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<document id="936E9F5F90D3801E28B870106D689406" ID-CLB-Dataset="56063" ID-DOI="10.5281/zenodo.5305725" ID-GBIF-Dataset="31cb15bf-28f5-4b27-86df-6a1b28cbce41" ID-ISSN="0374-1036" ID-Zenodo-Dep="5305725" ID-ZooBank="E24F1028-C6AC-4323-9ED5-C9B7FF3434ACD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630111997363" checkinUser="marcus" docAuthor="Sekerka, Lukáš" docDate="2016" docId="677FCF20FFC9FFE2FEB2FD6EFDC5FADA" docLanguage="en" docName="ActaEntMusNatPra.2016.56.1.275-344.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 56 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Helocassis ramosa Sekerka 2016, comb. nov." docType="treatment" docVersion="8" lastPageNumber="294" masterDocId="9B46B758FFDAFFF1FFFCFFE6FFBEFFB0" masterDocTitle="Taxonomic and nomenclatural changes in Cassidinae (Coleoptera: Chrysomelidae)" masterLastPageNumber="344" masterPageNumber="275" pageNumber="294" updateTime="1699159820365" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A224759FB79B909A6800F980771842DF">Taxonomic and nomenclatural changes in Cassidinae (Coleoptera: Chrysomelidae)</mods:title>
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<mods:namePart id="59469D7DE338F9DBA4513943208A8DD8">Sekerka, Lukáš</mods:namePart>
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<mods:title id="B5190058062472A16DEB91A5D0D34920">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="DCD6BAA04648A893F0CF8FFDF5DF9B1A">2016</mods:date>
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<mods:number id="C2FD10D29A062FD0095643221373C1A3">2016-07-15</mods:number>
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<treatment id="677FCF20FFC9FFE2FEB2FD6EFDC5FADA" ID-DOI="http://doi.org/10.5281/zenodo.5332681" ID-GBIF-Taxon="194566213" ID-Zenodo-Dep="5332681" LSID="urn:lsid:plazi:treatment:677FCF20FFC9FFE2FEB2FD6EFDC5FADA" httpUri="http://treatment.plazi.org/id/677FCF20FFC9FFE2FEB2FD6EFDC5FADA" lastPageNumber="294" pageId="19" pageNumber="294">
<subSubSection id="A7CC2DBDFFC9FFE2FEB2FD6EFCDAFD2F" box="[334,868,648,671]" pageId="19" pageNumber="294" type="nomenclature">
<paragraph id="EF697E36FFC9FFE2FEB2FD6EFCDAFD2F" blockId="19.[334,868,648,671]" box="[334,868,648,671]" pageId="19" pageNumber="294">
<heading id="B421C95AFFC9FFE2FEB2FD6EFCDAFD2F" bold="true" box="[334,868,648,671]" centered="true" fontSize="8" level="2" pageId="19" pageNumber="294" reason="3">
<emphasis id="DDA2A224FFC9FFE2FEB2FD6EFCDAFD2F" bold="true" box="[334,868,648,671]" pageId="19" pageNumber="294">
<taxonomicName id="28D605B5FFC9FFE2FEB2FD6EFD5CFD2F" authority="(Boheman, 1855)" authorityName="Sekerka" authorityYear="2016" baseAuthorityName="Boheman" baseAuthorityYear="1855" box="[334,738,648,671]" class="Insecta" family="Chrysomelidae" genus="Helocassis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="species" species="ramosa" status="comb. nov.">
<emphasis id="DDA2A224FFC9FFE2FEB2FD6EFDA5FD2F" bold="true" box="[334,539,648,671]" italics="true" pageId="19" pageNumber="294">Helocassis ramosa</emphasis>
(
<bibRefCitation id="8B4703C7FFC9FFE2FDD5FD6EFD65FD2F" author="Boheman" box="[553,731,648,671]" firstAuthor="Boheman" pageId="19" pageNumber="294" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C6911F5FFFC9FFE2FD15FD6EFCDAFD2F" box="[745,868,648,671]" pageId="19" pageNumber="294" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFC9FFE2FF8DFD48FD2EFD71" box="[113,656,686,705]" pageId="19" pageNumber="294" type="reference_group">
<paragraph id="EF697E36FFC9FFE2FF8DFD48FD2EFD71" blockId="19.[113,656,686,705]" box="[113,656,686,705]" pageId="19" pageNumber="294">
<treatmentCitationGroup id="CFC65918FFC9FFE2FF8DFD48FD2EFD71" box="[113,656,686,705]" pageId="19" pageNumber="294">
<treatmentCitation id="6E775827FFC9FFE2FF8DFD48FE6EFD71" author="BOHEMAN C. H." box="[113,464,686,705]" page="393" pageId="19" pageNumber="294" year="1855">
<taxonomicName id="28D605B5FFC9FFE2FF8DFD48FE6EFD71" authority="Boheman, 1855: 393" authorityName="Boheman" authorityPageNumber="393" authorityYear="1855" box="[113,464,686,705]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DDA2A224FFC9FFE2FF8DFD48FEA6FD71" box="[113,280,686,705]" italics="true" pageId="19" pageNumber="294">Coptocycla ramosa</emphasis>
<bibRefCitation id="8B4703C7FFC9FFE2FEE1FD48FE6EFD71" author="Boheman" box="[285,464,686,705]" firstAuthor="Boheman" page="393" pageId="19" pageNumber="294" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855: 393</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFC9FFE2FF8DFD31FC22FC8A" pageId="19" pageNumber="294" type="materials_examined">
<paragraph id="EF697E36FFC9FFE2FF8DFD31FD80FD5A" blockId="19.[113,1089,727,826]" box="[113,574,727,746]" pageId="19" pageNumber="294">
<emphasis id="DDA2A224FFC9FFE2FF8DFD31FF54FD5A" bold="true" box="[113,234,727,746]" pageId="19" pageNumber="294">
<typeStatus id="306DC094FFC9FFE2FF8DFD31FF20FD5A" box="[113,158,727,746]" pageId="19" pageNumber="294">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="97C13EA6FFC9FFE2FF13FD31FEFBFD5A" box="[239,325,727,746]" name="Colombia" pageId="19" pageNumber="294">Colombia</collectingCountry>
,
<collectingRegion id="2D12B0D4FFC9FFE2FEB2FD31FE48FD5A" box="[334,502,727,746]" country="Colombia" name="Tolima" pageId="19" pageNumber="294">Tolima Department</collectingRegion>
, Honda.
</paragraph>
<paragraph id="EF697E36FFC9FFE2FF8DFD14FC22FC8A" blockId="19.[113,1089,727,826]" pageId="19" pageNumber="294">
<emphasis id="DDA2A224FFC9FFE2FF8DFD14FEEAFCB5" bold="true" box="[113,340,754,773]" pageId="19" pageNumber="294">Type material examined.</emphasis>
<materialsCitation id="5FBE746BFFC9FFE2FEA5FD14FC22FC8A" ID-GBIF-Occurrence="3750508382" collectionCode="BMNH" country="Colombia" pageId="19" pageNumber="294" specimenCount="1" stateProvince="Tolima Department" typeStatus="holotype">
<typeStatus id="306DC094FFC9FFE2FEA5FD14FE13FCB3" box="[345,429,754,773]" pageId="19" pageNumber="294" type="holotype">HOLOTYPE</typeStatus>
: pinned: E. Coll. |
<superScript id="18A3D37EFFC9FFE2FDABFD14FD36FD4C" attach="left" box="[599,648,753,773]" fontSize="5" pageId="19" pageNumber="294">Chevt</superScript>
. [w, p, cb] || Type [w, p, s, c, rf] || N Grena | Honda | G Goudot [g, hw by Chevrolat, s] || 66 [g, p, s] || aspidom | ramosa | Bhn | Honda [w, hw by Chevrolat, s] || 67·56 [w, p, s] ||
<taxonomicName id="28D605B5FFC9FFE2FF35FCC1FE94FC8A" authorityName="Chevrolat" authorityYear="1836" box="[201,298,807,826]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="genus">Coptocycla</taxonomicName>
| ramosa, | Bohem | Type in coll. Chevr. [w, hw by Gahan, cb] (
<collectionCode id="89C7E6F3FFC9FFE2FCB1FCC1FC2CFC8A" box="[845,914,807,826]" pageId="19" pageNumber="294">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFC9FFE2FF8DFCB5FE3AFAFA" pageId="19" pageNumber="294" type="discussion">
<paragraph id="EF697E36FFC9FFE2FF8DFCB5FDCEFB5A" blockId="19.[113,1089,851,1386]" pageId="19" pageNumber="294">
<emphasis id="DDA2A224FFC9FFE2FF8DFCB5FF5EFCDA" bold="true" box="[113,224,851,874]" pageId="19" pageNumber="294">Remarks.</emphasis>
<bibRefCitation id="8B4703C7FFC9FFE2FF1AFCB5FE26FCDA" author="BOHEMAN" box="[230,408,851,874]" firstAuthor="BOHEMAN" pageId="19" pageNumber="294" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">BOHEMAN (1855)</bibRefCitation>
described this species presumably from a single specimen from the collection of Chevrolat. He compared it with
<taxonomicName id="28D605B5FFC9FFE2FD7FFC95FBADFC3A" authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[643,1043,883,906]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="DDA2A224FFC9FFE2FD7FFC95FCEDFC3A" box="[643,851,883,906]" italics="true" pageId="19" pageNumber="294">Coptocycla clavata</emphasis>
(Fabricius, 1798)
</taxonomicName>
and it was placed in the same group as other species latter transferred to the genus
<taxonomicName id="28D605B5FFC9FFE2FC31FC75FF44FC7A" authority="Spaeth, 1952" authorityName="Spaeth" authorityYear="1952" class="Insecta" family="Chrysomelidae" genus="Helocassis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="DDA2A224FFC9FFE2FC31FC75FBFFFC1A" box="[973,1089,915,938]" italics="true" pageId="19" pageNumber="294">Helocassis</emphasis>
Spaeth, 1952
</taxonomicName>
(in
<bibRefCitation id="8B4703C7FFC9FFE2FEDEFC55FE15FC7A" author="HINCKS" box="[290,427,947,970]" firstAuthor="HINCKS" pageId="19" pageNumber="294" pagination="327 - 358" refId="ref49715" refString="HINCKS W. D. 1952: The genera of the Cassidinae (Coleoptera: Chrysomelidae). Transactions of the Royal Entomological Society of London 103: 327 - 358." type="journal article" year="1952">HINCKS 1952</bibRefCitation>
).
<bibRefCitation id="8B4703C7FFC9FFE2FE47FC55FDEEFC7A" author="SPAETH" box="[443,592,947,970]" firstAuthor="SPAETH" pageId="19" pageNumber="294" pagination="135 - 158" refId="ref51390" refString="SPAETH F. 1937: Neue Cassidinen des Rijksmuseums in Leiden und meiner Sammlung. Temminckia 2: 135 - 158." type="journal article" year="1937">SPAETH (1937)</bibRefCitation>
however, placed
<taxonomicName id="28D605B5FFC9FFE2FCF9FC55FC6DFC7A" authorityName="Boheman" authorityYear="1855" box="[773,979,947,970]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DDA2A224FFC9FFE2FCF9FC55FC6DFC7A" box="[773,979,947,970]" italics="true" pageId="19" pageNumber="294">Coptocycla ramosa</emphasis>
</taxonomicName>
in
<taxonomicName id="28D605B5FFC9FFE2FC0DFC55FED8FC5A" authority="Spaeth, 1899" authorityName="Spaeth" authorityYear="1899" class="Insecta" family="Chrysomelidae" genus="Plagiometriona" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="DDA2A224FFC9FFE2FC0DFC55FF6FFC5A" italics="true" pageId="19" pageNumber="294">Plagiometriona</emphasis>
<bibRefCitation id="8B4703C7FFC9FFE2FF24FC35FED8FC5A" author="Spaeth" box="[216,358,979,1002]" firstAuthor="Spaeth" pageId="19" pageNumber="294" pagination="213 - 221" refId="ref50724" refString="SPAETH F. 1899: Beschreibung einiger neuer Cassiden nebst synonymischen Bemerkungen III. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 49: 213 - 221 + pl. 5." type="journal article" year="1899">Spaeth, 1899</bibRefCitation>
</taxonomicName>
, most likely based on the primary description. I have studied its
<typeStatus id="306DC094FFC9FFE2FBEFFC35FBFFFC5A" box="[1043,1089,979,1002]" pageId="19" pageNumber="294">type</typeStatus>
and the species in fact belongs to
<taxonomicName id="28D605B5FFC9FFE2FE2AFC15FDF4FBBA" authorityName="Spaeth" authorityYear="1952" box="[470,586,1011,1034]" class="Insecta" family="Chrysomelidae" genus="Helocassis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="DDA2A224FFC9FFE2FE2AFC15FDF4FBBA" box="[470,586,1011,1034]" italics="true" pageId="19" pageNumber="294">Helocassis</emphasis>
</taxonomicName>
as it has a head with frontoclypeal sulci, which are absent in
<taxonomicName id="28D605B5FFC9FFE2FF00FBF5FE1DFB9A" authorityName="Spaeth" authorityYear="1899" box="[252,419,1043,1066]" class="Insecta" family="Chrysomelidae" genus="Plagiometriona" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="DDA2A224FFC9FFE2FF00FBF5FE1DFB9A" box="[252,419,1043,1066]" italics="true" pageId="19" pageNumber="294">Plagiometriona</emphasis>
</taxonomicName>
. The species is morphologically most similar to
<taxonomicName id="28D605B5FFC9FFE2FC58FBF5FE90FBFA" authority="(Boheman, 1855)" baseAuthorityName="Boheman" baseAuthorityYear="1855" class="Insecta" family="Chrysomelidae" genus="Helocassis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="species" species="crucipennis">
<emphasis id="DDA2A224FFC9FFE2FC58FBF5FBFEFB9A" box="[932,1088,1043,1066]" italics="true" pageId="19" pageNumber="294">H. crucipennis</emphasis>
(
<bibRefCitation id="8B4703C7FFC9FFE2FF85FBD5FE99FBFA" author="Boheman" box="[121,295,1075,1098]" firstAuthor="Boheman" pageId="19" pageNumber="294" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855</bibRefCitation>
)
</taxonomicName>
but differs in the base of elytra much wider than the base of pronotum, humeral angles protruding anterad, and subpentagonal body outline with straight lateral sides of elytra while
<taxonomicName id="28D605B5FFC9FFE2FEE5FB95FE06FB3A" baseAuthorityName="Boheman" baseAuthorityYear="1855" box="[281,440,1139,1162]" class="Insecta" family="Chrysomelidae" genus="Helocassis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="species" species="crucipennis">
<emphasis id="DDA2A224FFC9FFE2FEE5FB95FE06FB3A" box="[281,440,1139,1162]" italics="true" pageId="19" pageNumber="294">H. crucipennis</emphasis>
</taxonomicName>
has oval body with weakly projecting humeral angles. The body shape is more similar to
<taxonomicName id="28D605B5FFC9FFE2FE42FB75FC99FB1A" authority="(Boheman, 1855)" baseAuthorityName="Boheman" baseAuthorityYear="1855" box="[446,807,1171,1194]" class="Insecta" family="Chrysomelidae" genus="Helocassis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="species" species="testudinaria">
<emphasis id="DDA2A224FFC9FFE2FE42FB75FDDCFB1A" box="[446,610,1171,1194]" italics="true" pageId="19" pageNumber="294">H. testudinaria</emphasis>
(
<bibRefCitation id="8B4703C7FFC9FFE2FD8EFB75FC9EFB1A" author="Boheman" box="[626,800,1171,1194]" firstAuthor="Boheman" pageId="19" pageNumber="294" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="28D605B5FFC9FFE2FCA3FB75FC71FB1A" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[863,975,1171,1194]" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="DDA2A224FFC9FFE2FCA3FB75FC71FB1A" box="[863,975,1171,1194]" italics="true" pageId="19" pageNumber="294">H. clavata</emphasis>
</taxonomicName>
, however, both have rugose surface of elytra and high postscutellar tubercle, which is very low in
<taxonomicName id="28D605B5FFC9FFE2FBDBFB55FF7FFB5A" authorityName="Boheman" authorityYear="1855" class="Insecta" family="Chrysomelidae" genus="Coptocycla" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="294" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DDA2A224FFC9FFE2FBDBFB55FF7FFB5A" italics="true" pageId="19" pageNumber="294">H. ramosa</emphasis>
</taxonomicName>
and it also has smooth surface of elytra.
</paragraph>
<paragraph id="EF697E36FFC9FFE2FF6CFB15FE3AFAFA" blockId="19.[113,1089,851,1386]" pageId="19" pageNumber="294">
<bibRefCitation id="8B4703C7FFC9FFE2FF6CFB15FEFFFABA" author="BOHEMAN" box="[144,321,1267,1290]" firstAuthor="BOHEMAN" pageId="19" pageNumber="294" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">BOHEMAN (1855)</bibRefCitation>
mentioned Columbia as the
<typeStatus id="306DC094FFC9FFE2FD7CFB15FD10FABA" box="[640,686,1267,1290]" pageId="19" pageNumber="294">type</typeStatus>
locality, however, the specimen bears more detailed locality data. The
<typeStatus id="306DC094FFC9FFE2FE3EFAF5FE4EFA9A" box="[450,496,1299,1322]" pageId="19" pageNumber="294">type</typeStatus>
locality is here specified to Honda situated in the
<collectingRegion id="2D12B0D4FFC9FFE2FC09FAF5FF51FAFA" country="Colombia" name="Tolima" pageId="19" pageNumber="294">Tolima Department</collectingRegion>
of
<collectingCountry id="97C13EA6FFC9FFE2FEEFFAD5FE3EFAFA" box="[275,384,1331,1354]" name="Colombia" pageId="19" pageNumber="294">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFC9FFE2FF8DFAB5FDC5FADA" box="[113,635,1363,1386]" pageId="19" pageNumber="294" type="distribution">
<paragraph id="EF697E36FFC9FFE2FF8DFAB5FDC5FADA" blockId="19.[113,1089,851,1386]" box="[113,635,1363,1386]" pageId="19" pageNumber="294">
<emphasis id="DDA2A224FFC9FFE2FF8DFAB5FEBAFADA" bold="true" box="[113,260,1363,1386]" pageId="19" pageNumber="294">Distribution.</emphasis>
<collectingCountry id="97C13EA6FFC9FFE2FEF6FAB5FEC9FADA" box="[266,375,1363,1386]" name="Colombia" pageId="19" pageNumber="294">Colombia</collectingCountry>
:
<collectingRegion id="2D12B0D4FFC9FFE2FE7DFAB5FE72FADA" box="[385,460,1363,1386]" country="Colombia" name="Tolima" pageId="19" pageNumber="294">Tolima</collectingRegion>
(present paper).
</paragraph>
</subSubSection>
</treatment>
</document>