. However, eight examined specimens of
|
||
<taxonomicName id="27AE4D7DF632C514FDA766E2779DCDCA" authorityName="Mandahl-Barth" authorityYear="1973" box="[604,756,1501,1527]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="dartevellei">
|
||
<emphasis id="D2DAEAECF632C514FDA766E27704CDCB" box="[604,621,1501,1526]" italics="true" pageId="11" pageNumber="312">P</emphasis>
|
||
.
|
||
<emphasis id="D2DAEAECF632C514FD8766E2779DCDCA" box="[636,756,1501,1527]" italics="true" pageId="11" pageNumber="312">dartevellei</emphasis>
|
||
</taxonomicName>
|
||
also lacked denticles on the inner marginal teeth, while two other specimens possessed up to 18 denticles on both marginal teeth (
|
||
<tableCitation id="AD2C0345F632C514FC5D653E7693CE26" box="[934,1018,1537,1563]" captionStart="TABLE 1" captionStartId="13.[151,239,154,178]" captionTargetPageId="13" captionText="TABLE 1. Pairwise correct classifications based on linear discriminant analyses of h/w and h/la ratios of Pseudocleopatra species. Values for species from the same river system, i.e. Volta River or Congo River, in bold. For abbreviations of measured shell variables, see Fig. 2." httpUri="http://table.plazi.org/id/B4D16676F634C512FF6C63A57771C8C7" pageId="11" pageNumber="312" tableUuid="B4D16676F634C512FF6C63A57771C8C7">Table 1</tableCitation>
|
||
). Thus, it may be possible that
|
||
<taxonomicName id="27AE4D7DF632C514FAA5653E7582CE03" authorityName="Thiele" authorityYear="1928" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="togoensis">
|
||
<emphasis id="D2DAEAECF632C514FAA5653E7006CE27" box="[1374,1391,1537,1562]" italics="true" pageId="11" pageNumber="312">P</emphasis>
|
||
.
|
||
<emphasis id="D2DAEAECF632C514FA86653E7582CE03" italics="true" pageId="11" pageNumber="312">togoensis</emphasis>
|
||
</taxonomicName>
|
||
is also polymorphic with regard to radula characters.
|
||
</paragraph>
|
||
<paragraph id="E01136FEF632C514FF3C65767031CE96" blockId="11.[151,1437,1140,1707]" pageId="11" pageNumber="312">
|
||
<taxonomicName id="27AE4D7DF632C514FF3C65767490CE5F" authorityName="Thiele" authorityYear="1928" box="[199,505,1609,1635]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="togoensis">
|
||
<emphasis id="D2DAEAECF632C514FF3C65767490CE5F" box="[199,505,1609,1635]" italics="true" pageId="11" pageNumber="312">Pseudocleopatra togoensis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="27AE4D7DF632C514FCC3657676C3CE5E" authorityName="Mandahl-Barth" authorityYear="1973" box="[824,938,1609,1635]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="11" pageNumber="312" phylum="Mollusca" rank="species" species="voltana">
|
||
<emphasis id="D2DAEAECF632C514FCC3657676C3CE5E" box="[824,938,1609,1635]" italics="true" pageId="11" pageNumber="312">P. voltana</emphasis>
|
||
</taxonomicName>
|
||
by the less elongate shape and the usually smoother shell without distinct spiral sculpture. Sinuous darker radial bands were only observed in some specimens of this species. Nothing is known about the ecology of the species and new records have not become known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |