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<document id="85F7E60DCA962015544835D81B0DFA89" ID-CLB-Dataset="22806" ID-DOI="10.11646/zootaxa.4407.1.2" ID-GBIF-Dataset="8e0aac3a-6556-4b96-a67e-fce9f0db0ba9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3766290" ID-ZooBank="B45E9ABD-CB52-4EE7-91D6-783E8E958AC5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587827908598" checkinUser="plazi" docAuthor="Short, Andrew Edward Z. &amp; Girón, Jennifer C." docDate="2018" docId="676A87E0FFB4B15B2392FA45469D0F2E" docLanguage="en" docName="zootaxa.4407.1.2.pdf.imf" docOrigin="Zootaxa 4407 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helochares (Hydrobaticus) trujillo Short &amp; Girón 2018, n. sp." docType="treatment" docUuid="68539169-5FAC-43B9-8839-69141BDB155E" docUuidSource="ZooBank" docVersion="6" lastPageNumber="46" masterDocId="9B53FF98FFA4B14A2305FFE3420C0E4A" masterDocTitle="Review of the Helochares (Hydrobaticus) MacLeay of the New World (Coleoptera: Hydrophilidae: Acidocerinae)" masterLastPageNumber="50" masterPageNumber="29" pageNumber="45" updateTime="1698835862916" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="DC33F55FEAF1CBA88B43ADBB8532B5B5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4B71B4A7579317BBFD92DCEE0A0703A3">
<mods:title id="5264C5105D9E56C8D3D5214C9D56258D">Review of the Helochares (Hydrobaticus) MacLeay of the New World (Coleoptera: Hydrophilidae: Acidocerinae)</mods:title>
</mods:titleInfo>
<mods:name id="02379BD12AD6854E78D733CFEDDFDFFB" type="personal">
<mods:role id="C2B3D07B51769914C7FF1D739706DF7D">
<mods:roleTerm id="E1883B40B9212E0E54499C506C6DB310">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7FC6D5C148D2C045797EC75EFE958702">Short, Andrew Edward Z.</mods:namePart>
</mods:name>
<mods:name id="50E857CD1DB5E9404695ED6A411C9F09" type="personal">
<mods:role id="F009A191F61589F7759107DC0250E4A8">
<mods:roleTerm id="C91AE50F72C044483239BE435332BD01">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B53728F9F49429796680E3EC1629BFBD">Girón, Jennifer C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="72742F31271E0D49A07B8CD4A9F0D6A8">text</mods:typeOfResource>
<mods:relatedItem id="10BA0186BC6A7F816B3B53CD0D91C423" type="host">
<mods:titleInfo id="468F678D7984082B20ED2FA3E9660C8D">
<mods:title id="61E45095763DE4C1D20C4196A084C548">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3243CD0E5E282F07DA8F5B79ECFBDA01">
<mods:date id="DAF38B0094000865235D984B90D0EF44">2018</mods:date>
<mods:detail id="F4F7E94F53D31EEDDB77461B54AA27AC" type="pubDate">
<mods:number id="C9F19DF853A4A36CA9908C80E484B717">2018-04-09</mods:number>
</mods:detail>
<mods:detail id="BA509B7FED70921DF41AAF0E19EF5DBB" type="volume">
<mods:number id="12635A3590B901F5E2F08777EAA29B0B">4407</mods:number>
</mods:detail>
<mods:detail id="AB22EA15150A08962B24D7568173A6B7" type="issue">
<mods:number id="B4D03A5F8A0EA35C8AAE5DC5FEF80443">1</mods:number>
</mods:detail>
<mods:extent id="F2C7D4324D3F001216E03142FE5BD400" unit="page">
<mods:start id="8116DDBF8037724B25FCBA814234DA60">29</mods:start>
<mods:end id="F5BB9BE076295C63A93CD134DC5D94EA">50</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="509CD034521555A74234E7E8BB2B9A55">journal article</mods:classification>
<mods:identifier id="B69941FFDAB8E1323B5AE872C577F180" type="CLB-Dataset">22806</mods:identifier>
<mods:identifier id="654E786C7C471B65D1175E9A96530159" type="DOI">10.11646/zootaxa.4407.1.2</mods:identifier>
<mods:identifier id="A6622D048B7FDE27E5B9D2FB786D39E8" type="GBIF-Dataset">8e0aac3a-6556-4b96-a67e-fce9f0db0ba9</mods:identifier>
<mods:identifier id="CA76455F2094FE3E501DC44678709D02" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5D1C33774BFE6E7171F1815B44EADB6F" type="Zenodo-Dep">3766290</mods:identifier>
<mods:identifier id="D4607412FA4A474DA7DC42E8197AD17C" type="ZooBank">B45E9ABD-CB52-4EE7-91D6-783E8E958AC5</mods:identifier>
</mods:mods>
<treatment id="676A87E0FFB4B15B2392FA45469D0F2E" ID-DOI="http://doi.org/10.5281/zenodo.3803378" ID-GBIF-Taxon="163743971" ID-Zenodo-Dep="3803378" LSID="urn:lsid:zoobank.org:act:68539169-5FAC-43B9-8839-69141BDB155E" httpUri="http://treatment.plazi.org/id/676A87E0FFB4B15B2392FA45469D0F2E" lastPageId="17" lastPageNumber="46" pageId="16" pageNumber="45">
<subSubSection id="A7D9657DFFB4B15A2392FA45408F0B8A" box="[151,643,1446,1472]" pageId="16" pageNumber="45" type="nomenclature">
<paragraph id="EF7C36F6FFB4B15A2392FA45408F0B8A" blockId="16.[151,643,1446,1506]" box="[151,643,1446,1472]" pageId="16" pageNumber="45">
<heading id="B434819AFFB4B15A2392FA45408F0B8A" bold="true" box="[151,643,1446,1472]" fontSize="11" level="1" pageId="16" pageNumber="45" reason="1">
<emphasis id="DDB7EAE4FFB4B15A2392FA45408F0B8A" bold="true" box="[151,643,1446,1472]" pageId="16" pageNumber="45">
<taxonomicName id="28C34D75FFB4B15A2392FA4540350B8A" ID-CoL="8XQKL" authority="Short &amp; Girón, 2018" authorityName="Short &amp; Girón" authorityYear="2018" box="[151,569,1446,1472]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov." subGenus="Hydrobaticus">
<emphasis id="DDB7EAE4FFB4B15A2392FA4540350B8A" bold="true" box="[151,569,1446,1472]" italics="true" pageId="16" pageNumber="45">Helochares (Hydrobaticus) trujillo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C684579FFFB4B15A2145FA45408F0B8A" box="[576,643,1446,1472]" pageId="16" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB4B15A2392FA2A43C90BA8" box="[151,453,1481,1506]" pageId="16" pageNumber="45" type="description">
<paragraph id="EF7C36F6FFB4B15A2392FA2A43C90BA8" blockId="16.[151,643,1446,1506]" box="[151,453,1481,1506]" pageId="16" pageNumber="45">
<figureCitation id="77F82A73FFB4B15A2392FA2A42F50BA8" box="[151,249,1481,1506]" captionStart="FIGURE 2" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[151,1436,265,1900]" captionTargetId="figure@6.[151,1436,265,1900]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. championi Sharp, (C) H. nexus n. sp., (D) H. zamora n. sp., (E) H. sp. A." figureDoi="http://doi.org/10.5281/zenodo.3766294" httpUri="https://zenodo.org/record/3766294/files/figure.png" pageId="16" pageNumber="45">Figs. 2A</figureCitation>
,
<figureCitation id="77F82A73FFB4B15A2200FA2A432F0BA8" box="[261,291,1481,1506]" captionStart="FIGURE 5" captionStartId="11.[151,250,1936,1958]" captionTargetBox="[151,1436,493,1914]" captionTargetId="figure@11.[151,1436,493,1914]" captionTargetPageId="11" captionText="FIGURE 5. Heads of Helochares (Hydrobaticus) spp. (A) H. maculicollis (Pennsylvania), (B) H. normatus (California), (C) H. normatus (Arizona), (D) H. nexus n. sp., (E) H. zamora n. sp., (F) H. trujillo n. sp., (G) H. championi, (H) H. sp. A., (I) H. politus n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766300" httpUri="https://zenodo.org/record/3766300/files/figure.png" pageId="16" pageNumber="45">5F</figureCitation>
,
<figureCitation id="77F82A73FFB4B15A222BFA2943580BA8" box="[302,340,1482,1506]" captionStart="FIGURE 7" captionStartId="13.[151,250,1200,1222]" captionTargetBox="[347,1229,206,1166]" captionTargetId="figure@13.[340,1247,193,1179]" captionTargetPageId="13" captionText="FIGURE 7. Aedeagi of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. zamora n. sp., (C) H. maculicollis (Florida), (D) H. maculicollis (Kansas), (E) H. maculicollis (Pennsylvania), (F) H. nexus n. sp. (Venezuela), (G) H. nexus n. sp. (Panama), (H) H. nexus n. sp. (Ecuador), (I) H. politus n. sp., (J) H. laevis n. sp., (K) H. championi." figureDoi="http://doi.org/10.5281/zenodo.3766304" httpUri="https://zenodo.org/record/3766304/files/figure.png" pageId="16" pageNumber="45">7A</figureCitation>
,
<figureCitation id="77F82A73FFB4B15A225AFA2943880BA8" box="[351,388,1482,1506]" captionStart="FIGURE 9" captionStartId="17.[151,250,1412,1434]" captionTargetBox="[151,1436,411,1391]" captionTargetId="figure@17.[151,1436,411,1391]" captionTargetPageId="17" captionText="FIGURE 9. Habitat of Helochares (Hydrobaticus) spp. (A) Habitat of H. trujillo n. sp. in Venezuela [collecting event VZ09- 0121-04X], (B) Habitat of H. normatus in California, (C) Habitat of H. nexus n. sp. in Venezuela [collecting event VZ09-0129- 01A], (D) Habitat of H. maculicollis in Kansas (USA)." figureDoi="http://doi.org/10.5281/zenodo.3766308" httpUri="https://zenodo.org/record/3766308/files/figure.png" pageId="16" pageNumber="45">9A</figureCitation>
,
<figureCitation id="77F82A73FFB4B15A228AFA2943CD0BA8" box="[399,449,1482,1506]" captionStart="FIGURE 10" captionStartId="18.[151,250,1999,2021]" captionTargetBox="[151,1436,373,1976]" captionTargetId="figure@18.[151,1436,373,1977]" captionTargetPageId="18" captionText="FIGURE 10. Distribution of New World Helochares (Hydrobaticus) spp." figureDoi="http://doi.org/10.5281/zenodo.3766310" httpUri="https://zenodo.org/record/3766310/files/figure.png" pageId="16" pageNumber="45">10B</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB4B15A2392F9F240C70894" pageId="16" pageNumber="45" type="materials_examined">
<paragraph id="EF7C36F6FFB4B15A2392F9F240C70894" blockId="16.[151,1437,1553,2010]" pageId="16" pageNumber="45">
<emphasis id="DDB7EAE4FFB4B15A2392F9F243460860" bold="true" box="[151,330,1553,1578]" pageId="16" pageNumber="45">Type material.</emphasis>
<materialsCitation id="5FAB3CABFFB4B15A2253F9F2413E0804" ID-GBIF-Occurrence="2608702056" collectingDate="2006-01-14" collectionCode="MIZA" collectorName="A. E. Z. Short" country="Venezuela" location="Monte Zerpa Area" pageId="16" pageNumber="45" specimenCount="1" stateProvince="Merida" typeStatus="holotype">
<emphasis id="DDB7EAE4FFB4B15A2253F9F240290860" bold="true" box="[342,549,1553,1578]" pageId="16" pageNumber="45">
<typeStatus id="30788854FFB4B15A2253F9F243CF0860" box="[342,451,1553,1578]" pageId="16" pageNumber="45" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
“VENEZUELA:
<collectingRegion id="2D07F814FFB4B15A21FCF9F241810860" box="[761,909,1553,1578]" country="Venezuela" name="Merida" pageId="16" pageNumber="45">Merida State</collectingRegion>
/
<collectingRegion id="2D07F814FFB4B15A20A7F9F241FA0860" box="[930,1014,1553,1578]" country="Venezuela" name="Merida" pageId="16" pageNumber="45">Merida</collectingRegion>
,
<location id="EA1C602DFFB4B15A2703F9F146D30860" LSID="urn:lsid:plazi:treatment:676A87E0FFB4B15B2392FA45469D0F2E:EA1C602DFFB4B15A2703F9F146D30860" box="[1030,1247,1554,1578]" country="Venezuela" name="Monte Zerpa Area" pageId="16" pageNumber="45" stateProvince="Merida">Monte Zerpa Area</location>
/ small roadside stream;
<date id="9B7D1036FFB4B15A23F4F9D643560804" box="[241,346,1589,1614]" pageId="16" pageNumber="45" value="2006-01-14">
<collectingDate id="8B39E9DEFFB4B15A23F4F9D643560804" box="[241,346,1589,1614]" pageId="16" pageNumber="45" value="2006-01-14">14.i.2006</collectingDate>
</date>
/ AS-06-027/ leg.
<collectorName id="42365320FFB4B15A212CF9D540B20804" box="[553,702,1589,1614]" pageId="16" pageNumber="45">A.E.Z. Short</collectorName>
”. (
<collectionCode id="89D2AE33FFB4B15A21D9F9D541260804" box="[732,810,1590,1614]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="16" pageNumber="45" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="5FAB3CABFFB4B15A2038F9D642A2083B" ID-GBIF-Occurrence="2608702068" collectorName="Same" country="Venezuela" location="Merida" pageId="16" pageNumber="45" specimenCount="1" stateProvince="Merida" typeStatus="paratype">
<emphasis id="DDB7EAE4FFB4B15A2038F9D647130804" bold="true" box="[829,1311,1589,1614]" pageId="16" pageNumber="45">
<typeStatus id="30788854FFB4B15A2038F9D641B90804" box="[829,949,1589,1614]" pageId="16" pageNumber="45" type="paratype">Paratypes</typeStatus>
(10):
<collectingCountry id="97D47666FFB4B15A20F8F9D646A30804" box="[1021,1199,1589,1614]" name="Venezuela" pageId="16" pageNumber="45">VENEZUELA</collectingCountry>
:
<collectingRegion id="2D07F814FFB4B15A27B8F9D647160804" box="[1213,1306,1589,1614]" country="Venezuela" name="Merida" pageId="16" pageNumber="45">Merida</collectingRegion>
:
</emphasis>
<collectorName id="42365320FFB4B15A2622F9D547690804" box="[1319,1381,1590,1614]" pageId="16" pageNumber="45">Same</collectorName>
data as
</materialsCitation>
<materialsCitation id="5FAB3CABFFB4B15A23BEF9BA47950838" ID-GBIF-Occurrence="2608702100" box="[187,1433,1625,1650]" collectionCode="SEMC, MIZA" country="Venezuela" latitude="8.63025" longLatPrecision="1" longitude="-71.1611" pageId="16" pageNumber="45" specimenCount="1" stateProvince="Trujillo" typeStatus="holotype">
<typeStatus id="30788854FFB4B15A23BEF9BA43110838" box="[187,285,1625,1650]" pageId="16" pageNumber="45" type="holotype">holotype</typeStatus>
[
<geoCoordinate id="8AF75031FFB4B15A2235F9B943B30838" box="[304,447,1626,1650]" degrees="8" direction="north" minutes="37" orientation="latitude" pageId="16" pageNumber="45" precision="1" seconds="48.9" value="8.63025">8°37'48.9&quot;N</geoCoordinate>
,
<geoCoordinate id="8AF75031FFB4B15A22D5F9BA40820838" box="[464,654,1625,1650]" degrees="71" direction="west" minutes="9" orientation="longitude" pageId="16" pageNumber="45" precision="1" seconds="39.9666" value="-71.1611">71°9'39.9666&quot;W</geoCoordinate>
] (5,
<collectionCode id="89D2AE33FFB4B15A21C9F9B941170838" box="[716,795,1626,1650]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="45" type="Museum">SEMC</collectionCode>
,
<collectionCode id="89D2AE33FFB4B15A2029F9B9417A0838" box="[812,886,1626,1650]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="16" pageNumber="45" type="Museum">MIZA</collectionCode>
, MALUZ, including DNA voucher SLE1192)
</materialsCitation>
.
<materialsCitation id="5FAB3CABFFB4B15A2392F99E40C40894" ID-GBIF-Occurrence="2608702034" collectingDate="2009-01-21" collectingDateMax="2009-07-13" collectingDateMin="2009-01-21" collectionCode="SEMC, MIZA" collectorName="Short &amp; Garcia &amp; Camacho, VZ" country="Venezuela" latitude="9.306217" location="Laguna Agua Negra" longLatPrecision="1" longitude="-70.17527" pageId="16" pageNumber="45" specimenCount="1" stateProvince="Trujillo" typeStatus="paratype">
<emphasis id="DDB7EAE4FFB4B15A2392F99E42F708DC" bold="true" box="[151,251,1661,1686]" pageId="16" pageNumber="45">
<collectingRegion id="2D07F814FFB4B15A2392F99E42FB08DC" box="[151,247,1661,1686]" country="Venezuela" name="Trujillo" pageId="16" pageNumber="45">Trujillo</collectingRegion>
:
</emphasis>
<location id="EA1C602DFFB4B15A2204F99D43EA08DC" LSID="urn:lsid:plazi:treatment:676A87E0FFB4B15B2392FA45469D0F2E:EA1C602DFFB4B15A2204F99D43EA08DC" box="[257,486,1662,1686]" country="Venezuela" latitude="9.306217" longLatPrecision="1" longitude="-70.17527" name="Laguna Agua Negra" pageId="16" pageNumber="45" stateProvince="Trujillo">Laguna Agua Negra</location>
,
<geoCoordinate id="8AF75031FFB4B15A22F5F99D40A308DC" box="[496,687,1662,1686]" degrees="9" direction="north" minutes="18" orientation="latitude" pageId="16" pageNumber="45" precision="1" seconds="22.3812" value="9.306217">9°18' 22.3812&quot;N</geoCoordinate>
,
<geoCoordinate id="8AF75031FFB4B15A21BCF99D418E08DC" box="[697,898,1662,1686]" degrees="70" direction="west" minutes="10" orientation="longitude" pageId="16" pageNumber="45" precision="1" seconds="30.9714" value="-70.17527">70°10'30.9714&quot;W</geoCoordinate>
,
<date id="9B7D1036FFB4B15A2088F99E41F508DC" box="[909,1017,1661,1686]" pageId="16" pageNumber="45" value="2009-01-21">
<collectingDate id="8B39E9DEFFB4B15A2088F99E41F508DC" box="[909,1017,1661,1686]" pageId="16" pageNumber="45" value="2009-01-21">21.i.2009</collectingDate>
</date>
, margin of lagoon, leg.
<collectorName id="42365320FFB4B15A2605F99E473308DC" box="[1280,1343,1661,1686]" pageId="16" pageNumber="45">Short</collectorName>
,
<collectorName id="42365320FFB4B15A264FF99D479408DC" box="[1354,1432,1662,1686]" pageId="16" pageNumber="45">García</collectorName>
, &amp;
<collectorName id="42365320FFB4B15A23B2F942435608F0" box="[183,346,1697,1722]" pageId="16" pageNumber="45">Camacho, VZ</collectorName>
09-0121-04X (2,
<collectionCode id="89D2AE33FFB4B15A2126F941407808F0" box="[547,628,1698,1722]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="45" type="Museum">SEMC</collectionCode>
); same locality but
<date id="9B7D1036FFB4B15A2058F94241D208F0" box="[861,990,1697,1722]" pageId="16" pageNumber="45" value="2009-07-13">
<collectingDate id="8B39E9DEFFB4B15A2058F94241D208F0" box="[861,990,1697,1722]" pageId="16" pageNumber="45" value="2009-07-13">13.vii.2009</collectingDate>
</date>
, leg.
<collectorName id="42365320FFB4B15A2727F942465308F0" box="[1058,1119,1697,1722]" pageId="16" pageNumber="45">Short</collectorName>
<emphasis id="DDB7EAE4FFB4B15A276CF94146A908F3" box="[1129,1189,1697,1721]" italics="true" pageId="16" pageNumber="45">et al.</emphasis>
, VZ09-0713-03A (3,
<collectionCode id="89D2AE33FFB4B15A2392F92542EA0894" box="[151,230,1734,1758]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="45" type="Museum">SEMC</collectionCode>
,
<collectionCode id="89D2AE33FFB4B15A23F4F92543300894" box="[241,316,1734,1758]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="16" pageNumber="45" type="Museum">MIZA</collectionCode>
; includes DNA voucher SLE0034)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB4B15A23C2F90A47810990" pageId="16" pageNumber="45" type="diagnosis">
<paragraph id="EF7C36F6FFB4B15A23C2F90A47810990" blockId="16.[151,1437,1553,2010]" pageId="16" pageNumber="45">
<emphasis id="DDB7EAE4FFB4B15A23C2F90A43C20948" bold="true" box="[199,462,1769,1794]" pageId="16" pageNumber="45">Differential diagnosis.</emphasis>
Body size
<quantity id="283B9B13FFB4B15A2149F90A40DA094B" box="[588,726,1769,1794]" metricMagnitude="-3" metricUnit="m" metricValue="6.199999999999999" metricValueMax="6.6" metricValueMin="5.8" pageId="16" pageNumber="45" unit="mm" value="6.199999999999999" valueMax="6.6" valueMin="5.8">5.8–6.6 mm</quantity>
. Clypeus pale yellow to light brown in color, slightly darkened posteromedially (
<figureCitation id="77F82A73FFB4B15A2259F8EE43BE096C" box="[348,434,1805,1830]" captionStart="FIGURE 5" captionStartId="11.[151,250,1936,1958]" captionTargetBox="[151,1436,493,1914]" captionTargetId="figure@11.[151,1436,493,1914]" captionTargetPageId="11" captionText="FIGURE 5. Heads of Helochares (Hydrobaticus) spp. (A) H. maculicollis (Pennsylvania), (B) H. normatus (California), (C) H. normatus (Arizona), (D) H. nexus n. sp., (E) H. zamora n. sp., (F) H. trujillo n. sp., (G) H. championi, (H) H. sp. A., (I) H. politus n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766300" httpUri="https://zenodo.org/record/3766300/files/figure.png" pageId="16" pageNumber="45">Fig. 5F</figureCitation>
). Elytra with ten distinct rows of serial punctures; rows 1–4 not reaching the base of the elytra; rows 1–2 weakly impressed into shallow striae along most of their length (
<figureCitation id="77F82A73FFB4B15A2746F8D246930900" box="[1091,1183,1841,1866]" captionStart="FIGURE 2" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[151,1436,265,1900]" captionTargetId="figure@6.[151,1436,265,1900]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. championi Sharp, (C) H. nexus n. sp., (D) H. zamora n. sp., (E) H. sp. A." figureDoi="http://doi.org/10.5281/zenodo.3766294" httpUri="https://zenodo.org/record/3766294/files/figure.png" pageId="16" pageNumber="45">Fig. 2A</figureCitation>
). Abdominal ventrites uniformly dark brown; pubescence dense and evenly distributed, the longest hairs shorter than those on the metaventrite. Aedeagus (
<figureCitation id="77F82A73FFB4B15A22AAF89A400409D8" box="[431,520,1913,1938]" captionStart="FIGURE 7" captionStartId="13.[151,250,1200,1222]" captionTargetBox="[347,1229,206,1166]" captionTargetId="figure@13.[340,1247,193,1179]" captionTargetPageId="13" captionText="FIGURE 7. Aedeagi of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. zamora n. sp., (C) H. maculicollis (Florida), (D) H. maculicollis (Kansas), (E) H. maculicollis (Pennsylvania), (F) H. nexus n. sp. (Venezuela), (G) H. nexus n. sp. (Panama), (H) H. nexus n. sp. (Ecuador), (I) H. politus n. sp., (J) H. laevis n. sp., (K) H. championi." figureDoi="http://doi.org/10.5281/zenodo.3766304" httpUri="https://zenodo.org/record/3766304/files/figure.png" pageId="16" pageNumber="45">Fig. 7A</figureCitation>
) nearly 2.4-times longer than its greatest width; apex of parameres truncate, with outer corners forming a rounded right angle; inner corners of apex of parameres forming an acute angle; lateral outer margins of parameres diverging on apical third; median lobe with apex gradually tapering to an acute point.
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB5B15B23C2FF7443D60E9E" pageId="17" pageNumber="46" type="description">
<paragraph id="EF7C36F6FFB5B15B23C2FF7443D60E9E" blockId="17.[151,1436,151,357]" pageId="17" pageNumber="46">
<emphasis id="DDB7EAE4FFB5B15B23C2FF7443550EFA" bold="true" box="[199,345,151,176]" pageId="17" pageNumber="46">Description.</emphasis>
In addition to differential diagnosis: dorsal coloration yellow to medium brown, pronotal disc moderately darker (
<figureCitation id="77F82A73FFB5B15B2276FF5F43C10E9E" box="[371,461,188,213]" captionStart="FIGURE 2" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[151,1436,265,1900]" captionTargetId="figure@6.[151,1436,265,1900]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. championi Sharp, (C) H. nexus n. sp., (D) H. zamora n. sp., (E) H. sp. A." figureDoi="http://doi.org/10.5281/zenodo.3766294" httpUri="https://zenodo.org/record/3766294/files/figure.png" pageId="17" pageNumber="46">Fig. 2A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB5B15B23C2FF3C41680EB2" box="[199,868,223,248]" pageId="17" pageNumber="46" type="distribution">
<paragraph id="EF7C36F6FFB5B15B23C2FF3C41680EB2" blockId="17.[151,1436,151,357]" box="[199,868,223,248]" pageId="17" pageNumber="46">
<emphasis id="DDB7EAE4FFB5B15B23C2FF3C436C0EB2" bold="true" box="[199,352,223,248]" pageId="17" pageNumber="46">Distribution.</emphasis>
<collectingCountry id="97D47666FFB5B15B2262FF3C43D60EB2" box="[359,474,223,248]" name="Venezuela" pageId="17" pageNumber="46">Venezuela</collectingCountry>
(
<collectingRegion id="2D07F814FFB5B15B22EDFF3C40370EB2" box="[488,571,223,248]" country="Venezuela" name="Merida" pageId="17" pageNumber="46">Merida</collectingRegion>
Andes region) (
<figureCitation id="77F82A73FFB5B15B21F4FF3C415B0EB2" box="[753,855,223,248]" captionStart="FIGURE 10" captionStartId="18.[151,250,1999,2021]" captionTargetBox="[151,1436,373,1976]" captionTargetId="figure@18.[151,1436,373,1977]" captionTargetPageId="18" captionText="FIGURE 10. Distribution of New World Helochares (Hydrobaticus) spp." figureDoi="http://doi.org/10.5281/zenodo.3766310" httpUri="https://zenodo.org/record/3766310/files/figure.png" pageId="17" pageNumber="46">Fig. 10B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB5B15B23C2FEE746B50F57" box="[199,1209,260,285]" pageId="17" pageNumber="46" type="etymology">
<paragraph id="EF7C36F6FFB5B15B23C2FEE746B50F57" blockId="17.[151,1436,151,357]" box="[199,1209,260,285]" pageId="17" pageNumber="46">
<emphasis id="DDB7EAE4FFB5B15B23C2FEE743420F57" bold="true" box="[199,334,260,285]" pageId="17" pageNumber="46">Etymology.</emphasis>
Named after one of the Venezuelan states in which this species was collected.
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB5B15B23C2FEC4469D0F2E" pageId="17" pageNumber="46" type="biology_ecology">
<paragraph id="EF7C36F6FFB5B15B23C2FEC4469D0F2E" blockId="17.[151,1436,151,357]" pageId="17" pageNumber="46">
<emphasis id="DDB7EAE4FFB5B15B23C2FEC4432B0F0A" bold="true" box="[199,295,295,320]" pageId="17" pageNumber="46">Biology.</emphasis>
All three collections of this species were made in densely vegetated, muddy margins of creeks and lagoons at high elevations (&gt;
<quantity id="283B9B13FFB5B15B22DCFEAF40380F2E" box="[473,564,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="46" unit="m" value="1800.0">1800 m</quantity>
). This species has been observed to carry its egg case.
</paragraph>
</subSubSection>
</treatment>
</document>
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