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<document id="5117167AA590F1FDA124B804D6177FBA" ID-CLB-Dataset="56063" ID-DOI="10.5281/zenodo.5305725" ID-GBIF-Dataset="31cb15bf-28f5-4b27-86df-6a1b28cbce41" ID-ISSN="0374-1036" ID-Zenodo-Dep="5305725" ID-ZooBank="E24F1028-C6AC-4323-9ED5-C9B7FF3434ACD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630111997363" checkinUser="marcus" docAuthor="Sekerka, Lukáš" docDate="2016" docId="677FCF20FFD4FFFFFE55FE1AFE8BFAC4" docLanguage="en" docName="ActaEntMusNatPra.2016.56.1.275-344.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 56 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Charidotis annularis" docType="treatment" docVersion="8" lastPageNumber="289" masterDocId="9B46B758FFDAFFF1FFFCFFE6FFBEFFB0" masterDocTitle="Taxonomic and nomenclatural changes in Cassidinae (Coleoptera: Chrysomelidae)" masterLastPageNumber="344" masterPageNumber="275" pageNumber="289" updateTime="1699159820365" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="49DC1FDAA1B338DBD7A8CCC9D6860AF1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EA4724447E20403FF82790B9F45E985F">
<mods:title id="86B84C26E219C6FF724A317301752791">Taxonomic and nomenclatural changes in Cassidinae (Coleoptera: Chrysomelidae)</mods:title>
</mods:titleInfo>
<mods:name id="3AC200313C72FBDB339BD3C8BC6F6D6B" type="personal">
<mods:role id="1BFBC69EC40D2ED2EB0A16406162ADD0">
<mods:roleTerm id="5BBFC4B5F6037B5D05B5EF765A55D5F7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1578945EB326162B3134923470D2591F">Sekerka, Lukáš</mods:namePart>
</mods:name>
<mods:typeOfResource id="EE2293B3F7AAE92ACAC59B740F12159B">text</mods:typeOfResource>
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<mods:title id="17DF150FA2FDDC358F9EBCC209F81820">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="F524F556862566D3C06010C9C8B041E4">
<mods:date id="A0E89F417A7731E5460BF2A3048D12B5">2016</mods:date>
<mods:detail id="C5BFCEA7ED3F9804755A3AA806891301" type="pubDate">
<mods:number id="CD16727F7F99AFA5BF12D55AC2AA425E">2016-07-15</mods:number>
</mods:detail>
<mods:detail id="2B58E2EDBC204D3B6735732CE4C8E32D" type="volume">
<mods:number id="36B858AEF0B14843DE172C91EB81AB4D">56</mods:number>
</mods:detail>
<mods:detail id="CDE6A25B7B9CA89CCB2F1CFEFC34C253" type="issue">
<mods:number id="B3306247ACDFF9B32CE8E9CCC04ED203">1</mods:number>
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<mods:classification id="EEFB58569A7C001D5CB42CA11166728A">journal article</mods:classification>
<mods:identifier id="34DB2F204BC306BAD1DE642A55C50F31" type="CLB-Dataset">56063</mods:identifier>
<mods:identifier id="E299FD20F13EFDEDB1EC389E0B6AB7EF" type="DOI">10.5281/zenodo.5305725</mods:identifier>
<mods:identifier id="615B31536C07B2A7AB40276D236B221F" type="GBIF-Dataset">31cb15bf-28f5-4b27-86df-6a1b28cbce41</mods:identifier>
<mods:identifier id="60C3DCEE2EBD3FF83FAF6939AC0A76B2" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="32B54A0E846762B10F6A6A2FB639471A" type="Zenodo-Dep">5305725</mods:identifier>
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</mods:mods>
<treatment id="677FCF20FFD4FFFFFE55FE1AFE8BFAC4" ID-DOI="http://doi.org/10.5281/zenodo.6449973" ID-GBIF-Taxon="194566200" ID-Zenodo-Dep="6449973" LSID="urn:lsid:plazi:treatment:677FCF20FFD4FFFFFE55FE1AFE8BFAC4" httpUri="http://treatment.plazi.org/id/677FCF20FFD4FFFFFE55FE1AFE8BFAC4" lastPageNumber="289" pageId="14" pageNumber="289">
<subSubSection id="A7CC2DBDFFD4FFFFFE55FE1AFCEAFDA3" box="[425,852,508,531]" pageId="14" pageNumber="289" type="nomenclature">
<paragraph id="EF697E36FFD4FFFFFE55FE1AFCEAFDA3" blockId="14.[425,852,508,531]" box="[425,852,508,531]" pageId="14" pageNumber="289">
<heading id="B421C95AFFD4FFFFFE55FE1AFCEAFDA3" bold="true" box="[425,852,508,531]" centered="true" fontSize="8" level="2" pageId="14" pageNumber="289" reason="3">
<taxonomicName id="28D605B5FFD4FFFFFE55FE1AFCEAFDA3" ID-CoL="TNGW" authority="(Boheman, 1855)" baseAuthorityName="Boheman" baseAuthorityYear="1855" box="[425,852,508,531]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="annularis">
<emphasis id="DDA2A224FFD4FFFFFE55FE1AFCEAFDA3" bold="true" box="[425,852,508,531]" pageId="14" pageNumber="289">
<emphasis id="DDA2A224FFD4FFFFFE55FE1AFD30FDA3" bold="true" box="[425,654,508,531]" italics="true" pageId="14" pageNumber="289">Charidotis annularis</emphasis>
(
<bibRefCitation id="8B4703C7FFD4FFFFFD60FE1AFCF0FDA3" author="Boheman" box="[668,846,508,531]" firstAuthor="Boheman" pageId="14" pageNumber="289" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFD4FFFFFF6BFDC4FCA9FDE0" pageId="14" pageNumber="289" type="reference_group">
<paragraph id="EF697E36FFD4FFFFFF6BFDC4FD76FD85" blockId="14.[151,791,546,592]" box="[151,712,546,565]" pageId="14" pageNumber="289">
<treatmentCitationGroup id="CFC65918FFD4FFFFFF6BFDC4FD76FD85" box="[151,712,546,565]" pageId="14" pageNumber="289">
<treatmentCitation id="6E775827FFD4FFFFFF6BFDC4FDB9FD85" author="BOHEMAN C. H." box="[151,519,546,565]" page="207" pageId="14" pageNumber="289" year="1855">
<taxonomicName id="28D605B5FFD4FFFFFF6BFDC4FDB9FD85" authority="Boheman, 1855: 207" authorityName="Boheman" authorityPageNumber="207" authorityYear="1855" box="[151,519,546,565]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="annularis">
<emphasis id="DDA2A224FFD4FFFFFF6BFDC4FEEEFD85" box="[151,336,546,565]" italics="true" pageId="14" pageNumber="289">Coptocycla annularis</emphasis>
<bibRefCitation id="8B4703C7FFD4FFFFFEA9FDC4FDB9FD85" author="Boheman" box="[341,519,546,565]" firstAuthor="Boheman" page="207" pageId="14" pageNumber="289" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855: 207</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EF697E36FFD4FFFFFF6BFDDBFCA9FDE0" blockId="14.[151,791,546,592]" box="[151,791,573,592]" pageId="14" pageNumber="289">
<treatmentCitationGroup id="CFC65918FFD4FFFFFF6BFDDBFCA9FDE0" box="[151,791,573,592]" pageId="14" pageNumber="289">
<treatmentCitation id="6E775827FFD4FFFFFF6BFDDBFDBDFDE0" author="BOHEMAN C. H." box="[151,515,573,592]" page="210" pageId="14" pageNumber="289" year="1855">
<taxonomicName id="28D605B5FFD4FFFFFF6BFDDBFDBDFDE0" authority="Boheman, 1855: 210" authorityName="Boheman" authorityPageNumber="210" authorityYear="1855" box="[151,515,573,592]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="DDA2A224FFD4FFFFFF6BFDDBFEF5FDE0" box="[151,331,573,592]" italics="true" pageId="14" pageNumber="289">Coptocycla Signoreti</emphasis>
<bibRefCitation id="8B4703C7FFD4FFFFFEACFDDBFDBDFDE0" author="Boheman" box="[336,515,573,592]" firstAuthor="Boheman" page="210" pageId="14" pageNumber="289" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855: 210</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description),
<emphasis id="DDA2A224FFD4FFFFFD3BFDDBFCA9FDE0" bold="true" box="[711,791,573,592]" pageId="14" pageNumber="289">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFD4FFFFFF6BFD80FE11FD54" pageId="14" pageNumber="289" type="materials_examined">
<paragraph id="EF697E36FFD4FFFFFF6BFD80FC34FDC9" blockId="14.[151,1128,614,740]" box="[151,906,614,633]" pageId="14" pageNumber="289">
<emphasis id="DDA2A224FFD4FFFFFF6BFD80FEA0FDC9" bold="true" box="[151,286,614,633]" pageId="14" pageNumber="289">Type localities.</emphasis>
<taxonomicName id="28D605B5FFD4FFFFFEDFFD80FE62FDC9" authorityName="Boheman" authorityYear="1855" box="[291,476,614,633]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="annularis">
<emphasis id="DDA2A224FFD4FFFFFEDFFD80FE62FDC9" box="[291,476,614,633]" italics="true" pageId="14" pageNumber="289">Coptocycla annularis</emphasis>
</taxonomicName>
: ‘Brasilia’;
<taxonomicName id="28D605B5FFD4FFFFFDBFFD80FD18FDC9" authorityName="Boheman" authorityYear="1855" box="[579,678,614,633]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="DDA2A224FFD4FFFFFDBFFD80FD18FDC9" box="[579,678,614,633]" italics="true" pageId="14" pageNumber="289">C. signoreti</emphasis>
</taxonomicName>
: ‘Brasilia ad Rio Janeiro’.
</paragraph>
<paragraph id="EF697E36FFD4FFFFFF6BFD67FD3BFD1F" blockId="14.[151,1128,614,740]" pageId="14" pageNumber="289">
<emphasis id="DDA2A224FFD4FFFFFF6BFD67FEC4FD24" bold="true" box="[151,378,641,660]" pageId="14" pageNumber="289">Type material examined.</emphasis>
<materialsCitation id="5FBE746BFFD4FFFFFE83FD67FD3BFD1F" ID-GBIF-Occurrence="3750508423" collectionCode="NHRS" country="Brazil" location="Rio Jan" pageId="14" pageNumber="289" specimenCode="000000524" specimenCount="1" stateProvince="Rio Jan" typeStatus="syntype">
<taxonomicName id="28D605B5FFD4FFFFFE83FD67FD86FD24" authorityName="Boheman" authorityYear="1855" box="[383,568,641,660]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="annularis">
<emphasis id="DDA2A224FFD4FFFFFE83FD67FD86FD24" box="[383,568,641,660]" italics="true" pageId="14" pageNumber="289">
<originalDetermination id="CCC8C6CAFFD4FFFFFE83FD67FD86FD24" box="[383,568,641,660]" pageId="14" pageNumber="289">Coptocycla annularis</originalDetermination>
</emphasis>
</taxonomicName>
:
<typeStatus id="306DC094FFD4FFFFFDBFFD67FD34FD22" box="[579,650,641,660]" pageId="14" pageNumber="289" type="syntype">SYNTYPE</typeStatus>
: pinned, ‘
<collectingRegion id="2D12B0D4FFD4FFFFFD22FD67FC9CFD24" box="[734,802,641,660]" pageId="14" pageNumber="289">Rio Jan</collectingRegion>
[w, p, s] || F. Sahlb. [w, p, cb] || Type. [w, p, s] || NHRS-SRAH |
<specimenCode id="BF70D64DFFD4FFFFFE8BFD7AFE69FD1F" box="[375,471,668,687]" pageId="14" pageNumber="289">000000524</specimenCode>
[w, p, cb]’ (
<collectionCode id="89C7E6F3FFD4FFFFFDC1FD7AFDC5FD1F" box="[573,635,668,687]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="14" pageNumber="289">NHRS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="EF697E36FFD4FFFFFF49FD50FE11FD54" blockId="14.[151,1128,614,740]" pageId="14" pageNumber="289">
<materialsCitation id="5FBE746BFFD4FFFFFF49FD50FE11FD54" ID-GBIF-Occurrence="3750508372" collectionCode="NHRS" country="Brazil" location="Rio Jan" pageId="14" pageNumber="289" specimenCode="000021832" specimenCount="1" stateProvince="Rio Jan" typeStatus="syntype">
<taxonomicName id="28D605B5FFD4FFFFFF49FD50FED9FD79" authorityName="Boheman" authorityYear="1855" box="[181,359,694,713]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="DDA2A224FFD4FFFFFF49FD50FED9FD79" box="[181,359,694,713]" italics="true" pageId="14" pageNumber="289">
<originalDetermination id="CCC8C6CAFFD4FFFFFF49FD50FED9FD79" box="[181,359,694,713]" pageId="14" pageNumber="289">Coptocycla signoreti</originalDetermination>
</emphasis>
</taxonomicName>
:
<typeStatus id="306DC094FFD4FFFFFE8FFD50FE04FD78" box="[371,442,694,713]" pageId="14" pageNumber="289" type="syntype">SYNTYPE</typeStatus>
: pinned, ‘
<collectingRegion id="2D12B0D4FFD4FFFFFDF2FD50FDECFD79" box="[526,594,694,713]" pageId="14" pageNumber="289">Rio Jan</collectingRegion>
[w, p, s] || F. Sahlb. [w, p, cb] || Type. [w, p, s] || NHRS-JLKB |
<specimenCode id="BF70D64DFFD4FFFFFF5DFD37FEBFFD54" box="[161,257,721,740]" pageId="14" pageNumber="289">000021832</specimenCode>
[w, p, cb]’ (
<collectionCode id="89C7E6F3FFD4FFFFFE9BFD37FE1BFD54" box="[359,421,721,740]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="14" pageNumber="289">NHRS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFD4FFFFFF6BFD1BFEF7FAA4" pageId="14" pageNumber="289" type="discussion">
<paragraph id="EF697E36FFD4FFFFFF6BFD1BFE2CFCC4" blockId="14.[151,1127,765,1396]" pageId="14" pageNumber="289">
<emphasis id="DDA2A224FFD4FFFFFF6BFD1BFEB8FCA4" bold="true" box="[151,262,765,788]" pageId="14" pageNumber="289">Remarks.</emphasis>
Both taxa were described in the same book by
<bibRefCitation id="8B4703C7FFD4FFFFFCFEFD1BFC0AFCA4" author="BOHEMAN" box="[770,948,765,788]" firstAuthor="BOHEMAN" pageId="14" pageNumber="289" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">BOHEMAN (1855)</bibRefCitation>
just a few pages apart.
<bibRefCitation id="8B4703C7FFD4FFFFFF27FCFBFECDFC84" author="SPAETH" box="[219,371,797,820]" firstAuthor="SPAETH" pageId="14" pageNumber="289" refId="ref50998" refString="SPAETH F. 1914: Chrysomelidae: 16. Cassidinae. In: SCHENKLING S. (ed.): Coleopterorum Catalogus. Pars 62. W. Junk, Berlin, 180 pp." type="book" year="1914">SPAETH (1914)</bibRefCitation>
transferred both to
<taxonomicName id="28D605B5FFD4FFFFFDB4FCFBFCD1FC84" authority="Boheman, 1854" authorityName="Boheman" authorityYear="1854" box="[584,879,797,820]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="genus">
<emphasis id="DDA2A224FFD4FFFFFDB4FCFBFD04FC84" box="[584,698,797,820]" italics="true" pageId="14" pageNumber="289">Charidotis</emphasis>
<bibRefCitation id="8B4703C7FFD4FFFFFD3EFCFBFCD1FC84" author="Boheman" box="[706,879,797,820]" firstAuthor="Boheman" pageId="14" pageNumber="289" refId="ref47930" refString="BOHEMAN C. H. 1854: Monographia Cassididarum. Tomus Secundus. Officina Norstedtiana, Holmiae, 506 pp. + 2 pls." type="book" year="1854">Boheman, 1854</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="8B4703C7FFD4FFFFFC86FCFBFBACFC84" author="SPAETH" box="[890,1042,797,820]" firstAuthor="SPAETH" pageId="14" pageNumber="289" pagination="66 - 99" refId="ref51359" refString="SPAETH F. 1936: Bestimmungstabelle der Gattung Charidotis (Col. Chrys. Cassid.). Sbornik Entomologickeho Oddeleni Narodniho Musea v Praze 14: 66 - 99." type="journal article" year="1936">SPAETH (1936)</bibRefCitation>
revised the genus and separated the two taxa only by colour:
<taxonomicName id="28D605B5FFD4FFFFFD34FCDBFCF2FCE4" authorityName="Boheman" authorityYear="1855" box="[712,844,829,852]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="annularis">
<emphasis id="DDA2A224FFD4FFFFFD34FCDBFCF2FCE4" box="[712,844,829,852]" italics="true" pageId="14" pageNumber="289">C. annularis</emphasis>
</taxonomicName>
with red disc of elytra and
<taxonomicName id="28D605B5FFD4FFFFFF6BFCBBFEAAFCC4" authorityName="Boheman" authorityYear="1855" box="[151,276,861,884]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="DDA2A224FFD4FFFFFF6BFCBBFEAAFCC4" box="[151,276,861,884]" italics="true" pageId="14" pageNumber="289">C. signoreti</emphasis>
</taxonomicName>
with black.
</paragraph>
<paragraph id="EF697E36FFD4FFFFFF49FC9BFD34FB24" blockId="14.[151,1127,765,1396]" pageId="14" pageNumber="289">
Examination of both types as well as additional historic specimens from various institutions revealed that the colour of elytra is variable in these taxa. Even the
<typeStatus id="306DC094FFD4FFFFFC8DFC7BFC7AFC04" box="[881,964,925,948]" pageId="14" pageNumber="289" type="syntype">syntype</typeStatus>
of
<taxonomicName id="28D605B5FFD4FFFFFC15FC7BFBD9FC04" authorityName="Boheman" authorityYear="1855" box="[1001,1127,925,948]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="DDA2A224FFD4FFFFFC15FC7BFBD9FC04" box="[1001,1127,925,948]" italics="true" pageId="14" pageNumber="289">C. signoreti</emphasis>
</taxonomicName>
has variegated elytra, basally and centrally more black and laterally reddish-brown. Uniformly black specimens are very rare in collections and as result
<taxonomicName id="28D605B5FFD4FFFFFCDEFC3BFC1FFC44" authorityName="Boheman" authorityYear="1855" box="[802,929,989,1012]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="DDA2A224FFD4FFFFFCDEFC3BFC1FFC44" box="[802,929,989,1012]" italics="true" pageId="14" pageNumber="289">C. signoreti</emphasis>
</taxonomicName>
was listed only in catalogues while
<taxonomicName id="28D605B5FFD4FFFFFEAFFC1BFE64FBA4" authorityName="Boheman" authorityYear="1855" box="[339,474,1021,1044]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="annularis">
<emphasis id="DDA2A224FFD4FFFFFEAFFC1BFE64FBA4" box="[339,474,1021,1044]" italics="true" pageId="14" pageNumber="289">C. annularis</emphasis>
</taxonomicName>
was repeatedly reported (see
<bibRefCitation id="8B4703C7FFD4FFFFFCDCFC1BFC6AFBA4" author="BOROWIEC" box="[800,980,1021,1044]" firstAuthor="BOROWIEC" pageId="14" pageNumber="289" refId="ref48283" refString="BOROWIEC L. 1999 a: A world catalogue of the Cassidinae (Coleoptera: Chrysomelidae). Biologica Silesiae, Wroclaw, 476 pp." type="book" year="1999" yearSuffix="a">BOROWIEC 1999a</bibRefCitation>
,
<bibRefCitation id="8B4703C7FFD4FFFFFC1DFC1BFED5FB84" author="BOROWIEC &amp; SWIETOJANSKA" firstAuthor="BOROWIEC" pageId="14" pageNumber="289" refId="ref48617" refString="BOROWIEC L. &amp; SWIETOJANSKA J. 2016: Cassidinae of the world - an interactive manual (Coleoptera: Chrysomelidae). Permanent electronic publication available from: http: // culex. biol. uni. wroc. pl / cassidae / katalog % 20 internetowy / index. htm (last accessed on 18 th May 2016)." type="url" year="2016">BOROWIEC &amp; ŚWIĘTOJAŃSKA 2016</bibRefCitation>
). In my opinion both nominal taxa represent colour morphs of a single species as all other morphological characters are identical. Such variability in colouration is not uncommon in similar species of
<taxonomicName id="28D605B5FFD4FFFFFDE8FBBBFD38FBC4" authorityName="Boheman" authorityYear="1854" box="[532,646,1117,1140]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="genus">
<emphasis id="DDA2A224FFD4FFFFFDE8FBBBFD38FBC4" box="[532,646,1117,1140]" italics="true" pageId="14" pageNumber="289">Charidotis</emphasis>
</taxonomicName>
and it is known for instance in
<taxonomicName id="28D605B5FFD4FFFFFC30FBBBFEFEFB24" authority="Boheman, 1855" authorityName="Boheman" authorityYear="1855" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="auroguttata">
<emphasis id="DDA2A224FFD4FFFFFC30FBBBFBD8FBC4" box="[972,1126,1117,1140]" italics="true" pageId="14" pageNumber="289">C. auroguttata</emphasis>
<bibRefCitation id="8B4703C7FFD4FFFFFF6BFB9BFEFEFB24" author="Boheman" box="[151,320,1149,1172]" firstAuthor="Boheman" pageId="14" pageNumber="289" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="28D605B5FFD4FFFFFE88FB9BFD38FB24" authority="Spaeth, 1902" authorityName="Spaeth" authorityYear="1902" box="[372,646,1149,1172]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="porosula">
<emphasis id="DDA2A224FFD4FFFFFE88FB9BFE4FFB24" box="[372,497,1149,1172]" italics="true" pageId="14" pageNumber="289">C. porosula</emphasis>
<bibRefCitation id="8B4703C7FFD4FFFFFE04FB9BFD38FB24" author="Spaeth" box="[504,646,1149,1172]" firstAuthor="Spaeth" pageId="14" pageNumber="289" pagination="81 - 102" refId="ref50784" refString="SPAETH F. 1902: Neue Cassiden aus Peru. Deutsche Entomologische Zeitschrift 1902: 81 - 102." type="journal article" year="1902">Spaeth, 1902</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="EF697E36FFD4FFFFFF49FB7BFEF7FAA4" blockId="14.[151,1127,765,1396]" pageId="14" pageNumber="289">
Since both taxa were described in the same publication I followed the Principle of the first reviser (
<bibRefCitation id="8B4703C7FFD4FFFFFF13FB5BFED1FB64" author="ICZN" box="[239,367,1213,1236]" firstAuthor="ICZN" pageId="14" pageNumber="289" refId="ref49784" refString="ICZN 1999: International code of zoological nomenclature. Fourth edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Article 24.2) and chose the name
<taxonomicName id="28D605B5FFD4FFFFFD1AFB5BFCD3FB64" authorityName="Boheman" authorityYear="1855" box="[742,877,1213,1236]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="289" phylum="Arthropoda" rank="species" species="annularis">
<emphasis id="DDA2A224FFD4FFFFFD1AFB5BFCD3FB64" box="[742,877,1213,1236]" italics="true" pageId="14" pageNumber="289">C. annularis</emphasis>
</taxonomicName>
as valid for this taxon, as it has been used recently and the species is more frequently identified in the collections under this name.
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFD4FFFFFF6BFAFBFE8BFAC4" pageId="14" pageNumber="289" type="distribution">
<paragraph id="EF697E36FFD4FFFFFF6BFAFBFE8BFAC4" blockId="14.[151,1127,765,1396]" pageId="14" pageNumber="289">
<emphasis id="DDA2A224FFD4FFFFFF6BFAFBFE94FA84" bold="true" box="[151,298,1309,1332]" pageId="14" pageNumber="289">Distribution.</emphasis>
<collectingCountry id="97C13EA6FFD4FFFFFECEFAFBFEC9FA84" box="[306,375,1309,1332]" name="Brazil" pageId="14" pageNumber="289">Brazil</collectingCountry>
:
<collectingRegion id="2D12B0D4FFD4FFFFFE7EFAFBFE7EFA84" box="[386,448,1309,1332]" country="Brazil" name="Bahia" pageId="14" pageNumber="289">Bahia</collectingRegion>
(
<bibRefCitation id="8B4703C7FFD4FFFFFE2EFAFBFDE3FA84" author="SPAETH" box="[466,605,1309,1332]" firstAuthor="SPAETH" pageId="14" pageNumber="289" pagination="66 - 99" refId="ref51359" refString="SPAETH F. 1936: Bestimmungstabelle der Gattung Charidotis (Col. Chrys. Cassid.). Sbornik Entomologickeho Oddeleni Narodniho Musea v Praze 14: 66 - 99." type="journal article" year="1936">SPAETH 1936</bibRefCitation>
),
<collectingRegion id="2D12B0D4FFD4FFFFFD8DFAFBFCBCFA84" box="[625,770,1309,1332]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="289">Minas Gerais</collectingRegion>
(
<bibRefCitation id="8B4703C7FFD4FFFFFCEEFAFBFC77FA84" author="BOROWIEC" box="[786,969,1309,1332]" firstAuthor="BOROWIEC" pageId="14" pageNumber="289" pagination="615 - 722" refId="ref48513" refString="BOROWIEC L. 2009 c: New records of Neotropical tortoise beetles (Coleoptera: Chrysomelidae: Cassidinae). Genus 20: 615 - 722." type="journal article" year="2009" yearSuffix="c">BOROWIEC 2009c</bibRefCitation>
),
<collectingRegion id="2D12B0D4FFD4FFFFFC21FAFBFB98FA84" box="[989,1062,1309,1332]" country="Brazil" name="Parana" pageId="14" pageNumber="289">Paraná</collectingRegion>
(FERNANDES &amp; LINZMEIER 2012),
<collectingRegion id="2D12B0D4FFD4FFFFFE3AFADBFDDDFAE4" box="[454,611,1341,1364]" country="Brazil" name="Rio de Janeiro" pageId="14" pageNumber="289">Rio de Janeiro</collectingRegion>
(
<bibRefCitation id="8B4703C7FFD4FFFFFD8FFADBFCA9FAE4" author="BOHEMAN" box="[627,791,1341,1364]" firstAuthor="BOHEMAN" pageId="14" pageNumber="289" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">BOHEMAN 1855</bibRefCitation>
,
<bibRefCitation id="8B4703C7FFD4FFFFFCDFFADBFC58FAE4" author="FLINTE" box="[803,998,1341,1364]" etAl="et al." firstAuthor="FLINTE" pageId="14" pageNumber="289" pagination="571 - 614" refId="ref49297" refString="FLINTE V., BOROWIEC L., FERITAS S., VIANA J. H., FERNANDES F. R., NOGUEIRA-DE-SA F., MACEDO M. V. &amp; MONTEIRO R. F. 2009: Tortoise beetles of the State of Rio de Janeiro, Brazil (Coleoptera: Chrysomelidae: Cassidinae). Genus 20: 571 - 614." type="journal article" year="2009">FLINTE et al. 2009</bibRefCitation>
),
<collectingRegion id="2D12B0D4FFD4FFFFFC06FADBFBD9FAE4" box="[1018,1127,1341,1364]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="289">São Paulo</collectingRegion>
(
<bibRefCitation id="8B4703C7FFD4FFFFFF5CFABBFE94FAC4" author="SPAETH" box="[160,298,1373,1396]" firstAuthor="SPAETH" pageId="14" pageNumber="289" pagination="66 - 99" refId="ref51359" refString="SPAETH F. 1936: Bestimmungstabelle der Gattung Charidotis (Col. Chrys. Cassid.). Sbornik Entomologickeho Oddeleni Narodniho Musea v Praze 14: 66 - 99." type="journal article" year="1936">SPAETH 1936</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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