treatments-xml/data/68/03/E4/6803E4A591CF5A698FD4306B53CF8AA9.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1056.68722" ID-GBIF-Dataset="b4bae190-3a51-417c-bc86-1d036552cb02" ID-PMC="PMC8387304" ID-Pensoft-Pub="1313-2970-1056-59" ID-Pensoft-UUID="2D2F52E3B5EF5014BE22CCFDF71E1961" ID-PubMed="34512091" ID-ZooBank="A506736774764471B083284562438CB8" ModsDocID="1313-2970-1056-59" checkinTime="1629388992862" checkinUser="pensoft" docAuthor="Aswaj, Punnath, Sahanashree, Ramakrishnaiah, Udayakantha, Warnakulasuriyage Sudesh, Aniruddha, Marathe &amp; Priyadarsanan, Dharma Rajan" docDate="2021" docId="6803E4A591CF5A698FD4306B53CF8AA9" docLanguage="en" docName="ZooKeys 1056: 59-72" docOrigin="ZooKeys 1056" docPubDate="2021-08-18" docSource="http://dx.doi.org/10.3897/zookeys.1056.68722" docTitle="Syscia indica Aswaj &amp; Sahanashree &amp; Udayakantha &amp; Aniruddha &amp; Priyadarsanan 2021, sp. nov." docType="treatment" docUuid="3A9C67DF-8692-40EF-A3FD-3102D10C1253" docUuidSource="ZooBank" docVersion="4" id="2D2F52E3B5EF5014BE22CCFDF71E1961" lastPageNumber="59" masterDocId="2D2F52E3B5EF5014BE22CCFDF71E1961" masterDocTitle="Two new species of doryline ants (Hymenoptera, Formicidae) with 11 - segmented antennae from India" masterLastPageNumber="72" masterPageNumber="59" pageNumber="59" updateTime="1668150855576" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Two new species of doryline ants (Hymenoptera, Formicidae) with 11 - segmented antennae from India</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aswaj, Punnath</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8034-6578</mods:nameIdentifier>
<mods:affiliation>Ashoka Trust for Research in Ecology and the Environment, Royal Enclave, Srirampura, Jakkur Post, Bengaluru - 560064, Karnataka, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sahanashree, Ramakrishnaiah</mods:namePart>
<mods:affiliation>Ashoka Trust for Research in Ecology and the Environment, Royal Enclave, Srirampura, Jakkur Post, Bengaluru - 560064, Karnataka, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Udayakantha, Warnakulasuriyage Sudesh</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6766-5504</mods:nameIdentifier>
<mods:affiliation>Department of Zoology and Environmental Management, University of Kelaniya, Sri Lanka</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aniruddha, Marathe</mods:namePart>
<mods:affiliation>Centre for Ecological Sciences, Indian Institute of Science, Bengaluru - 560012, Karnataka, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Priyadarsanan, Dharma Rajan</mods:namePart>
<mods:affiliation>Ashoka Trust for Research in Ecology and the Environment, Royal Enclave, Srirampura, Jakkur Post, Bengaluru - 560064, Karnataka, India</mods:affiliation>
<mods:nameIdentifier type="email">priyan@atree.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1056</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>59</mods:start>
<mods:end>72</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1056.68722</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1056.68722</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1056-59</mods:identifier>
<mods:identifier type="ZooBank">A506736774764471B083284562438CB8</mods:identifier>
<mods:identifier type="Pensoft-UUID">2D2F52E3B5EF5014BE22CCFDF71E1961</mods:identifier>
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<treatment ID-GBIF-Taxon="184603608" LSID="urn:lsid:zoobank.org:act:3A9C67DF-8692-40EF-A3FD-3102D10C1253" httpUri="http://treatment.plazi.org/id/6803E4A591CF5A698FD4306B53CF8AA9" lastPageNumber="59" pageId="0" pageNumber="59">
<subSubSection pageId="0" pageNumber="59" type="nomenclature">
<paragraph pageId="0" pageNumber="59">
<taxonomicName LSID="http://zoobank.org/3A9C67DF-8692-40EF-A3FD-3102D10C1253" authority="Aswaj &amp; Sahanashree &amp; Udayakantha &amp; Aniruddha &amp; Priyadarsanan, 2021" authorityName="Aswaj &amp; Sahanashree &amp; Udayakantha &amp; Aniruddha &amp; Priyadarsanan" authorityYear="2021" class="Insecta" family="Formicidae" genus="Syscia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Syscia indica" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">Syscia indica</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="59">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="description">
<paragraph pageId="0" pageNumber="59">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">Fig. 3A-F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="materials_examined">
<paragraph pageId="0" pageNumber="59">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="59">
<materialsCitation collectingDate="2013-04-10" collectorName="Aniruddha Marathe" country="India" county="One" elevation="1600" latitude="27.0434" location="Original" longLatPrecision="6" longitude="92.4302" municipality="Point" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="59">
<typeStatus>Holotype</typeStatus>
</emphasis>
<collectingCounty>One</collectingCounty>
worker, 
<collectingMunicipality>Point</collectingMunicipality>
mounted. 
<location LSID="urn:lsid:plazi:treatment:6803E4A591CF5A698FD4306B53CF8AA9:BB3D016C5C40B37581E207FF299837FF" country="India" county="One" latitude="27.0434" longLatPrecision="6" longitude="92.4302" municipality="Point" name="Original" stateProvince="Arunachal Pradesh">Original</location>
label: &quot; 
<collectingCountry name="India">India</collectingCountry>
, 
<collectingRegion country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
, West Kameng, Eaglenest WLS, 
<geoCoordinate degrees="27.0434" direction="north" orientation="latitude" precision="5" value="27.0434">27.0434°N</geoCoordinate>
, 
<geoCoordinate degrees="92.4302" direction="east" orientation="longitude" precision="5" value="92.4302">92.4302°E</geoCoordinate>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</elevation>
</quantity>
, Winkler extraction method, 
<collectingDate value="2013-04-10">10.IV.2013</collectingDate>
, 
<collectorName>Aniruddha Marathe</collectorName>
leg.&quot; [NBAIR/HYM-FOR/1721-2].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="description">
<paragraph pageId="0" pageNumber="59">Worker description.</paragraph>
<paragraph pageId="0" pageNumber="59">Measurements and indices (holotype): BL 3.33; HL 0.68; HW 0.50; SL 0.25; PRW 0.35; WL 0.85; PL 0.36, PH 0.31, PW 0.29, PPL 0.41, PPH 0.47, PPW 0.43. Indices: CI 74, SI 50, PI1 116, PI2 81, PPI1 87, PPI2 105, WI 148 (all measurements in mm).</paragraph>
<paragraph pageId="0" pageNumber="59">
<emphasis bold="true" pageId="0" pageNumber="59">
<emphasis italics="true" pageId="0" pageNumber="59">Head</emphasis>
.
</emphasis>
In full-face view, rectangular, distinctly longer than broad; lateral margin weakly convex; posterior margin weakly concave (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3A</figureCitation>
). Antennae 11-segmented, apical segment (XI) longer than IX-X combined; scape short, reaching mid-length of head when folded back. Antennal segment II almost as long as broad, segments III-X slightly broader than long (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3F</figureCitation>
). Frontal carinae short and narrow, reaching less than half of head length. Eyes and ocelli completely absent (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3A</figureCitation>
). Mandibles triangular; masticatory margin serrated, without distinct teeth (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3F</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="59">
<emphasis italics="true" pageId="0" pageNumber="59">Mesosoma</emphasis>
.
</emphasis>
In dorsal view, elongate with almost parallel lateral sides (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3C</figureCitation>
). In profile view, dorsal outline weakly convex; promesonotal suture and metanotal groove absent; mesopleuron demarcated from propleuron by a distinct pronotomesopleural suture; mesopleuron not clearly differentiated from metapleuron. Propodeal declivity feebly concave, encircled with a distinct thin rim (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3B</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="59">
<emphasis italics="true" pageId="0" pageNumber="59">Metasoma</emphasis>
.
</emphasis>
In dorsal view, petiole distinctly longer than broad; almost parallel sided (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3D</figureCitation>
). In profile view, petiole subrectangular; slightly longer than high with weakly convex dorsal outline. Subpetiolar process subrectangular; ventral outline strongly concave (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3B</figureCitation>
). Postpetiole in dorsal view, larger than petiole; slightly longer than broad (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3D</figureCitation>
). In profile view, postpetiole slightly shorter than high; dorsal outline weakly convex. Postpetiolar sternite in profile view low, ventral margin weakly convex; anteroventrally produced into a blunt angle (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3B</figureCitation>
). Abdominal tergite IV (first gastral tergite) in dorsal view, elongate with anterior margin weakly concave; lateral sides weakly convex (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3E</figureCitation>
). Base of cinctus of first gastral tergite with cross ribs (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Syscia indica sp. nov., holotype worker A head in full-face view B body in profile view C mesosoma in dorsal view D petiole and postpetiole in dorsal view E girdling constriction and gaster in dorsal view F antenna and Mandibles." figureDoi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59">3E</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="59">
<emphasis italics="true" pageId="0" pageNumber="59">Sculpture</emphasis>
.
</emphasis>
Body with numerous, relatively small, closely-spaced foveolae. Ventrolateral surface of petiole with two distinct almost parallel carinae just above the subpetiolar process. Antennal scape, outer surface of mandible and legs with fine dense micropunctures. 
<emphasis bold="true" pageId="0" pageNumber="59">
<emphasis italics="true" pageId="0" pageNumber="59">Pilosity</emphasis>
.
</emphasis>
Body covered with erect and sub-erect hairs. Dorsum of postpetiole and first gastral tergite with sparse erect hairs mixed with dense long decumbent hairs. Antennae and legs with dense short decumbent hairs. 
<emphasis italics="true" pageId="0" pageNumber="59">Body coloration</emphasis>
. Body reddish brown. Antennae and legs yellowish brown.
</paragraph>
<caption doi="10.3897/zookeys.1056.68722.figure3" httpUri="https://binary.pensoft.net/fig/578940" pageId="0" pageNumber="59" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="59">
<emphasis bold="true" pageId="0" pageNumber="59">Figure 3.</emphasis>
<taxonomicName authorityName="Aswaj &amp; Sahanashree &amp; Udayakantha &amp; Aniruddha &amp; Priyadarsanan" authorityYear="2021" class="Insecta" family="Formicidae" genus="Syscia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Syscia indica" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">Syscia indica</emphasis>
</taxonomicName>
sp. nov., holotype worker 
<emphasis bold="true" pageId="0" pageNumber="59">A</emphasis>
head in full-face view 
<emphasis bold="true" pageId="0" pageNumber="59">B</emphasis>
body in profile view 
<emphasis bold="true" pageId="0" pageNumber="59">C</emphasis>
mesosoma in dorsal view 
<emphasis bold="true" pageId="0" pageNumber="59">D</emphasis>
petiole and postpetiole in dorsal view 
<emphasis bold="true" pageId="0" pageNumber="59">E</emphasis>
girdling constriction and gaster in dorsal view 
<emphasis bold="true" pageId="0" pageNumber="59">F</emphasis>
antenna and Mandibles.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="recognition">
<paragraph pageId="0" pageNumber="59">Recognition.</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Aswaj &amp; Sahanashree &amp; Udayakantha &amp; Aniruddha &amp; Priyadarsanan" authorityYear="2021" class="Insecta" family="Formicidae" genus="Syscia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Syscia indica" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">Syscia indica</emphasis>
</taxonomicName>
sp. nov. is similar to 
<taxonomicName lsidName="S. chaladthanyakiji" pageId="0" pageNumber="59" rank="species" species="chaladthanyakiji">
<emphasis italics="true" pageId="0" pageNumber="59">S. chaladthanyakiji</emphasis>
</taxonomicName>
in general appearance and 11-segmented antennae. However, 
<taxonomicName lsidName="S. indica" pageId="0" pageNumber="59" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">S. indica</emphasis>
</taxonomicName>
can be separated from 
<taxonomicName lsidName="S. chaladthanyakiji" pageId="0" pageNumber="59" rank="species" species="chaladthanyakiji">
<emphasis italics="true" pageId="0" pageNumber="59">S. chaladthanyakiji</emphasis>
</taxonomicName>
by the following characteristics: 1) weak sculpture with relatively small and closely-spaced foveolae in 
<taxonomicName lsidName="S. indica" pageId="0" pageNumber="59" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">S. indica</emphasis>
</taxonomicName>
(stronger sculpturing with well-defined macropunctures in 
<taxonomicName lsidName="S. chaladthanyakiji" pageId="0" pageNumber="59" rank="species" species="chaladthanyakiji">
<emphasis italics="true" pageId="0" pageNumber="59">S. chaladthanyakiji</emphasis>
</taxonomicName>
); 2) abdominal segment IV (first gastral tergite) with large number of closely-spaced foveolae in 
<taxonomicName lsidName="S. indica" pageId="0" pageNumber="59" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">S. indica</emphasis>
</taxonomicName>
(first gastral tergite with lesser number of relatively widely spaced foveolae in 
<taxonomicName lsidName="S. chaladthanyakiji" pageId="0" pageNumber="59" rank="species" species="chaladthanyakiji">
<emphasis italics="true" pageId="0" pageNumber="59">S. chaladthanyakiji</emphasis>
</taxonomicName>
); 3) remaining gastral segments with feeble foveolae in 
<taxonomicName lsidName="S. indica" pageId="0" pageNumber="59" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">S. indica</emphasis>
</taxonomicName>
(remaining gastral segments anteriorly with an unscupltured area in 
<emphasis italics="true" pageId="0" pageNumber="59">
<taxonomicName lsidName="S. chaladthanyakiji" pageId="0" pageNumber="59" rank="species" species="chaladthanyakiji">S. chaladthanyakiji</taxonomicName>
)
</emphasis>
; 4) head in full-face view oval shaped with convex lateral sides and moderately concave posterior margin in 
<taxonomicName lsidName="S. indica" pageId="0" pageNumber="59" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">S. indica</emphasis>
</taxonomicName>
(head in full-face view rectangular, almost parallel sides, posterior margin weakly concave in 
<taxonomicName lsidName="S. chaladthanyakiji" pageId="0" pageNumber="59" rank="species" species="chaladthanyakiji">
<emphasis italics="true" pageId="0" pageNumber="59">S. chaladthanyakiji</emphasis>
</taxonomicName>
); 5) CI 74, SI 50 in 
<taxonomicName lsidName="S. indica" pageId="0" pageNumber="59" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">S. indica</emphasis>
</taxonomicName>
(CI 80-84, SI 56-63 in 
<emphasis italics="true" pageId="0" pageNumber="59">
<taxonomicName lsidName="S. chaladthanyakiji" pageId="0" pageNumber="59" rank="species" species="chaladthanyakiji">S. chaladthanyakiji</taxonomicName>
)
</emphasis>
. 
<taxonomicName authorityName="Aswaj &amp; Sahanashree &amp; Udayakantha &amp; Aniruddha &amp; Priyadarsanan" authorityYear="2021" class="Insecta" family="Formicidae" genus="Syscia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Syscia indica" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">Syscia indica</emphasis>
</taxonomicName>
sp. nov. is also similar to the Sri Lankan and Chinese 
<taxonomicName lsidName="S. typhla" pageId="0" pageNumber="59" rank="species" species="typhla">
<emphasis italics="true" pageId="0" pageNumber="59">S. typhla</emphasis>
</taxonomicName>
in general appearance. However, the new species can be easily separated from 
<taxonomicName lsidName="S. typhla" pageId="0" pageNumber="59" rank="species" species="typhla">
<emphasis italics="true" pageId="0" pageNumber="59">S. typhla</emphasis>
</taxonomicName>
by the 1) presence of 11-segmented antennae (9-segmented in 
<taxonomicName lsidName="S. typhla" pageId="0" pageNumber="59" rank="species" species="typhla">
<emphasis italics="true" pageId="0" pageNumber="59">S. typhla</emphasis>
</taxonomicName>
); 2) foveolae on cephalic dorsum smaller in diameter and closely spaced in 
<taxonomicName lsidName="S. indica" pageId="0" pageNumber="59" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">S. indica</emphasis>
</taxonomicName>
(foveolae comparatively larger in diameter and widely spaced in 
<taxonomicName lsidName="S. typhla" pageId="0" pageNumber="59" rank="species" species="typhla">
<emphasis italics="true" pageId="0" pageNumber="59">S. typhla</emphasis>
</taxonomicName>
); 3) posterior cephalic margin nearly transverse in 
<taxonomicName lsidName="S. Indica" pageId="0" pageNumber="59" rank="species" species="Indica">
<emphasis italics="true" pageId="0" pageNumber="59">S. Indica</emphasis>
</taxonomicName>
(posterior cephalic margin strongly concave in 
<taxonomicName lsidName="S. typhla" pageId="0" pageNumber="59" rank="species" species="typhla">
<emphasis italics="true" pageId="0" pageNumber="59">S. typhla</emphasis>
</taxonomicName>
); 4) base of cinctus of first gastral tergite with cross ribs in 
<taxonomicName lsidName="S. indica" pageId="0" pageNumber="59" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="59">S. indica</emphasis>
</taxonomicName>
(base of cinctus of first gastral tergite without cross ribs in 
<taxonomicName lsidName="S. typhla" pageId="0" pageNumber="59" rank="species" species="typhla">
<emphasis italics="true" pageId="0" pageNumber="59">S. typhla</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="habitat">
<paragraph pageId="0" pageNumber="59">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="59">
The holotype was collected from the Eaglenest Wildlife Sanctuary at an elevation of 1600 m above sea level. The region has a canopy cover of about 85% and no indications of anthropogenic disturbances. The soil had a pH of 6.85 and temperature 16°C at the time of collection. The specimen was collected using a Winkler extractor from sifted leaf litter of 1 m2. We were able to capture one additional ant genus 
<taxonomicName authorityName="Westwood" authorityYear="1840" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" lsidName="Carebara" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="59">Carebara</emphasis>
</taxonomicName>
in the same trap, which consisted of 295 ml leaf litter.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="distribution">
<paragraph pageId="0" pageNumber="59">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="59">Indomalaya: India (Arunachal Pradesh).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="etymology">
<paragraph pageId="0" pageNumber="59">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="59">
The specific epithet 
<taxonomicName authorityName="Aswaj &amp; Sahanashree &amp; Udayakantha &amp; Aniruddha &amp; Priyadarsanan" authorityYear="2021" class="Insecta" family="Formicidae" genus="Syscia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Syscia indica" order="Hymenoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="indica">Syscia indica</taxonomicName>
is a Latin singular feminine adjective in the nominative case and refers to the country where the species was collected.
</paragraph>
</subSubSection>
</treatment>
</document>