150 lines
19 KiB
XML
150 lines
19 KiB
XML
<document id="470A3A4B540E9BD6CC627AE2B01F25F0" ID-CLB-Dataset="35155" ID-DOI="http://dx.doi.org/10.3897/zookeys.335.5567" ID-GBIF-Dataset="f0c6f528-5c7c-4e61-87c3-dbf43b9350ef" ID-PMC="PMC3800789" ID-Pensoft-Pub="1313-2970-335-1" ID-PubMed="24146563" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-335-1" ModsDocOrigin="ZooKeys 335" ModsDocTitle="Shallow water marine gammaridean amphipods of Pulau Tioman, Malaysia, with the description of a new species" checkinTime="1451246890703" checkinUser="pensoft" docAuthor="Azman, B. A. R. & Othman, B. H. R." docDate="2013" docId="67A002947C6414D0C20E28F316D014ED" docLanguage="en" docName="ZooKeys 335: 1-31" docOrigin="ZooKeys 335" docSource="http://dx.doi.org/10.3897/zookeys.335.5567" docTitle="Ericthonius pugnax Dana 1853" docType="treatment" docVersion="5" lastPageNumber="9" masterDocId="6B111C28A76E1D660F77FF9FFFCFFFCB" masterDocTitle="Shallow water marine gammaridean amphipods of Pulau Tioman, Malaysia, with the description of a new species" masterLastPageNumber="31" masterPageNumber="1" pageNumber="9" updateTime="1732742891443" updateUser="ExternalLinkService">
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<mods:mods id="9AECB4AB2F28A4B2CC0B414C17E7E68F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1E7AD96CF797A2D2381008A821D45C6C">Shallow water marine gammaridean amphipods of Pulau Tioman, Malaysia, with the description of a new species</mods:title>
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<mods:namePart id="878B1C872541EF9D3C08F7903B6DCF46">Azman, B. A. R.</mods:namePart>
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<mods:namePart id="6D80F1159CEC4A78D937D72BA4A4D894">Othman, B. H. R.</mods:namePart>
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<mods:typeOfResource id="FA4FD7EC46F81F95C85A552224802140">text</mods:typeOfResource>
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<mods:date id="A634BD45A75AE7E39DBE300C07D10A7F">2013</mods:date>
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<mods:number id="D96290C229FF05F4880726135508C952">335</mods:number>
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<mods:start id="DB5FE9B9AFD12B9BB686D7F3C169252C">1</mods:start>
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<mods:classification id="FD5B3737917B47FC5EAA9E6612757FB0">journal article</mods:classification>
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<mods:identifier id="F6A556BE193503869D21F565651BD50F" type="DOI">http://dx.doi.org/10.3897/zookeys.335.5567</mods:identifier>
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<mods:identifier id="C1276CA7B3D7B221D611CDEAE049DF35" type="Pensoft-Pub">1313-2970-335-1</mods:identifier>
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</mods:mods>
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<treatment id="67A002947C6414D0C20E28F316D014ED" ID-GBIF-Taxon="152048371" LSID="urn:lsid:plazi:treatment:67A002947C6414D0C20E28F316D014ED" httpUri="http://treatment.plazi.org/id/67A002947C6414D0C20E28F316D014ED" lastPageNumber="9" pageId="8" pageNumber="9" scope_family="Ischyroceridae">
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<subSubSection id="8826EEC49DF3E65DA8B50740FE7AA6B0" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="871AEAA393DE6600F82767741B7EC79D" pageId="8" pageNumber="9">
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<taxonomicName id="95A2D95305215C7215AF9E539D2E48D9" ID-CoL="6GNZY" LSID="http://species-id.net/wiki/Ericthonius_pugnax" authority="Dana, 1853" authorityName="Dana" authorityYear="1853" class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax (Dana, 1853)</taxonomicName>
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Figure 6
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</paragraph>
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</subSubSection>
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<subSubSection id="25050A95FEC2353FC7EEC09901AE3F19" pageId="8" pageNumber="9" type="reference_group">
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<paragraph id="3D4B6D76EF16DD1AEFC12AC288FE71A0" pageId="8" pageNumber="9">
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<taxonomicName id="5F564714F9E299B8784E290DD3781F27" genus="Pyctilus" lsidName="Pyctilus macrodactylus" pageId="8" pageNumber="9" rank="species" species="macrodactylus">Pyctilus macrodactylus</taxonomicName>
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Synonymy: Dana, 1853: 974.
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</paragraph>
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<paragraph id="DC5C643AD30B22589B84472533649F16" pageId="8" pageNumber="9">
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<taxonomicName id="7934FB2CA7601459215307FE93A52497" class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius macrodactylus" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macrodactylus">Ericthonius macrodactylus</taxonomicName>
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Stebbing, 1906: 672;
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<bibRefCitation id="A063ADD4C119796008F012C58D111229" author="Walker, AO" journalOrPublisher="Report of the Government of Ceylon Pearl Oyster Fishery of the Gulf of Manaar" pageId="21" pageNumber="22" pagination="229 - 300" title="Report on Amphipoda collected by Professor Herdman, at Ceylon, in 1902." volume="17" year="1904">Walker 1904</bibRefCitation>
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: 292, fig. 48.
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</paragraph>
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<paragraph id="6FDAB0C38F4E6B9B5050CB30C94799C8" pageId="8" pageNumber="9">
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<taxonomicName id="F76CF407B55E850FCED9D21AF7ECF6A6" class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax</taxonomicName>
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Stebbing, 1906: 672;
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<bibRefCitation id="3CF56AD2075006FE58422B76DB67CD82" pageId="8" pageNumber="9">Pirlot 1938</bibRefCitation>
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: 352;
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<bibRefCitation id="DB37CEF59821E6659CA90698320A8083" author="Hurley, DE" journalOrPublisher="Transactions of the Royal Society of New Zealand" pageId="18" pageNumber="19" pagination="431 - 460" volume="82" year="1954">Hurley 1954</bibRefCitation>
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: 445, figs 40-61;
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<bibRefCitation id="506C7D383F0B2A827ADF65CE80B183B6" author="Nagata, K" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="19" pageNumber="20" pagination="163 - 182" title="Preliminary notes on benthic gammaridean Amphipoda from the Zostera region of Mihara Bay, Seto Inland Sea, Japan." volume="8" year="1960">Nagata 1960</bibRefCitation>
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: 179, pl. 179, figs 99-102;
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<bibRefCitation id="5186477738BE4A7EB9E3224C03D6FF18" author="Nagata, K" journalOrPublisher="Publication of Seto Marine Biological Laboratory" pageId="19" pageNumber="20" pagination="291 - 322" title="Studies on marine Gammaridean Amphipoda of the Seto Inland Sea III." volume="13" year="1965 b">Nagata 1965b</bibRefCitation>
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: 320, fig. 40;
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<bibRefCitation id="B72C599DC98C771E5E9A09754C194B35" pageId="8" pageNumber="9">Nayar 1967</bibRefCitation>
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:162;
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<bibRefCitation id="D9648E76058DFD8F12AF106F4B3B155C" author="Ledoyer, M" journalOrPublisher="Recueil des Travaux de la Station Marine Endoume, Fascicule hors serie Supplement" pageId="18" pageNumber="19" pagination="179 - 182" title="Amphipodes tubicoles des feuilles des herbiers de phanerogames marines de la region de Tulear (Madagascar)." volume="9" year="1969">Ledoyer 1969</bibRefCitation>
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: 179, fig. 1;
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<bibRefCitation id="6EF3D14623B2304151B1DB0AD68B3868" author="Hirayama, A" journalOrPublisher="Publication of Seto Marine Biological Laboratory" pageId="17" pageNumber="18" pagination="1 - 53" title="Taxonomic studies on the shallow water Gammaridean Amphipoda of west Kyushu, Japan. IV. Dexaminidae (Guernea), Eophiliantidae, Eusoridae, Haustoriidae, Hyalidae and Ischyroceridae." volume="30" year="1985">Hirayama 1985</bibRefCitation>
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: 52,
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<bibRefCitation id="4572694800D082DB51631306E3CE971C" author="Ledoyer, M" journalOrPublisher="Memorie del Museo Civico di Storia Naturale di Verona" pageId="18" pageNumber="19" pagination="45 - 84" title="Les Oedicerotidae (Crustacea Amphipoda) De La Mer Mediterranee." volume="9" year="1982 a">Ledoyer 1982a</bibRefCitation>
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: 628, fig. 239;
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<bibRefCitation id="67164BF0B79AC643DA374CA005918510" author="Moore, PG" journalOrPublisher="Journal of Natural History" pageId="19" pageNumber="20" pagination="705 - 732" title="Taxonomic observations on the genera Xenocheria Haswell and Ericthonius Milne-Edwards (Crustacea: Amphipoda) from Australian coastal waters." url="10.1080/00222938800770461" volume="22" year="1988">Moore 1988</bibRefCitation>
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: 727-730, fig. 14;
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<bibRefCitation id="832ED30A7298B1B3850F757BEBF98B38" author="Kim, W" journalOrPublisher="Korean J. Zool." pageId="18" pageNumber="19" pagination="232 - 252" title="The Marine Amphipod Crustaceans of Ulreung Island, Korea: Part 1." volume="34" year="1991">Kim and Kim 1991</bibRefCitation>
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: 246-247, figs 13-14.
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</paragraph>
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</subSubSection>
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<subSubSection id="AE5073ABDD0725E8131C0ADBEEFE94FA" pageId="8" pageNumber="9" type="material">
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<paragraph id="CF4052F6D7B5D769D5079433E723D2DA" pageId="8" pageNumber="9">Material.</paragraph>
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<paragraph id="4D88D5B456CD4CD556381F0BD999DFBC" pageId="8" pageNumber="9">
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2 specimens, TIO 5, Monkey Bay, Pulau Tioman, 2
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<geoCoordinate id="071F96BF90DCC48BF3A8E5EC7F6DA79F" direction="north" orientation="latitude" precision="15" value="2.8258333">2°49'33"N</geoCoordinate>
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,
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<geoCoordinate id="817B13CAF3539242C1A2BE9D1A3E919D" direction="east" orientation="longitude" precision="15" value="104.1625">104°9'45"E</geoCoordinate>
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, coral rubble, Azman, Josim, 22 August 1996; 86 specimens, TIO 15, Renggis, Pulau Tioman,
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<geoCoordinate id="06EEDC09A97C72A4FAF56662780646DD" direction="north" orientation="latitude" precision="15" value="2.8097222">2°48'35"N</geoCoordinate>
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,
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<geoCoordinate id="956F5887D59A51477A361EA8DDE14D44" direction="east" orientation="longitude" precision="15" value="104.134995">104°8'6"E</geoCoordinate>
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, seagrass, Azman, Rayida, 15 July 1999, UKM I.D. 5712-5719; 2 specimens, TIO 33, Tomok, Pulau Tioman,
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<geoCoordinate id="C171538839B486CE54B53B211BFC2867" direction="north" orientation="latitude" precision="15" value="2.7938888">2°47'38"N</geoCoordinate>
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,
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<geoCoordinate id="DB2AECB3CACE788AA5F987B0B0AC7D8E" direction="east" orientation="longitude" precision="15" value="104.12112">104°7'16"E</geoCoordinate>
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, live corals (
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<taxonomicName id="762B659BC5829958B4391D7D912C2461" class="Anthozoa" family="Poritidae" genus="Porites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porites" order="Scleractinia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">Porites</taxonomicName>
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sp.,
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<taxonomicName id="4170B741F74456D98A2CE0829101F8B8" class="Anthozoa" family="Acroporidae" genus="Montipora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montipora" order="Scleractinia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">Montipora</taxonomicName>
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sp.,
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<taxonomicName id="1B622B7C96260CEBEDB042A07C6D2DE5" class="Anthozoa" family="Acroporidae" genus="Acropora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acropora" order="Scleractinia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">Acropora</taxonomicName>
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sp.), Azman, B.A.R, Kee, A.A., Zuhaimi, S., Maekawa, T., Okashita, T., 22 March 2004.
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</paragraph>
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</subSubSection>
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<subSubSection id="52A730E92A6752E31DA6823A61CE5CEA" pageId="8" pageNumber="9" type="remarks">
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<paragraph id="C0C15062D861EFB1DD958AAE71BDA7F3" pageId="8" pageNumber="9">Remarks.</paragraph>
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<paragraph id="38F5E70B2FE3B5E717D4F52ABB453FCE" pageId="8" pageNumber="9">
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The Pulau Tioman specimens agree well with figures of
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<taxonomicName id="CAEABD30D690B5DB95DF2CD5AD996DF0" class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax</taxonomicName>
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from
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<bibRefCitation id="2E045700B21E72D7C3EC3350A4337013" author="Walker, AO" journalOrPublisher="Report of the Government of Ceylon Pearl Oyster Fishery of the Gulf of Manaar" pageId="21" pageNumber="22" pagination="229 - 300" title="Report on Amphipoda collected by Professor Herdman, at Ceylon, in 1902." volume="17" year="1904">Walker (1904)</bibRefCitation>
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,
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<bibRefCitation id="EA661C0F779D085DB4A7C288026DB266" author="Nagata, K" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="19" pageNumber="20" pagination="163 - 182" title="Preliminary notes on benthic gammaridean Amphipoda from the Zostera region of Mihara Bay, Seto Inland Sea, Japan." volume="8" year="1960">Nagata (1960</bibRefCitation>
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, 1965c),
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<bibRefCitation id="14B0B2B9C4A06C83DA2EC8DDA38C354D" author="Ledoyer, M" journalOrPublisher="Centre National de la Recherche Scientifique-Paris" pageId="19" pageNumber="20" title="Faune de Madagascar. Crustaces Amphipodes Gammariens; Families des Haustoriidae a' Vitjazianidae." volume="59" year="1982 c">Ledoyer (1982c)</bibRefCitation>
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and
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<bibRefCitation id="B1712C4D27693C5F29367C9DA43F2BBD" author="Moore, PG" journalOrPublisher="Journal of Natural History" pageId="19" pageNumber="20" pagination="705 - 732" title="Taxonomic observations on the genera Xenocheria Haswell and Ericthonius Milne-Edwards (Crustacea: Amphipoda) from Australian coastal waters." url="10.1080/00222938800770461" volume="22" year="1988">Moore (1988)</bibRefCitation>
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. This species seems to have a wide-ranging distribution as far as the Arabian Sea. From closer examination of the specimens, the males not only have distally bidentate carpal spur of gnathopod 2 but also a mixture of single dentate carpal spur (see also
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<bibRefCitation id="154BDF15BAFDBF8C9B6C83D2CA911F26" author="Nagata, K" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="19" pageNumber="20" pagination="163 - 182" title="Preliminary notes on benthic gammaridean Amphipoda from the Zostera region of Mihara Bay, Seto Inland Sea, Japan." volume="8" year="1960">Nagata 1960</bibRefCitation>
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;
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<bibRefCitation id="D6C80B313C350FBC460AED65115525F5" author="Ledoyer, M" journalOrPublisher="Recueil des Travaux de la Station Marine Endoume, Fascicule hors serie Supplement" pageId="18" pageNumber="19" pagination="179 - 182" title="Amphipodes tubicoles des feuilles des herbiers de phanerogames marines de la region de Tulear (Madagascar)." volume="9" year="1969">Ledoyer 1969</bibRefCitation>
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,
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<bibRefCitation id="26EBA73EBCBE91647996F8951BFBF9A3" author="Ledoyer, M" journalOrPublisher="Mexico" pageId="19" pageNumber="20" pagination="171 - 200" title="Faune mobile des herbiers de phanerogames marines (Holodule et Thalassia) de la lagune de terminus (Mexique, Campeche). II. Les Gammariens (Crustacea). An. Inst. Cienc. del Mar y Limnol. Univ. Nal. Auton." volume="13" year="1986">1986</bibRefCitation>
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). As mentioned by
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<bibRefCitation id="A56CF1DC590E2A7A9FE5E7C497E5F962" author="Moore, PG" journalOrPublisher="Journal of Natural History" pageId="19" pageNumber="20" pagination="705 - 732" title="Taxonomic observations on the genera Xenocheria Haswell and Ericthonius Milne-Edwards (Crustacea: Amphipoda) from Australian coastal waters." url="10.1080/00222938800770461" volume="22" year="1988">Moore (1988)</bibRefCitation>
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the carpal bidentate spur of ganthopod 2 will eventually be lost in hyperadult males. Unfortunately the stipulation over hyper adult morphological changes has brought several descriptions of new species that actually represented a single species (i.e
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<taxonomicName id="05B1A501964DE445CD435B91D0276655" class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius macrodactylus" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macrodactylus">Ericthonius macrodactylus</taxonomicName>
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). Prior to this,
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<bibRefCitation id="1971E81F4D33E54639D6F350F8BD6662" author="Ledoyer, M" journalOrPublisher="Recueil des Travaux de la Station Marine Endoume, Fascicule hors serie Supplement" pageId="18" pageNumber="19" pagination="179 - 182" title="Amphipodes tubicoles des feuilles des herbiers de phanerogames marines de la region de Tulear (Madagascar)." volume="9" year="1969">Ledoyer (1969)</bibRefCitation>
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synonymized
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<taxonomicName id="16155E2746DCF02D18382FCCF63B9A96" class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius macrodactylus" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macrodactylus">Ericthonius macrodactylus</taxonomicName>
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as a hyperadult form of
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<taxonomicName id="29C5EC54F086D4EC7E3FD566A6E9561A" class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax</taxonomicName>
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. Examination of the Pulau Tioman specimens confirms the presence of this hyper adult morphology: that the basis of pereopod 5 possesses a strongly developed posterodistal lobe. The Pulau Tioman specimens are undoubtedly assigned to
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<taxonomicName id="B53016D4CB5280D554FA4A82F257C2C4" class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax</taxonomicName>
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and constitute the first record of this species for Peninsular Malaysia.
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</paragraph>
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<caption id="BEFE103C7CED9AD384E4AF4724638031" pageId="8" pageNumber="9">
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<paragraph id="7605D8DAA634D8AFD34EABBE58B88A7C" pageId="8" pageNumber="9">
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Figure 6.
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<taxonomicName id="D7200AD3E4C57A7525F33131FBB97586" class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax</taxonomicName>
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(Dana, 1853), male, (UKMMZ-1136), 3.8 mm. Pulau Tioman, South China Sea. Scale for G2 ♂ represents 0.6 mm; A1, A2 HD and ABD scales = 0.5 mm, G1 ♂ scale = 0.25 mm; P5 scale = 0.2 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |