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<taxonomicName id="28AA4D0B4D160445F741B0CB6488FE55" authority="Victor, 1838" authorityName="Victor" authorityYear="1838" box="[573,1015,469,496]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F741B0CB6421FE4A" bold="true" box="[573,862,469,496]" italics="true" pageId="19" pageNumber="20">Cholovocera formicaria</emphasis>
<bibRefCitation id="8B3B4B794D160445F619B0C86488FE55" author="Victor T." box="[869,1015,470,496]" pageId="19" pageNumber="20" pagination="175 - 188" refId="ref42813" refString="Victor T. 1838. Coleopteres du Caucase et des Provinces transcaucasiennes recueillis et decrits. Bulletin de la Societe imperiale des Naturalistes de Moscou 11 (2): 175 - 188." type="journal article" year="1838">Victor, 1838</bibRefCitation>
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<figureCitation id="77912A0D4D160445F4FAB0E4669BFDB1" box="[390,484,506,532]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[264,1318,1324,1816]" captionTargetId="figure-432@10.[187,1400,1277,1877]" captionTargetPageId="10" captionText="Fig. 4. Prosternal processes. A. Cholovocera afghana Johnson, 1977. B. Ch. attae (Kraatz, 1858). C. Ch. balcánica (Karaman, 1936). D. Ch. gallica (Schaufuss, 1876). E. Ch. formicaria Victor, 1838. F. Ch. formiceticola (Rosenhauer, 1856). G. Ch. occulta sp. nov. H. Ch. punctata (Märkel, 1845)." figureDoi="http://doi.org/10.5281/zenodo.10424520" httpUri="https://zenodo.org/record/10424520/files/figure.png" pageId="19" pageNumber="20">Figs 4E</figureCitation>
,
<figureCitation id="77912A0D4D160445F48CB0E5656DFDB0" box="[496,530,507,533]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[301,1319,679,1805]" captionTargetId="figure-184@13.[265,1322,663,1845]" captionTargetPageId="13" captionText="Fig. 7. Spermathecae, lateral view. A. Cholovocera afghana Johnson, 1977. B. Ch. balcánica (Karaman, 1936). C. Ch. occulta sp. nov. D. Ch. gallica (Schaufuss, 1876). E. Ch. formicaria Victor, 1838. F. Ch. formiceticola (Rosenhauer, 1856). G. Ch. punctata (Märkel, 1845). Abbreviations: See Material and methods. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424528" httpUri="https://zenodo.org/record/10424528/files/figure.png" pageId="19" pageNumber="20">7E</figureCitation>
,
<figureCitation id="77912A0D4D160445F763B0E4653CFDB1" box="[543,579,506,532]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[303,1293,291,1782]" captionTargetId="figure-70@18.[247,1340,265,1810]" captionTargetPageId="18" captionText="Fig. 9. Known geographical distribution of the species ofCholovocera Victor, 1838.A. Ch. formiceticola (Rosenhauer, 1856) (red dots) and Ch. gallica (Schaufuss, 1876) (yellow dots). B. Ch. punctata (Märkel, 1845) (red dots) and Ch. occulta sp. nov. (yellow dots). C. Ch. balcanica (Karaman, 1936) (red dots), Ch. attae (Kraatz, 1858) (yellow dots), Ch. formicaria (dark blue dots) and Ch. afghana Johnson, 1977 (orange dot)." figureDoi="http://doi.org/10.5281/zenodo.10424533" httpUri="https://zenodo.org/record/10424533/files/figure.png" pageId="19" pageNumber="20">9C</figureCitation>
,
<figureCitation id="77912A0D4D160445F72CB0E465FCFDB1" box="[592,643,506,532]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[321,1272,530,1820]" captionTargetId="figure-115@41.[311,1276,513,1878]" captionTargetPageId="41" captionText="Fig. 11. Male habitus, dorsal view. A. Cholovocera formicaria Victor, 1838. B. Ch. formiceticola (Rosenhauer, 1856). C. Ch. occulta sp. nov., paratype. D. Ch. punctata (Märkel, 1845). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10424537" httpUri="https://zenodo.org/record/10424537/files/figure.png" pageId="19" pageNumber="20">11A</figureCitation>
,
<figureCitation id="77912A0D4D160445F7ECB0E465BDFDB1" box="[656,706,506,532]" captionStart="Fig" captionStartId="42.[189,232,1892,1918]" captionTargetBox="[341,1310,800,1791]" captionTargetId="figure-179@42.[221,1367,721,1840]" captionTargetPageId="42" captionText="Fig. 12. Terminal female antennomeres. A. Cholovocera afghana Johnson, 1977. B. Ch. attae (Kraatz, 1858). C. Ch. balcánica (Karaman, 1936). D. Ch. gallica (Schaufuss, 1876). E. Ch. formicaria Victor, 1838. F. Ch. formiceticola (Rosenhauer, 1856). G. Ch. occulta sp. nov. H. Ch. punctata (Märkel, 1845). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424539" httpUri="https://zenodo.org/record/10424539/files/figure.png" pageId="19" pageNumber="20">12E</figureCitation>
,
<figureCitation id="77912A0D4D160445F7B2B0E4647CFDB1" box="[718,771,506,532]" captionStart="Fig" captionStartId="44.[189,232,1928,1954]" captionTargetBox="[247,1330,760,1834]" captionTargetId="figure-280@44.[217,1371,719,1875]" captionTargetPageId="44" captionText="Fig. 13. Terminal male antennomeres.A. Cholovocera attae (Kraatz, 1858). B. Ch. balcánica(Karaman, 1936). C. Ch. gallica (Schaufuss, 1876). D. Ch. formicaria Victor, 1838. E. Ch. formiceticola (Rosenhauer, 1856). F. Ch. occulta sp. nov. G. Ch. punctata (Märkel, 1845). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424541" httpUri="https://zenodo.org/record/10424541/files/figure.png" pageId="19" pageNumber="20">13D</figureCitation>
,
<figureCitation id="77912A0D4D160445F673B0E46422FDB1" box="[783,861,506,532]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[293,1322,589,1725]" captionTargetId="figure-210@46.[250,1337,543,1774]" captionTargetPageId="46" captionText="Fig. 14. Metatibiae. A. Cholovocera afghana Johnson, 1977, ♀. B. Ch. attae (Kraatz, 1858), ♀. C. Ch. attae (Kraatz, 1858), ♂. D. Ch. balcanica (Karaman, 1936), ♀. E. Ch. balcanica (Karaman, 1936), ♂. F. Ch. gallica (Schaufuss, 1876), ♀. G. Ch. gallica (Schaufuss, 1876), ♂. H. Ch. formicaria Victor, 1838, ♀. I. Ch. formicaria Victor, 1838, ♂. J. Ch. formiceticola (Rosenhauer, 1856), ♀. K. Ch. formiceticola (Rosenhauer, 1856), ♂. L. Ch. occulta sp. nov., ♀. M. Ch. occulta sp. nov., ♂. N. Ch. punctata (Märkel, 1845), ♀. O. Ch. punctata (Märkel, 1845), ♂. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424543" httpUri="https://zenodo.org/record/10424543/files/figure.png" pageId="19" pageNumber="20">14HI</figureCitation>
,
<figureCitation id="77912A0D4D160445F615B0E464F7FDB0" box="[873,904,506,533]" captionStart="Fig" captionStartId="53.[189,232,1056,1082]" captionTargetBox="[456,1134,285,992]" captionTargetId="figure-59@53.[422,1166,272,1017]" captionTargetPageId="53" captionText="Fig. 15. Aedeagus and paramere of Cholovocera formicaria Victor, 1838. Male from type locality (Derbent). AB. Ventral views. CD. Lateral views. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10424545" httpUri="https://zenodo.org/record/10424545/files/figure.png" pageId="19" pageNumber="20">15</figureCitation>
,
<figureCitation id="77912A0D4D160445F6E9B0E46493FDB1" box="[917,1004,506,532]" captionStart="Fig" captionStartId="59.[189,232,1716,1742]" captionTargetBox="[256,1301,225,1672]" captionTargetId="figure-138@59.[256,1301,217,1680]" captionTargetPageId="59" captionText="Fig. 23. Types and relevant specimens of species of Cholovocera Victor, 1838. A. Syntypes of Ch. formicaria Victor, 1838 on original Motchoulskys mounting card (courtesy of A.A. Gusakov). B. Male specimen of Ch. formicaria Victor, 1838 from the type locality. C. Lectotype of Ch. punctata (Märkel, 1845) on left, and a specimen from the J.W. Helfers Collection (NMPC) on right, showing identical pins and mounting cards. D. Single label attached to five females of Ch. punctata (Märkel, 1845) from the Helfer Collection. E. Label of Ch. punctata (Märkel, 1845) lectotype. F. Lectotype (above) and paralectotype (below) of Ch. formiceticola (Rosenhauer, 1856). G. Label of lectotype of Ch. formiceticola (Rosenhauer, 1856). H. Five syntypes of Ch. balcanica (Karaman, 1936), from left: second, third and fourth pins (courtesy of V. Krpach)." figureDoi="http://doi.org/10.5281/zenodo.10424571" httpUri="https://zenodo.org/record/10424571/files/figure.png" pageId="19" pageNumber="20">23AB</figureCitation>
,
<figureCitation id="77912A0D4D160445F684B0E463D2FDB1" box="[1016,1197,506,532]" captionStart="Fig" captionStartId="60.[189,232,1787,1813]" captionTargetBox="[257,1331,228,1700]" captionTargetId="figure-1@60.[257,1331,225,1744]" captionTargetPageId="60" captionText="Fig. 24. A. A syntype of Cholovocera formicaria Victor, 1838, dorsal view (courtesy of K.V. Makarov). B. Same specimen, ventral view (courtesy of K.V. Makarov). C. Holotype of Ch. brevicornis Johnson, 1977 (= Displotera beloni (Wasmann, 1899)), dorsal view. D. Same specimen, ocular area, lateral view. E. Ocular area of Ch. formiceticola (Rosenhauer, 1856) (Lisbon), lateral view. F. Aedeagus of Ch. formicaria major, lectotype, ventral view. G. Same specimen, lateral view. H. Aedeagus of Co punctata sardoa, lectotype, ventral view. I. Same specimen, lateral view. Scale bars: AC = 0.5 mm; DI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424576" httpUri="https://zenodo.org/record/10424576/files/figure.png" pageId="19" pageNumber="20">24AB, 24FG</figureCitation>
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<treatmentCitation id="6E0B10994D160445F5C1B32265CFFDF3" author="Victor T." box="[189,688,572,598]" page="179" pageId="19" pageNumber="20" year="1838">
<taxonomicName id="28AA4D0B4D160445F5C1B32265CFFDF3" authority="Victor, 1838: 179" authorityName="Victor" authorityPageNumber="179" authorityYear="1838" box="[189,688,572,598]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F5C1B32266A6FDF3" box="[189,473,572,598]" italics="true" pageId="19" pageNumber="20">Cholovocera formicaria</emphasis>
<bibRefCitation id="8B3B4B794D160445F49CB32265CFFDF3" author="Victor T." box="[480,688,572,598]" pageId="19" pageNumber="20" pagination="175 - 188" refId="ref42813" refString="Victor T. 1838. Coleopteres du Caucase et des Provinces transcaucasiennes recueillis et decrits. Bulletin de la Societe imperiale des Naturalistes de Moscou 11 (2): 175 - 188." type="journal article" year="1838">Victor, 1838: 179</bibRefCitation>
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, pl. III.
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<taxonomicName id="28AA4D0B4D160445F5C1B36166B5FD3C" authorityName="Motschulsky" authorityYear="1838" box="[189,458,639,665]" class="Insecta" family="Latridiidae" genus="Colovocera" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F5C1B36166B5FD3C" box="[189,458,639,665]" italics="true" pageId="19" pageNumber="20">Colovocera formicaria</emphasis>
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<treatmentCitation id="6E0B10994D160445F494B36165D0FD3C" author="Belon M. - J." box="[488,687,639,665]" page="194" pageId="19" pageNumber="20" year="1879">
<bibRefCitation id="8B3B4B794D160445F494B36165D0FD3C" author="Belon M. - J." box="[488,687,639,665]" pageId="19" pageNumber="20" pagination="157 - 365" refId="ref38155" refString="Belon M. - J. 1879. Famille des Lathridiens, 1 re partie. In: Mulsant E. (ed.) Histoire naturelle des coleopteres de France. Annales de la Societe linneenne de Lyon, n. s. 26: 157 - 365." type="book chapter" year="1879">Belon 1879: 194</bibRefCitation>
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.
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<taxonomicName id="28AA4D0B4D160445F5C1B3BD6695FD18" authorityName="Motchoulsky" authorityYear="1845" box="[189,490,675,701]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="DDDEEA9A4D160445F5C1B3BD6695FD18" box="[189,490,675,701]" italics="true" pageId="19" pageNumber="20">Cholovocera subterranea</emphasis>
</taxonomicName>
<treatmentCitation id="6E0B10994D160445F77BB3BD645FFD18" author="Motchoulsky V. de" box="[519,800,675,702]" page="111" pageId="19" pageNumber="20" year="1845">
<bibRefCitation id="8B3B4B794D160445F77BB3BD645FFD18" author="Motchoulsky V. de" box="[519,800,675,702]" pageId="19" pageNumber="20" pagination="3 - 127" refId="ref40872" refString="Motchoulsky V. de 1845. Remarques sur la collection de coleopteres russes. Bulletin de la Societe imperiale des Naturalistes de Moscou 18 (1): 3 - 127." type="journal article" year="1845">Motchoulsky 1845: 111</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="EF1536884D160445F5C1B3D96591FD44" blockId="19.[189,898,639,810]" box="[189,750,711,737]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="CFBA11A64D160445F5C1B3D96591FD44" box="[189,750,711,737]" pageId="19" pageNumber="20">
<taxonomicName id="28AA4D0B4D160445F5C1B3D966C2FD44" authorityName="Victor" authorityYear="1838" box="[189,445,711,737]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F5C1B3D966C2FD44" box="[189,445,711,737]" italics="true" pageId="19" pageNumber="20">Coluocera formicaria</emphasis>
</taxonomicName>
<treatmentCitation id="6E0B10994D160445F4A7B3D96594FD44" author="Heyden L. von &amp; Reitter E. &amp; Weise J." box="[475,747,711,737]" page="80" pageId="19" pageNumber="20" year="1883">
<bibRefCitation id="8B3B4B794D160445F4A7B3D96594FD44" author="Heyden L. von &amp; Reitter E. &amp; Weise J." box="[475,747,711,737]" pageId="19" pageNumber="20" refId="ref39594" refString="Heyden L. von, Reitter E. &amp; Weise J. 1883. Catalogus coleopterorum Europae et Caucasi. 3 th ed. Libraria Nicolai, Berlin." type="book" year="1883">
Heyden
<emphasis id="DDDEEA9A4D160445F742B3D66507FD44" box="[574,632,711,737]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
1883: 80
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="EF1536884D160445F5C1B3F564FDFCA0" blockId="19.[189,898,639,810]" box="[189,898,747,774]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="CFBA11A64D160445F5C1B3F564FDFCA0" box="[189,898,747,774]" pageId="19" pageNumber="20">
<taxonomicName id="28AA4D0B4D160445F5C1B3F56559FCA0" authorityName="Reitter" authorityYear="1887" box="[189,550,747,774]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="variety" species="formicaria" variety="major">
<emphasis id="DDDEEA9A4D160445F5C1B3F566C2FCA0" box="[189,445,747,773]" italics="true" pageId="19" pageNumber="20">Coluocera formicaria</emphasis>
v.
<emphasis id="DDDEEA9A4D160445F49CB3F26559FCA0" box="[480,550,748,773]" italics="true" pageId="19" pageNumber="20">major</emphasis>
</taxonomicName>
<treatmentCitation id="6E0B10994D160445F738B3F56479FCA0" author="Reitter E." box="[580,774,747,773]" page="10" pageId="19" pageNumber="20" year="1887">
<bibRefCitation id="8B3B4B794D160445F738B3F56479FCA0" author="Reitter E." box="[580,774,747,773]" pageId="19" pageNumber="20" refId="ref41271" refString="Reitter E. 1887. Bestimmungs-Tabellen der europaischen Coleopteren. III Helft. II Auflage. Enthaltend die Familien: Scaphidiidae, Lathridiidae und Dermestidae. E. Reitter, M ˆ dling." type="book" year="1887">Reitter 1887: 10</bibRefCitation>
</treatmentCitation>
.
<emphasis id="DDDEEA9A4D160445F66DB3F564FDFCA0" bold="true" box="[785,898,747,773]" pageId="19" pageNumber="20">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="EF1536884D160445F5C1B211645FFC8C" blockId="19.[189,898,639,810]" box="[189,800,783,810]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="CFBA11A64D160445F5C1B211645FFC8C" box="[189,800,783,810]" pageId="19" pageNumber="20">
<taxonomicName id="28AA4D0B4D160445F5C1B21166DDFC8C" box="[189,418,783,809]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="major">
<emphasis id="DDDEEA9A4D160445F5C1B21166DDFC8C" box="[189,418,783,809]" italics="true" pageId="19" pageNumber="20">Cholovocera major</emphasis>
</taxonomicName>
<treatmentCitation id="6E0B10994D160445F4C3B21165EFFC8C" author="Rucker W. H." box="[447,656,783,809]" page="13" pageId="19" pageNumber="20" year="2011">
<bibRefCitation id="8B3B4B794D160445F4C3B21165EFFC8C" author="Rucker W. H." box="[447,656,783,809]" pageId="19" pageNumber="20" pagination="11 - 20" refId="ref41653" refString="Rucker W. H. 2011 a. Ergebnisse von Typenstudien der Gattungen Metophthalmus Motschulsky, 1850 (Coleoptera: Latridiidae), Cholovocera Motschulsky, 1838, Reitteria Leder, 1872, Merophysia Lucas, 1852 (Endomychidae: Merophysiinae) und Holoparamecus Curtis, 1833 (Endomychidae: Holoparamecinae). Latridiidae - Mitteilungsblatt fur Systematik und Taxonomie der Latridiidae 8: 11 - 20." type="journal article" year="2011">Rücker 2011a: 13</bibRefCitation>
</treatmentCitation>
, figs 1820.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D160445F5C1B24F64C7FBCB" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="EF1536884D160445F5C1B24F66B6FCCE" blockId="19.[189,457,849,875]" box="[189,457,849,875]" pageId="19" pageNumber="20">
<heading id="B45D81E44D160445F5C1B24F66B6FCCE" bold="true" box="[189,457,849,875]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="DDDEEA9A4D160445F5C1B24F66B6FCCE" bold="true" box="[189,457,849,875]" pageId="19" pageNumber="20">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D160445F5C1B29F640FFBA2" blockId="19.[188,1399,897,1031]" pageId="19" pageNumber="20">
<taxonomicName id="28AA4D0B4D160445F5C1B29F66A7FC3E" authorityName="Victor" authorityYear="1838" box="[189,472,897,923]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F5C1B29F66A7FC3E" box="[189,472,897,923]" italics="true" pageId="19" pageNumber="20">Cholovocera formicaria</emphasis>
</taxonomicName>
is morphologically and geographically close to
<taxonomicName id="28AA4D0B4D160445F178B29F63EBFC3E" authorityName="Johnson" authorityYear="1977" box="[1028,1172,897,923]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="afghana">
<emphasis id="DDDEEA9A4D160445F178B29F63EBFC3E" box="[1028,1172,897,923]" italics="true" pageId="19" pageNumber="20">Ch. afghana</emphasis>
</taxonomicName>
and
<taxonomicName id="28AA4D0B4D160445F1B7B29F6211FC3E" box="[1227,1390,897,923]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D160445F1B7B29F6211FC3E" box="[1227,1390,897,923]" italics="true" pageId="19" pageNumber="20">Ch. balcanica</emphasis>
</taxonomicName>
, but it can be distinguished from both species by the shape of the metatibiae. Those of
<taxonomicName id="28AA4D0B4D160445F1E2B2BB6234FC1A" authorityName="Victor" authorityYear="1838" box="[1182,1355,933,959]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F1E2B2BB6234FC1A" box="[1182,1355,933,959]" italics="true" pageId="19" pageNumber="20">Ch. formicaria</emphasis>
</taxonomicName>
are club-shaped and sinuous (
<figureCitation id="77912A0D4D160445F48EB2D76505FC46" box="[498,634,969,995]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[293,1322,589,1725]" captionTargetId="figure-210@46.[250,1337,543,1774]" captionTargetPageId="46" captionText="Fig. 14. Metatibiae. A. Cholovocera afghana Johnson, 1977, ♀. B. Ch. attae (Kraatz, 1858), ♀. C. Ch. attae (Kraatz, 1858), ♂. D. Ch. balcanica (Karaman, 1936), ♀. E. Ch. balcanica (Karaman, 1936), ♂. F. Ch. gallica (Schaufuss, 1876), ♀. G. Ch. gallica (Schaufuss, 1876), ♂. H. Ch. formicaria Victor, 1838, ♀. I. Ch. formicaria Victor, 1838, ♂. J. Ch. formiceticola (Rosenhauer, 1856), ♀. K. Ch. formiceticola (Rosenhauer, 1856), ♂. L. Ch. occulta sp. nov., ♀. M. Ch. occulta sp. nov., ♂. N. Ch. punctata (Märkel, 1845), ♀. O. Ch. punctata (Märkel, 1845), ♂. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424543" httpUri="https://zenodo.org/record/10424543/files/figure.png" pageId="19" pageNumber="20">Fig. 14HI</figureCitation>
), but straight and gradually tapering in
<taxonomicName id="28AA4D0B4D160445F12BB2D76397FC46" authorityName="Johnson" authorityYear="1977" box="[1111,1256,969,995]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="afghana">
<emphasis id="DDDEEA9A4D160445F12BB2D76397FC46" box="[1111,1256,969,995]" italics="true" pageId="19" pageNumber="20">Ch. afghana</emphasis>
</taxonomicName>
(
<figureCitation id="77912A0D4D160445F186B2D76216FC46" box="[1274,1385,969,995]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[293,1322,589,1725]" captionTargetId="figure-210@46.[250,1337,543,1774]" captionTargetPageId="46" captionText="Fig. 14. Metatibiae. A. Cholovocera afghana Johnson, 1977, ♀. B. Ch. attae (Kraatz, 1858), ♀. C. Ch. attae (Kraatz, 1858), ♂. D. Ch. balcanica (Karaman, 1936), ♀. E. Ch. balcanica (Karaman, 1936), ♂. F. Ch. gallica (Schaufuss, 1876), ♀. G. Ch. gallica (Schaufuss, 1876), ♂. H. Ch. formicaria Victor, 1838, ♀. I. Ch. formicaria Victor, 1838, ♂. J. Ch. formiceticola (Rosenhauer, 1856), ♀. K. Ch. formiceticola (Rosenhauer, 1856), ♂. L. Ch. occulta sp. nov., ♀. M. Ch. occulta sp. nov., ♂. N. Ch. punctata (Märkel, 1845), ♀. O. Ch. punctata (Märkel, 1845), ♂. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424543" httpUri="https://zenodo.org/record/10424543/files/figure.png" pageId="19" pageNumber="20">Fig. 14A</figureCitation>
), and much shorter and wide in
<taxonomicName id="28AA4D0B4D160445F75CB2F365BAFBA2" box="[544,709,1005,1031]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D160445F75CB2F365BAFBA2" box="[544,709,1005,1031]" italics="true" pageId="19" pageNumber="20">Ch. balcanica</emphasis>
</taxonomicName>
(
<figureCitation id="77912A0D4D160445F7A9B2F3641BFBA2" box="[725,868,1005,1031]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[293,1322,589,1725]" captionTargetId="figure-210@46.[250,1337,543,1774]" captionTargetPageId="46" captionText="Fig. 14. Metatibiae. A. Cholovocera afghana Johnson, 1977, ♀. B. Ch. attae (Kraatz, 1858), ♀. C. Ch. attae (Kraatz, 1858), ♂. D. Ch. balcanica (Karaman, 1936), ♀. E. Ch. balcanica (Karaman, 1936), ♂. F. Ch. gallica (Schaufuss, 1876), ♀. G. Ch. gallica (Schaufuss, 1876), ♂. H. Ch. formicaria Victor, 1838, ♀. I. Ch. formicaria Victor, 1838, ♂. J. Ch. formiceticola (Rosenhauer, 1856), ♀. K. Ch. formiceticola (Rosenhauer, 1856), ♂. L. Ch. occulta sp. nov., ♀. M. Ch. occulta sp. nov., ♂. N. Ch. punctata (Märkel, 1845), ♀. O. Ch. punctata (Märkel, 1845), ♂. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424543" httpUri="https://zenodo.org/record/10424543/files/figure.png" pageId="19" pageNumber="20">Fig. 14DE</figureCitation>
).
</paragraph>
<paragraph id="EF1536884D160445F5C1B52E64C7FBCB" blockId="19.[189,1396,1072,1135]" pageId="19" pageNumber="20">
Furthermore, the aedeagus and the paramere are useful characters to distinguish
<taxonomicName id="28AA4D0B4D160445F126B52E620BFBEF" authorityName="Victor" authorityYear="1838" box="[1114,1396,1072,1098]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F126B52E620BFBEF" box="[1114,1396,1072,1098]" italics="true" pageId="19" pageNumber="20">Cholovocera formicaria</emphasis>
</taxonomicName>
from
<taxonomicName id="28AA4D0B4D160445F581B54A66DDFBCB" box="[253,418,1108,1134]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D160445F581B54A66DDFBCB" box="[253,418,1108,1134]" italics="true" pageId="19" pageNumber="20">Ch. balcanica</emphasis>
</taxonomicName>
and
<taxonomicName id="28AA4D0B4D160445F4A7B54A6522FBCB" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Schaufuss" baseAuthorityYear="1876" box="[475,605,1108,1134]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="gallica">
<emphasis id="DDDEEA9A4D160445F4A7B54A6522FBCB" box="[475,605,1108,1134]" italics="true" pageId="19" pageNumber="20">Ch. gallica</emphasis>
</taxonomicName>
(
<figureCitation id="77912A0D4D160445F711B54A65BEFBCA" box="[621,705,1108,1135]" captionStart="Fig" captionStartId="53.[189,232,1056,1082]" captionTargetBox="[456,1134,285,992]" captionTargetId="figure-59@53.[422,1166,272,1017]" captionTargetPageId="53" captionText="Fig. 15. Aedeagus and paramere of Cholovocera formicaria Victor, 1838. Male from type locality (Derbent). AB. Ventral views. CD. Lateral views. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10424545" httpUri="https://zenodo.org/record/10424545/files/figure.png" pageId="19" pageNumber="20">Fig. 15</figureCitation>
against
<figureCitation id="77912A0D4D160445F65FB54A64D4FBCB" box="[803,939,1108,1134]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="55.[189,232,1963,1989]" captionStartId-1="56.[189,232,1095,1121]" captionTargetBox-0="[485,1124,1125,1899]" captionTargetBox-1="[479,1118,301,1026]" captionTargetId-0="figure-57@55.[426,1183,1077,1935]" captionTargetId-1="figure-60@56.[434,1154,263,1062]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionText-0="Fig. 20. Aedeagus and paramere of Cholovocera gallica (Schaufuss, 1876). Lectotype (Southern France). AB. Ventral views. C. Lateral view. Scale bar = 0.1 mm." captionText-1="Fig. 21. Aedeagus and paramere of Cholovocera balcanica (Karaman, 1936). Male from type locality (Skopje). AB. Ventral views. CD. Lateral views. Scale bar = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10424559" figureDoi-1="http://doi.org/10.5281/zenodo.10424565" httpUri-0="https://zenodo.org/record/10424559/files/figure.png" httpUri-1="https://zenodo.org/record/10424565/files/figure.png" pageId="19" pageNumber="20">Figs 2021</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D160442F5C1B588630BFECF" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="EF1536884D160445F5C1B5886613FB15" blockId="19.[189,364,1174,1200]" box="[189,364,1174,1200]" pageId="19" pageNumber="20">
<heading id="B45D81E44D160445F5C1B5886613FB15" bold="true" box="[189,364,1174,1200]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="DDDEEA9A4D160445F5C1B5886613FB15" bold="true" box="[189,364,1174,1200]" pageId="19" pageNumber="20">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D160445F5C1B5DB64D3F9A5" blockId="19.[185,1399,1221,1537]" pageId="19" pageNumber="20">
<taxonomicName id="28AA4D0B4D160445F5C1B5DB66AAFB45" authorityName="Victor" authorityYear="1838" box="[189,469,1221,1248]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F5C1B5DB66AAFB45" box="[189,469,1221,1248]" italics="true" pageId="19" pageNumber="20">Cholovocera formicaria</emphasis>
</taxonomicName>
:
<specimenCount id="F9ACFD014D160445F49EB5D8651EFB7A" box="[482,609,1221,1248]" count="2" pageId="19" pageNumber="20" type="female">two female</specimenCount>
<typeStatus id="3011882A4D160445F719B5D865B3FB45" box="[613,716,1222,1248]" pageId="19" pageNumber="20" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="28AA4D0B4D160445F794B5DB64ECFB45" authorityName="Victor" authorityYear="1838" box="[744,915,1221,1248]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F794B5DB64ECFB45" box="[744,915,1221,1248]" italics="true" pageId="19" pageNumber="20">Ch. formicaria</emphasis>
</taxonomicName>
(
<figureCitation id="77912A0D4D160445F6DFB5DB636FFB45" box="[931,1040,1221,1248]" captionStart="Fig" captionStartId="59.[189,232,1716,1742]" captionTargetBox="[256,1301,225,1672]" captionTargetId="figure-138@59.[256,1301,217,1680]" captionTargetPageId="59" captionText="Fig. 23. Types and relevant specimens of species of Cholovocera Victor, 1838. A. Syntypes of Ch. formicaria Victor, 1838 on original Motchoulskys mounting card (courtesy of A.A. Gusakov). B. Male specimen of Ch. formicaria Victor, 1838 from the type locality. C. Lectotype of Ch. punctata (Märkel, 1845) on left, and a specimen from the J.W. Helfers Collection (NMPC) on right, showing identical pins and mounting cards. D. Single label attached to five females of Ch. punctata (Märkel, 1845) from the Helfer Collection. E. Label of Ch. punctata (Märkel, 1845) lectotype. F. Lectotype (above) and paralectotype (below) of Ch. formiceticola (Rosenhauer, 1856). G. Label of lectotype of Ch. formiceticola (Rosenhauer, 1856). H. Five syntypes of Ch. balcanica (Karaman, 1936), from left: second, third and fourth pins (courtesy of V. Krpach)." figureDoi="http://doi.org/10.5281/zenodo.10424571" httpUri="https://zenodo.org/record/10424571/files/figure.png" pageId="19" pageNumber="20">Figs23A</figureCitation>
,
<figureCitation id="77912A0D4D160445F164B5D86311FB45" box="[1048,1134,1222,1248]" captionStart="Fig" captionStartId="60.[189,232,1787,1813]" captionTargetBox="[257,1331,228,1700]" captionTargetId="figure-1@60.[257,1331,225,1744]" captionTargetPageId="60" captionText="Fig. 24. A. A syntype of Cholovocera formicaria Victor, 1838, dorsal view (courtesy of K.V. Makarov). B. Same specimen, ventral view (courtesy of K.V. Makarov). C. Holotype of Ch. brevicornis Johnson, 1977 (= Displotera beloni (Wasmann, 1899)), dorsal view. D. Same specimen, ocular area, lateral view. E. Ocular area of Ch. formiceticola (Rosenhauer, 1856) (Lisbon), lateral view. F. Aedeagus of Ch. formicaria major, lectotype, ventral view. G. Same specimen, lateral view. H. Aedeagus of Co punctata sardoa, lectotype, ventral view. I. Same specimen, lateral view. Scale bars: AC = 0.5 mm; DI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424576" httpUri="https://zenodo.org/record/10424576/files/figure.png" pageId="19" pageNumber="20">24AB</figureCitation>
) held in Motschulskys Collection at the University of Moscow (A.A. Gusakov pers. comm.
<date id="9B1410484D160445F6B6B5F4633EFAA1" box="[970,1089,1258,1284]" pageId="19" pageNumber="20" value="2021-05">May 2021</date>
) were not available for our examination. Instead, we have examined a male of
<taxonomicName id="28AA4D0B4D160445F664B41064B8FA8D" authorityName="Victor" authorityYear="1838" box="[792,967,1294,1320]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F664B41064B8FA8D" box="[792,967,1294,1320]" italics="true" pageId="19" pageNumber="20">Ch. formicaria</emphasis>
</taxonomicName>
from “Derbent” (
<figureCitation id="77912A0D4D160445F1E4B410627CFA8D" box="[1176,1283,1294,1320]" captionStart="Fig" captionStartId="59.[189,232,1716,1742]" captionTargetBox="[256,1301,225,1672]" captionTargetId="figure-138@59.[256,1301,217,1680]" captionTargetPageId="59" captionText="Fig. 23. Types and relevant specimens of species of Cholovocera Victor, 1838. A. Syntypes of Ch. formicaria Victor, 1838 on original Motchoulskys mounting card (courtesy of A.A. Gusakov). B. Male specimen of Ch. formicaria Victor, 1838 from the type locality. C. Lectotype of Ch. punctata (Märkel, 1845) on left, and a specimen from the J.W. Helfers Collection (NMPC) on right, showing identical pins and mounting cards. D. Single label attached to five females of Ch. punctata (Märkel, 1845) from the Helfer Collection. E. Label of Ch. punctata (Märkel, 1845) lectotype. F. Lectotype (above) and paralectotype (below) of Ch. formiceticola (Rosenhauer, 1856). G. Label of lectotype of Ch. formiceticola (Rosenhauer, 1856). H. Five syntypes of Ch. balcanica (Karaman, 1936), from left: second, third and fourth pins (courtesy of V. Krpach)." figureDoi="http://doi.org/10.5281/zenodo.10424571" httpUri="https://zenodo.org/record/10424571/files/figure.png" pageId="19" pageNumber="20">Fig. 23B</figureCitation>
), the type locality, held in the Märkel Collection in
<collectionCode id="89BBAE4D4D160445F7E7B42C6593FAE9" box="[667,748,1330,1356]" pageId="19" pageNumber="20">SMTD</collectionCode>
(Dresden), which may have been part of Motschulskys type series. However, considering its different labelling from the
<typeStatus id="3011882A4D160445F6AAB4486342FAD5" box="[982,1085,1366,1392]" pageId="19" pageNumber="20" type="syntype">syntypes</typeStatus>
(
<figureCitation id="77912A0D4D160445F12CB448639CFAD5" box="[1104,1251,1366,1392]" captionStart="Fig" captionStartId="59.[189,232,1716,1742]" captionTargetBox="[256,1301,225,1672]" captionTargetId="figure-138@59.[256,1301,217,1680]" captionTargetPageId="59" captionText="Fig. 23. Types and relevant specimens of species of Cholovocera Victor, 1838. A. Syntypes of Ch. formicaria Victor, 1838 on original Motchoulskys mounting card (courtesy of A.A. Gusakov). B. Male specimen of Ch. formicaria Victor, 1838 from the type locality. C. Lectotype of Ch. punctata (Märkel, 1845) on left, and a specimen from the J.W. Helfers Collection (NMPC) on right, showing identical pins and mounting cards. D. Single label attached to five females of Ch. punctata (Märkel, 1845) from the Helfer Collection. E. Label of Ch. punctata (Märkel, 1845) lectotype. F. Lectotype (above) and paralectotype (below) of Ch. formiceticola (Rosenhauer, 1856). G. Label of lectotype of Ch. formiceticola (Rosenhauer, 1856). H. Five syntypes of Ch. balcanica (Karaman, 1936), from left: second, third and fourth pins (courtesy of V. Krpach)." figureDoi="http://doi.org/10.5281/zenodo.10424571" httpUri="https://zenodo.org/record/10424571/files/figure.png" pageId="19" pageNumber="20">Fig. 23AB</figureCitation>
), we cannot be absolutely sure of its status. Nevertheless, it is an authenticated specimen of
<taxonomicName id="28AA4D0B4D160445F10DB4646260FA31" authorityName="Victor" authorityYear="1838" box="[1137,1311,1402,1428]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F10DB4646260FA31" box="[1137,1311,1402,1428]" italics="true" pageId="19" pageNumber="20">Ch. formicaria</emphasis>
</taxonomicName>
, which allows us to make a comparison between this species and the male
<typeStatus id="3011882A4D160445F6B3B4816355FA1C" box="[975,1066,1439,1465]" pageId="19" pageNumber="20" type="syntype">syntype</typeStatus>
of
<taxonomicName id="28AA4D0B4D160445F12CB4806397FA1D" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[1104,1256,1438,1464]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D160445F12CB4806397FA1D" box="[1104,1256,1438,1464]" italics="true" pageId="19" pageNumber="20">Ch. punctata</emphasis>
</taxonomicName>
to conclude that they are different species.
<bibRefCitation id="8B3B4B794D160445F758B4DC65BCFA79" author="Victor T." box="[548,707,1474,1500]" pageId="19" pageNumber="20" pagination="175 - 188" refId="ref42813" refString="Victor T. 1838. Coleopteres du Caucase et des Provinces transcaucasiennes recueillis et decrits. Bulletin de la Societe imperiale des Naturalistes de Moscou 11 (2): 175 - 188." type="journal article" year="1838">Victor (1838)</bibRefCitation>
also mentioned to have found
<taxonomicName id="28AA4D0B4D160445F151B4DC63A4FA79" authorityName="Victor" authorityYear="1838" box="[1069,1243,1474,1500]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D160445F151B4DC63A4FA79" box="[1069,1243,1474,1500]" italics="true" pageId="19" pageNumber="20">Ch. formicaria</emphasis>
</taxonomicName>
in “Kahétie” (
<collectingCountry id="97BD76184D160445F5B9B4F86656F9A5" box="[197,297,1510,1536]" name="Georgia" pageId="19" pageNumber="20">Georgia</collectingCountry>
), but no specimen with such a label have been located.
</paragraph>
<paragraph id="EF1536884D160445F5C1B7376337F976" blockId="19.[189,1401,1577,1748]" pageId="19" pageNumber="20">
<taxonomicName id="28AA4D0B4D160445F5C1B737653AF9E6" authorityName="Reitter" authorityYear="1887" box="[189,581,1577,1603]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="variety" species="formicaria" variety="major">
<emphasis id="DDDEEA9A4D160445F5C1B73766BFF9E6" box="[189,448,1577,1603]" italics="true" pageId="19" pageNumber="20">Coluocera formicaria</emphasis>
var.
<emphasis id="DDDEEA9A4D160445F483B734653AF9E6" box="[511,581,1578,1603]" italics="true" pageId="19" pageNumber="20">major</emphasis>
</taxonomicName>
:
<typeStatus id="3011882A4D160445F72BB73765BCF9E6" box="[599,707,1577,1603]" pageId="19" pageNumber="20" type="lectotype">lectotype</typeStatus>
(male) and
<specimenCount id="F9ACFD014D160445F62AB737634AF9E6" box="[854,1077,1577,1603]" count="2" pageId="19" pageNumber="20" type="generic" typeStatus="paralectotypes">two paralectotypes</specimenCount>
(male and female) held in
<collectionCode id="89BBAE4D4D160445F5C3B7536663F9C2" box="[191,284,1613,1639]" pageId="19" pageNumber="20">HNHM</collectionCode>
. The type locality given by
<bibRefCitation id="8B3B4B794D160445F70DB7536466F9C2" author="Reitter E." box="[625,793,1613,1639]" pageId="19" pageNumber="20" refId="ref41271" refString="Reitter E. 1887. Bestimmungs-Tabellen der europaischen Coleopteren. III Helft. II Auflage. Enthaltend die Familien: Scaphidiidae, Lathridiidae und Dermestidae. E. Reitter, M ˆ dling." type="book" year="1887">Reitter (1887)</bibRefCitation>
is “Talish Gebiete” [Talish Region]. However, the label attached to the
<typeStatus id="3011882A4D160445F4D3B76F6564F92E" box="[431,539,1649,1675]" pageId="19" pageNumber="20" type="lectotype">lectotype</typeStatus>
reads “Rasano” (see
<bibRefCitation id="8B3B4B794D160445F66FB76F64C6F92E" author="Rucker W. H." box="[787,953,1649,1675]" pageId="19" pageNumber="20" pagination="11 - 20" refId="ref41653" refString="Rucker W. H. 2011 a. Ergebnisse von Typenstudien der Gattungen Metophthalmus Motschulsky, 1850 (Coleoptera: Latridiidae), Cholovocera Motschulsky, 1838, Reitteria Leder, 1872, Merophysia Lucas, 1852 (Endomychidae: Merophysiinae) und Holoparamecus Curtis, 1833 (Endomychidae: Holoparamecinae). Latridiidae - Mitteilungsblatt fur Systematik und Taxonomie der Latridiidae 8: 11 - 20." type="journal article" year="2011">Rücker 2011a</bibRefCitation>
: fig. 18, bottom, left), which is located in the region of the Talish Mountains. According to the ICZN Code (1999), the type locality is what is written on the label associated with the type specimen, in this case “Rasano”.
</paragraph>
<materialsCitation id="5FC23CD54D160445F596B7E563E0F89F" collectionCode="ZMUM" country="Russia" location="Derbent" pageId="19" pageNumber="20" specimenCount="2" specimenCount-female="2" stateProvince="Dagestan" typeStatus="syntype">
<paragraph id="EF1536884D160445F596B7E5656CF8B0" blockId="19.[189,1187,1787,1850]" box="[234,531,1787,1814]" pageId="19" pageNumber="20">
<emphasis id="DDDEEA9A4D160445F596B7E5656CF8B0" bold="true" box="[234,531,1787,1814]" pageId="19" pageNumber="20">
<typeStatus id="3011882A4D160445F596B7E56623F8B0" box="[234,348,1787,1813]" pageId="19" pageNumber="20" type="syntype">Syntypes</typeStatus>
, not examined
</emphasis>
</paragraph>
<paragraph id="EF1536884D160445F5C1B63E63DCF89F" blockId="19.[189,1187,1787,1850]" box="[189,1187,1824,1850]" pageId="19" pageNumber="20">
<collectingCountry id="97BD76184D160445F5C1B63E665BF89F" box="[189,292,1824,1850]" name="Russia" pageId="19" pageNumber="20">RUSSIA</collectingCountry>
<emphasis id="DDDEEA9A4D160445F43DB63E66B9F89F" bold="true" box="[321,454,1824,1850]" pageId="19" pageNumber="20">
<collectingRegion id="2D6EF86A4D160445F43DB63E66CAF89F" box="[321,437,1824,1850]" country="Russia" name="Dagestan" pageId="19" pageNumber="20">Dagestan</collectingRegion>
</emphasis>
<specimenCount id="F9ACFD014D160445F4B2B63E656FF89F" box="[462,528,1824,1850]" count="2" pageId="19" pageNumber="20" type="female">2 ♀♀</specimenCount>
;
<location id="EA7560534D160445F763B63E65FEF89F" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D160445F763B63E65FEF89F" box="[543,641,1824,1850]" country="Russia" name="Derbent" pageId="19" pageNumber="20" stateProvince="Dagestan">Derbent</location>
, “Litt. m. csp.” [Caspian Sea region];
<collectionCode id="89BBAE4D4D160445F13CB63E63E0F89F" box="[1088,1183,1824,1850]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="19" pageNumber="20">ZMUM</collectionCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="5FC23CD54D160445F596B67C65AFF84D" collectionCode="HNHM" collectorName="Leder" country="Azerbaijan" latitude="38.670002" location="Caspi. - M. - Gebiet" longLatPrecision="1" longitude="48.31278" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Lankaran" typeStatus="lectotype">
<paragraph id="EF1536884D160445F596B67C6467F8D9" blockId="19.[189,1399,1889,2025]" box="[234,792,1889,1916]" pageId="19" pageNumber="20">
<emphasis id="DDDEEA9A4D160445F596B67C6467F8D9" bold="true" box="[234,792,1889,1916]" pageId="19" pageNumber="20">
<typeStatus id="3011882A4D160445F596B67C6619F8D9" box="[234,358,1890,1916]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="28AA4D0B4D160445F4F1B67F6467F8D9" authorityName="Reitter" authorityYear="1887" box="[397,792,1889,1916]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="variety" species="formicaria" variety="major">
<emphasis id="DDDEEA9A4D160445F4F1B67F65ECF8DE" bold="true" box="[397,659,1889,1916]" italics="true" pageId="19" pageNumber="20">Coluocera formicaria</emphasis>
var.
<emphasis id="DDDEEA9A4D160445F7ACB67C6467F8D9" bold="true" box="[720,792,1890,1916]" italics="true" pageId="19" pageNumber="20">major</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="EF1536884D160445F5C1B69865ABF84D" blockId="19.[189,1399,1889,2025]" pageId="19" pageNumber="20">
<collectingCountry id="97BD76184D160445F5C1B698660DF805" box="[189,370,1926,1952]" name="Azerbaijan" pageId="19" pageNumber="20">AZERBAIJAN</collectingCountry>
<emphasis id="DDDEEA9A4D160445F4E8B698655BF805" bold="true" box="[404,548,1926,1952]" pageId="19" pageNumber="20">
<collectingRegion id="2D6EF86A4D160445F4E8B698656DF805" box="[404,530,1926,1952]" country="Azerbaijan" name="Lankaran" pageId="19" pageNumber="20">Lankaran</collectingRegion>
</emphasis>
<specimenCount id="F9ACFD014D160445F750B6986526F805" box="[556,601,1926,1952]" pageId="19" pageNumber="20">
1
<emphasis id="DDDEEA9A4D160445F73FB6986526F805" bold="true" box="[579,601,1926,1952]" pageId="19" pageNumber="20"></emphasis>
</specimenCount>
; “
<location id="EA7560534D160445F709B698642EF805" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D160445F709B698642EF805" box="[629,849,1926,1952]" country="Azerbaijan" latitude="38.670002" longLatPrecision="1" longitude="48.31278" name="Caspi. - M. - Gebiet" pageId="19" pageNumber="20" stateProvince="Lankaran">Caspi.M.Gebiet</location>
” [
<location id="EA7560534D160445F617B6986480F805" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D160445F617B6986480F805" box="[875,1023,1926,1952]" country="Azerbaijan" latitude="38.670002" longLatPrecision="1" longitude="48.31278" name="Caspian Sea" pageId="19" pageNumber="20" stateProvince="Lankaran">Caspian Sea</location>
region],
<location id="EA7560534D160445F111B69963BCF804" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D160445F111B69963BCF804" box="[1133,1219,1927,1953]" country="Azerbaijan" latitude="38.670002" longLatPrecision="1" longitude="48.31278" name="Rasano" pageId="19" pageNumber="20" stateProvince="Lankaran">Rasano</location>
[
<location id="EA7560534D160445F1AFB698624CF805" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D160445F1AFB698624CF805" box="[1235,1331,1926,1952]" country="Azerbaijan" latitude="38.670002" longLatPrecision="1" longitude="48.31278" name="Ancient" pageId="19" pageNumber="20" stateProvince="Lankaran">Ancient</location>
town in ruins,
<quantity id="28529B6D4D160445F450B6B46608F861" box="[300,375,1962,1989]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="20" unit="km" value="15.0">
<locationDeviation id="FEA72FAA4D160445F450B6B46608F861" box="[300,375,1962,1989]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="20" unit="km" value="15.0">15 km</locationDeviation>
</quantity>
southwest of
<collectingRegion id="2D6EF86A4D160445F758B6B46518F861" box="[548,615,1962,1988]" country="Azerbaijan" name="Lerik" pageId="19" pageNumber="20">Lerik</collectingRegion>
,
<geoCoordinate id="8A9E504F4D160445F70AB6B7645BF860" box="[630,804,1961,1989]" degrees="38" direction="north" minutes="40" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="12.0" value="38.670002">38°4012.0″ N</geoCoordinate>
,
<geoCoordinate id="8A9E504F4D160445F64FB6B764A1F860" box="[819,990,1961,1989]" degrees="48" direction="east" minutes="18" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="46.0" value="48.31278">48°18'46.0″ E</geoCoordinate>
. See
<bibRefCitation id="8B3B4B794D160445F15DB6B563A1F861" author="Lazarev M. A." box="[1057,1246,1962,1989]" pageId="19" pageNumber="20" pagination="878 - 888" refId="ref40303" refString="Lazarev M. A. 2017. A new subspecies of Dorcadion (Cribridorcadion) talyschense Ganglbauer, 1884 from Iran (Coleoptera, Cerambycidae). Humanity Space. International almanac 6 (5): 878 - 888." type="journal article" year="2017">Lazarev (2017)</bibRefCitation>
];
<collectorName id="425F535E4D160445F188B6B46247F861" box="[1268,1336,1962,1988]" pageId="19" pageNumber="20">Leder</collectorName>
leg.;
<collectionCode id="89BBAE4D4D160445F5C1B6D16665F84C" box="[189,282,1999,2025]" collectionName="Hungary, Budapest, Hungarian Natural History Museum" pageId="19" pageNumber="20">HNHM</collectionCode>
. Designated by
<bibRefCitation id="8B3B4B794D160445F4A9B6D065AFF84D" author="Rucker W. H." box="[469,720,1998,2024]" pageId="19" pageNumber="20" pagination="11 - 20" refId="ref41653" refString="Rucker W. H. 2011 a. Ergebnisse von Typenstudien der Gattungen Metophthalmus Motschulsky, 1850 (Coleoptera: Latridiidae), Cholovocera Motschulsky, 1838, Reitteria Leder, 1872, Merophysia Lucas, 1852 (Endomychidae: Merophysiinae) und Holoparamecus Curtis, 1833 (Endomychidae: Holoparamecinae). Latridiidae - Mitteilungsblatt fur Systematik und Taxonomie der Latridiidae 8: 11 - 20." type="journal article" year="2011">Rückers (2011a: 13)</bibRefCitation>
.
</paragraph>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F596B0176310FECF" collectionCode="HNHM" collectorName="Rasano &amp; Leder" country="Azerbaijan" determinerName="X. Espadaler" location="Caspi. - M. - Gebiet" pageId="20" pageNumber="21" specimenCount="2" stateProvince="Lankaran" typeStatus="paralectotype">
<paragraph id="EF1536884D110442F596B017642AFE87" blockId="20.[188,1399,264,362]" box="[234,853,264,291]" pageId="20" pageNumber="21">
<emphasis id="DDDEEA9A4D110442F596B017642AFE87" bold="true" box="[234,853,264,291]" pageId="20" pageNumber="21">
<typeStatus id="3011882A4D110442F596B01766DDFE86" box="[234,418,265,291]" pageId="20" pageNumber="21" type="paralectotype">Paralectotypes</typeStatus>
of
<taxonomicName id="28AA4D0B4D110442F4B6B016642AFE87" authorityName="Reitter" authorityYear="1887" box="[458,853,264,291]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="variety" species="formicaria" variety="major">
<emphasis id="DDDEEA9A4D110442F4B6B01665B0FE87" bold="true" box="[458,719,264,291]" italics="true" pageId="20" pageNumber="21">Coluocera formicaria</emphasis>
var.
<emphasis id="DDDEEA9A4D110442F671B016642AFE87" bold="true" box="[781,853,264,290]" italics="true" pageId="20" pageNumber="21">major</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="EF1536884D110442F5C1B032630BFECF" blockId="20.[188,1399,264,362]" pageId="20" pageNumber="21">
<collectingCountry id="97BD76184D110442F5C1B032660DFEE3" box="[189,370,300,326]" name="Azerbaijan" pageId="20" pageNumber="21">AZERBAIJAN</collectingCountry>
<emphasis id="DDDEEA9A4D110442F4F0B0326575FEE3" bold="true" box="[396,522,300,326]" pageId="20" pageNumber="21">
<collectingRegion id="2D6EF86A4D110442F4F0B0326575FEE3" box="[396,522,300,326]" country="Azerbaijan" name="Lankaran" pageId="20" pageNumber="21">Lankaran</collectingRegion>
</emphasis>
<specimenCount id="F9ACFD014D110442F75DB0326515FEE2" box="[545,618,300,327]" pageId="20" pageNumber="21">
2
<emphasis id="DDDEEA9A4D110442F749B032651EFEE3" bold="true" box="[565,609,300,326]" pageId="20" pageNumber="21">♂♂</emphasis>
;
</specimenCount>
<location id="EA7560534D110442F707B0326428FEE3" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D110442F707B0326428FEE3" box="[635,855,300,326]" country="Azerbaijan" name="Caspi. - M. - Gebiet" pageId="20" pageNumber="21" stateProvince="Lankaran">Caspi.M.Gebiet</location>
” [
<location id="EA7560534D110442F612B0326343FEE2" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D110442F612B0326343FEE2" box="[878,1084,300,327]" country="Azerbaijan" name="Caspian Sea area" pageId="20" pageNumber="21" stateProvince="Lankaran">Caspian Sea area</location>
],
<collectorName id="425F535E4D110442F132B03363D7FEE2" box="[1102,1192,301,327]" pageId="20" pageNumber="21">Rasano</collectorName>
;
<collectorName id="425F535E4D110442F1CFB0326388FEE3" box="[1203,1271,300,326]" pageId="20" pageNumber="21">Leder</collectorName>
leg.; [each specimen associated with a
<taxonomicName id="28AA4D0B4D110442F77FB04E65E4FECC" authorityName="Mayr" authorityYear="1855" box="[515,667,336,361]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D110442F77FB04E65E4FECC" box="[515,667,336,361]" italics="true" pageId="20" pageNumber="21">Tetramorium</emphasis>
</taxonomicName>
worker
<vernacularName id="61A946A64D110442F781B04E645BFECF" authorityName="Latreille" authorityYear="1802" box="[765,804,336,362]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">ant</vernacularName>
, det. X. Espadaler];
<collectionCode id="89BBAE4D4D110442F16EB04E6310FECF" box="[1042,1135,336,362]" collectionName="Hungary, Budapest, Hungarian Natural History Museum" pageId="20" pageNumber="21">HNHM</collectionCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="A7B065034D110442F596B08C63FAFD3A" pageId="20" pageNumber="21" type="discussion">
<paragraph id="EF1536884D110442F596B08C664FFE09" blockId="20.[189,1399,402,605]" box="[234,304,402,428]" pageId="20" pageNumber="21">
<emphasis id="DDDEEA9A4D110442F596B08C664FFE09" bold="true" box="[234,304,402,428]" pageId="20" pageNumber="21">Notes</emphasis>
</paragraph>
<paragraph id="EF1536884D110442F5C1B0AB65C8FDF8" blockId="20.[189,1399,402,605]" pageId="20" pageNumber="21">
As it can be seen in
<bibRefCitation id="8B3B4B794D110442F4DEB0AB653BFE6A" author="Rucker W. H." box="[418,580,437,463]" pageId="20" pageNumber="21" pagination="11 - 20" refId="ref41653" refString="Rucker W. H. 2011 a. Ergebnisse von Typenstudien der Gattungen Metophthalmus Motschulsky, 1850 (Coleoptera: Latridiidae), Cholovocera Motschulsky, 1838, Reitteria Leder, 1872, Merophysia Lucas, 1852 (Endomychidae: Merophysiinae) und Holoparamecus Curtis, 1833 (Endomychidae: Holoparamecinae). Latridiidae - Mitteilungsblatt fur Systematik und Taxonomie der Latridiidae 8: 11 - 20." type="journal article" year="2011">Rücker 2011a</bibRefCitation>
(fig. 18), there is a label reading “
<typeStatus id="3011882A4D110442F6B5B0AB6337FE6A" box="[969,1096,437,463]" pageId="20" pageNumber="21" type="holotype">Holotypus</typeStatus>
” attached to the specimen that Rücker designated as the
<typeStatus id="3011882A4D110442F756B0C665E6FE57" box="[554,665,472,498]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
. However, this specimen cannot be regarded as the
<typeStatus id="3011882A4D110442F06CB0C66208FE57" box="[1296,1399,472,498]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
because it was not designated in the original description, which included more than
<specimenCount id="F9ACFD014D110442F1E7B0E36240FDB3" box="[1179,1343,508,535]" count="1" pageId="20" pageNumber="21" type="generic">one specimen</specimenCount>
, i.e.,
<typeStatus id="3011882A4D110442F5C1B33E6658FD9F" box="[189,295,544,570]" pageId="20" pageNumber="21" type="syntype">syntypes</typeStatus>
. Examining the handwriting of the
<typeStatus id="3011882A4D110442F7ACB33E6435FD9F" box="[720,842,544,570]" pageId="20" pageNumber="21" type="holotype">Holotypus</typeStatus>
label, we conclude that it was added at a later date than the description by
<bibRefCitation id="8B3B4B794D110442F776B35D65CCFDF8" author="Reitter E." box="[522,691,579,605]" pageId="20" pageNumber="21" refId="ref41271" refString="Reitter E. 1887. Bestimmungs-Tabellen der europaischen Coleopteren. III Helft. II Auflage. Enthaltend die Familien: Scaphidiidae, Lathridiidae und Dermestidae. E. Reitter, M ˆ dling." type="book" year="1887">Reitter (1887)</bibRefCitation>
.
</paragraph>
<paragraph id="EF1536884D110442F5C1B39B63FAFD3A" blockId="20.[189,1157,645,672]" box="[189,1157,645,672]" pageId="20" pageNumber="21">
The same comment given above under the
<typeStatus id="3011882A4D110442F7C9B39B645BFD3A" box="[693,804,645,671]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
, applies to the
<typeStatus id="3011882A4D110442F6AFB39B63FFFD3A" box="[979,1152,645,671]" pageId="20" pageNumber="21" type="paralectotype">paralectotypes</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D110442F5C1B3D8622BF998" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="EF1536884D110442F5C1B3D86542FD44" blockId="20.[189,573,710,737]" box="[189,573,710,737]" pageId="20" pageNumber="21">
<heading id="B45D81E44D110442F5C1B3D86542FD44" bold="true" box="[189,573,710,737]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="DDDEEA9A4D110442F5C1B3D86542FD44" bold="true" box="[189,573,710,737]" pageId="20" pageNumber="21">Additional material, non-types</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D110442F5C1B3E864A4FC96" blockId="20.[189,1398,756,820]" pageId="20" pageNumber="21">
<materialsCitation id="5FC23CD54D110442F5C1B3E86444FCB5" box="[189,827,758,784]" country="Russia" location="Derbent" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="Dagestan">
<collectingCountry id="97BD76184D110442F5C1B3E8665BFCB5" box="[189,292,758,784]" name="Russia" pageId="20" pageNumber="21">RUSSIA</collectingCountry>
<emphasis id="DDDEEA9A4D110442F438B3E866C7FCB5" bold="true" box="[324,440,758,784]" pageId="20" pageNumber="21">
<collectingRegion id="2D6EF86A4D110442F438B3E866C7FCB5" box="[324,440,758,784]" country="Russia" name="Dagestan" pageId="20" pageNumber="21">Dagestan</collectingRegion>
</emphasis>
<specimenCount id="F9ACFD014D110442F4ADB3E86681FCB5" box="[465,510,758,784]" count="1" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
;
<location id="EA7560534D110442F773B3E8650EFCB5" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D110442F773B3E8650EFCB5" box="[527,625,758,784]" country="Russia" name="Derbent" pageId="20" pageNumber="21" stateProvince="Dagestan">Derbent</location>
;
<collectionCode id="89BBAE4D4D110442F702B3E865A4FCB5" box="[638,731,758,784]" pageId="20" pageNumber="21">MCNM</collectionCode>
198707
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F629B3E863F0FCB5" box="[853,1167,756,784]" collectionCode="SMTD" country="Russia" location="Derbent" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1">
1 ♂,
<specimenCount id="F9ACFD014D110442F6EEB3E864C0FCB5" box="[914,959,758,784]" count="1" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
;
<location id="EA7560534D110442F6B3B3E8634EFCB5" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D110442F6B3B3E8634EFCB5" box="[975,1073,758,784]" country="Russia" name="Derbent" pageId="20" pageNumber="21">Derbent</location>
;
<collectionCode id="89BBAE4D4D110442F142B3E863F0FCB5" box="[1086,1167,758,784]" pageId="20" pageNumber="21">SMTD</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F1D6B3E86656FC96" country="Turkey" location="East Anatolia" pageId="20" pageNumber="21" specimenCount="1">
<collectingCountry id="97BD76184D110442F1D6B3E8625FFCB5" box="[1194,1312,758,784]" name="Turkey" pageId="20" pageNumber="21">TURKEY</collectingCountry>
<emphasis id="DDDEEA9A4D110442F043B3E86656FC96" bold="true" pageId="20" pageNumber="21">
<location id="EA7560534D110442F043B3E86656FC96" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D110442F043B3E86656FC96" country="Turkey" name="East Anatolia" pageId="20" pageNumber="21">East Anatolia</location>
</emphasis>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F43DB20764A8FC96" box="[321,983,793,820]" collectingDate="1986-06-15" country="Turkey" elevation="1650" location="Digor" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="2" stateProvince="Kars">
<specimenCount id="F9ACFD014D110442F43DB20766FCFC91" box="[321,387,793,820]" count="2" pageId="20" pageNumber="21" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="2D6EF86A4D110442F4EFB20466B2FC91" box="[403,461,794,820]" country="Turkey" name="Kars" pageId="20" pageNumber="21">Kars</collectingRegion>
,
<location id="EA7560534D110442F4A5B207655EFC96" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D110442F4A5B207655EFC96" box="[473,545,793,819]" country="Turkey" name="Digor" pageId="20" pageNumber="21" stateProvince="Kars">Digor</location>
;
<elevation id="6487D1BB4D110442F751B20765B9FC96" box="[557,710,793,820]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="20" pageNumber="21" unit="m" value="1650.0">
<quantity id="28529B6D4D110442F751B20765F9FC91" box="[557,646,793,820]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="20" pageNumber="21" unit="m" value="1650.0">1650 m</quantity>
a.s.l.
</elevation>
;
<date id="9B1410484D110442F7AFB2076412FC96" box="[723,877,793,820]" pageId="20" pageNumber="21" value="1986-06-15">
<collectingDate id="8B50E9A04D110442F7AFB2076412FC96" box="[723,877,793,820]" pageId="20" pageNumber="21" value="1986-06-15">15 Jun. 1986</collectingDate>
</date>
;
<collectionCode id="89BBAE4D4D110442F605B20764A8FC96" box="[889,983,793,819]" pageId="20" pageNumber="21">MHNG</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="EF1536884D110442F5C1B2456379FA85" blockId="20.[189,1399,857,1312]" pageId="20" pageNumber="21">
<materialsCitation id="5FC23CD54D110442F5C1B2456256FCD0" box="[189,1321,857,888]" collectionCode="NMPC" collectorName="Couril" country="Azerbaijan" location="Goygol" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Goygol">
<collectingCountry id="97BD76184D110442F5C1B245660DFCD0" box="[189,370,859,885]" name="Azerbaijan" pageId="20" pageNumber="21">AZERBAIJAN</collectingCountry>
<emphasis id="DDDEEA9A4D110442F4F3B2426697FCDD" bold="true" box="[399,488,860,888]" pageId="20" pageNumber="21">Ĝyĝl</emphasis>
• 1 ♂; “Caucasus, Helenendorf” [modern
<collectingRegion id="2D6EF86A4D110442F6A5B2456348FCD0" box="[985,1079,859,885]" country="Azerbaijan" name="Goygol" pageId="20" pageNumber="21">Göygöl</collectingRegion>
];
<collectorName id="425F535E4D110442F136B24563EAFCD0" box="[1098,1173,859,885]" pageId="20" pageNumber="21">Couřil</collectorName>
leg.;
<collectionCode id="89BBAE4D4D110442F1AAB2456256FCD0" box="[1238,1321,859,885]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F03EB24565BCFC3D" collectionCode="SMTD" collectorName="Helenendorf" country="Azerbaijan" location="Ganja" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Ganca">
1 ♂; “Caucasus“,
<collectorName id="425F535E4D110442F42FB2616693FC3C" box="[339,492,895,921]" pageId="20" pageNumber="21">Helenendorf</collectorName>
“;
<collectionCode id="89BBAE4D4D110442F77DB2616529FC3C" box="[513,598,895,921]" pageId="20" pageNumber="21">SMTD</collectionCode>
.
<emphasis id="DDDEEA9A4D110442F709B26065BCFC3D" bold="true" box="[629,707,894,920]" pageId="20" pageNumber="21">
<collectingRegion id="2D6EF86A4D110442F709B26065BCFC3D" box="[629,707,894,920]" country="Azerbaijan" name="Ganca" pageId="20" pageNumber="21">Ganja</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F7A7B261660CFC18" collectionCode="SDEI" country="Azerbaijan" location="Ganja" pageId="20" pageNumber="21" specimenCode="SDEI 10850, 10851" specimenCount="1" specimenCount-female="1" stateProvince="Ganca">
1 ♂,
<specimenCount id="F9ACFD014D110442F66FB2616441FC3C" box="[787,830,895,921]" count="1" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
; “Caucasus, Elisabetpol” [modern
<collectingRegion id="2D6EF86A4D110442F1ABB261625BFC3C" box="[1239,1316,895,921]" country="Azerbaijan" name="Ganca" pageId="20" pageNumber="21">Ganja</collectingRegion>
];
<specimenCode id="BF0C9EF34D110442F049B2616679FC18" collectionCode="SDEI" pageId="20" pageNumber="21">SDEI 10850</specimenCode>
to
<specimenCode id="BF0C9EF34D110442F456B2BC660CFC18" box="[298,371,930,957]" collectionCode="SDEI" pageId="20" pageNumber="21">10851</specimenCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F4F7B2BC633BFC19" box="[395,1092,930,957]" collectionCode="NHMW" country="Azerbaijan" location="Lankaran" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Lankaran">
<specimenCount id="F9ACFD014D110442F4F7B2BC656CFC19" box="[395,531,930,956]" count="1" pageId="20" pageNumber="21" type="generic">1 specimen</specimenCount>
; “Caucasus, Elisabetpol”;
<collectionCode id="89BBAE4D4D110442F63AB2BD64D8FC18" box="[838,935,931,957]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="21" type="Museum">NHMW</collectionCode>
.
<emphasis id="DDDEEA9A4D110442F6BAB2BC633BFC19" bold="true" box="[966,1092,930,956]" pageId="20" pageNumber="21">
<collectingRegion id="2D6EF86A4D110442F6BAB2BC633BFC19" box="[966,1092,930,956]" country="Azerbaijan" name="Lankaran" pageId="20" pageNumber="21">Lankaran</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F127B2BC662EFC45" collectionCode="NHMB" collectorName="Leder" country="Azerbaijan" pageId="20" pageNumber="21" specimenCount="3">
1 ♂,
<specimenCount id="F9ACFD014D110442F1E8B2BC6258FC19" box="[1172,1319,930,956]" count="3" pageId="20" pageNumber="21" type="generic">3 specimens</specimenCount>
;
<collectorName id="425F535E4D110442F04EB2BC6209FC19" box="[1330,1398,930,956]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<collectionCode id="89BBAE4D4D110442F585B2D8662EFC45" box="[249,337,966,992]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="20" pageNumber="21">NHMB</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F411B2D865B8FC45" box="[365,711,966,994]" collectionCode="SDEI" collectorName="Leder" country="Azerbaijan" pageId="20" pageNumber="21" specimenCode="SDEI 11874" specimenCount="1" specimenCount-female="1">
<specimenCount id="F9ACFD014D110442F411B2D866E5FC47" box="[365,410,966,994]" count="1" pageId="20" pageNumber="21" type="female">
1
<emphasis id="DDDEEA9A4D110442F4FAB2D866E5FC47" bold="true" box="[390,410,966,994]" pageId="20" pageNumber="21"></emphasis>
</specimenCount>
;
<collectorName id="425F535E4D110442F4D0B2D8668FFC45" box="[428,496,966,992]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<specimenCode id="BF0C9EF34D110442F749B2D865B8FC45" box="[565,711,966,992]" collectionCode="SDEI" pageId="20" pageNumber="21">SDEI 11874</specimenCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F79FB2D86321FC45" box="[739,1118,966,992]" pageId="20" pageNumber="21">
<specimenCount id="F9ACFD014D110442F79FB2D86411FC45" box="[739,878,966,992]" count="1" pageId="20" pageNumber="21" type="generic">1 specimen</specimenCount>
; Leder leg.;
<collectionCode id="89BBAE4D4D110442F178B2D86321FC45" box="[1028,1118,966,992]" pageId="20" pageNumber="21">MHNG</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F106B2D8666FFBA6" collectionCode="NMPC" collectorName="Leder" country="Azerbaijan" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="2">
2 ♂,
<specimenCount id="F9ACFD014D110442F1CBB2D8639AFC45" box="[1207,1253,966,992]" count="2" pageId="20" pageNumber="21" type="female">2 ♀</specimenCount>
;
<collectorName id="425F535E4D110442F18AB2D86245FC45" box="[1270,1338,966,992]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<collectionCode id="89BBAE4D4D110442F5C1B2F7666FFBA6" box="[189,272,1001,1027]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F45AB2F7622EFBA1" box="[294,1361,999,1028]" collectionCode="SDEI" collectorName="Leder" country="Azerbaijan" location="Talish area" pageId="20" pageNumber="21" specimenCode="SDEI 11875-11876" specimenCount="1" specimenCount-female="1">
1 ♂,
<specimenCount id="F9ACFD014D110442F423B2F766F6FBA1" box="[351,393,1001,1028]" count="1" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
; “Talyschgeb[er]g [
<location id="EA7560534D110442F713B2F76589FBA1" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D110442F713B2F76589FBA1" box="[623,758,1001,1028]" country="Azerbaijan" name="Talish area" pageId="20" pageNumber="21">Talish area</location>
], Transcaucas[us].”;
<collectorName id="425F535E4D110442F69BB2F76354FBA6" box="[999,1067,1001,1027]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<specimenCode id="BF0C9EF34D110442F117B2F7622EFBA1" box="[1131,1361,1001,1028]" collectionCode="SDEI" pageId="20" pageNumber="21">SDEI 1187511876</specimenCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F014B2F764B8FB82" collectionCode="SFUN" collectorName="Leder" country="Azerbaijan" pageId="20" pageNumber="21" specimenCount="3">
2 ♂♂,
<specimenCount id="F9ACFD014D110442F58AB51366F7FB82" box="[246,392,1037,1063]" count="3" pageId="20" pageNumber="21" type="generic">3 specimens</specimenCount>
; “Talyschgebg., Transcaucas.”;
<collectorName id="425F535E4D110442F785B5136442FB82" box="[761,829,1037,1063]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<collectionCode id="89BBAE4D4D110442F600B51364B8FB82" box="[892,967,1037,1063]" pageId="20" pageNumber="21">SFUN</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F6A1B51366E1FBEE" pageId="20" pageNumber="21">
1 ♂; “Talyschgebg., Transcaucas.”;
<collectorName id="425F535E4D110442F5C1B52E667EFBEF" box="[189,257,1072,1098]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<collectionCode id="89BBAE4D4D110442F431B52F66E1FBEE" box="[333,414,1073,1099]" pageId="20" pageNumber="21">SMTD</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F4BDB52F63BDFBEE" box="[449,1218,1072,1099]" pageId="20" pageNumber="21">
<specimenCount id="F9ACFD014D110442F4BDB52F6525FBEF" box="[449,602,1072,1099]" count="2" pageId="20" pageNumber="21" type="generic">2 specimens</specimenCount>
; “Talyschgebg., Transcaucas.”; Leder leg.;
<collectionCode id="89BBAE4D4D110442F113B52F63BDFBEE" box="[1135,1218,1073,1099]" pageId="20" pageNumber="21">MFNB</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F199B52F64FCFBCB" collectionCode="SFUN" collectorName="Leder" country="Azerbaijan" location="Ordubad" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Ordubad">
<specimenCount id="F9ACFD014D110442F199B52F620CFBEF" box="[1253,1395,1072,1099]" count="1" pageId="20" pageNumber="21" type="generic">1 specimen</specimenCount>
; “Caspi.M.Gebiet, Rasano”;
<collectorName id="425F535E4D110442F75CB54A651BFBCB" box="[544,612,1108,1134]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<collectionCode id="89BBAE4D4D110442F7DFB54A658DFBCB" box="[675,754,1108,1134]" pageId="20" pageNumber="21">SFUN</collectionCode>
.
<emphasis id="DDDEEA9A4D110442F673B54A64FCFBCB" bold="true" box="[783,899,1108,1134]" pageId="20" pageNumber="21">
<collectingRegion id="2D6EF86A4D110442F673B54A64FCFBCB" box="[783,899,1108,1134]" country="Azerbaijan" name="Ordubad" pageId="20" pageNumber="21">Ordubad</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F6E6B54A636EFB34" collectionCode="NMPC" collectorName="Leder" country="Azerbaijan" location="Caucasus" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="1">
<specimenCount id="F9ACFD014D110442F6E6B54A64BBFBCB" box="[922,964,1108,1134]" count="1" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
,
<specimenCount id="F9ACFD014D110442F6AEB54A6326FBCB" box="[978,1113,1108,1134]" count="1" pageId="20" pageNumber="21" type="generic">1 specimen</specimenCount>
; “Caucasus, Araxesthal” [Arax River Valley];
<collectorName id="425F535E4D110442F4CEB5696689FB34" box="[434,502,1143,1169]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<collectionCode id="89BBAE4D4D110442F74AB56665F2FB37" box="[566,653,1144,1170]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
.
<emphasis id="DDDEEA9A4D110442F7D6B569636EFB34" bold="true" box="[682,1041,1143,1170]" pageId="20" pageNumber="21">
<location id="EA7560534D110442F7D6B569645CFB34" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D110442F7D6B569645CFB34" box="[682,803,1143,1169]" country="Azerbaijan" name="Caucasus" pageId="20" pageNumber="21">Caucasus</location>
, no specific locality
</emphasis>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F155B5666576FB10" collectionCode="SDEI" collectorName="Leder" country="Azerbaijan" pageId="20" pageNumber="21" specimenCode="SDEI 10842- 10843" specimenCount="1">
2 ♂♂; “Kaukas”;
<collectorName id="425F535E4D110442F184B5696243FB34" box="[1272,1340,1143,1169]" pageId="20" pageNumber="21">Leder</collectorName>
leg.; [18]86;
<specimenCode id="BF0C9EF34D110442F466B5856576FB10" box="[282,521,1179,1205]" collectionCode="SDEI" pageId="20" pageNumber="21">SDEI 10842 10843</specimenCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F75DB58563DEFB10" box="[545,1185,1177,1206]" collectionCode="SDEI" collectorName="Leder" country="Azerbaijan" pageId="20" pageNumber="21" specimenCode="SDEI 10847- 10849" specimenCount="2" specimenCount-female="2">
1 ♂,
<specimenCount id="F9ACFD014D110442F726B58565E4FB13" box="[602,667,1179,1206]" count="2" pageId="20" pageNumber="21" type="female">2 ♀♀</specimenCount>
; “Kaukas”;
<collectorName id="425F535E4D110442F656B5856411FB10" box="[810,878,1179,1205]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<specimenCode id="BF0C9EF34D110442F6D3B58563DEFB10" box="[943,1185,1179,1205]" collectionCode="SDEI" pageId="20" pageNumber="21">SDEI 10847 10849</specimenCode>
</materialsCitation>
” •
<materialsCitation id="5FC23CD54D110442F1BEB58566E3FB7C" collectionCode="NMPC" collectorName="Leder" country="Azerbaijan" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1">
<specimenCount id="F9ACFD014D110442F1BEB5856391FB13" box="[1218,1262,1179,1206]" count="1" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
; “Kaukas”;
<collectorName id="425F535E4D110442F5C1B5A1667EFB7C" box="[189,257,1215,1241]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<collectionCode id="89BBAE4D4D110442F435B5A166E3FB7C" box="[329,412,1215,1241]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F4C7B5A1647AFB7C" box="[443,773,1213,1241]" pageId="20" pageNumber="21">
1 ♂; “Cauc. Sept.”;
<collectionCode id="89BBAE4D4D110442F7CEB5A1647AFB7C" box="[690,773,1215,1241]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="21">NMPC</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F658B5A1625FFB7C" box="[804,1312,1215,1241]" collectionCode="ZFMK" collectorName="Leder" country="Azerbaijan" pageId="20" pageNumber="21" specimenCount="1">
<specimenCount id="F9ACFD014D110442F658B5A164CFFB7C" box="[804,944,1215,1241]" count="1" pageId="20" pageNumber="21" type="generic">1 specimen</specimenCount>
; “Kaukas”;
<collectorName id="425F535E4D110442F13FB5A163F8FB7C" box="[1091,1159,1215,1241]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<collectionCode id="89BBAE4D4D110442F1B3B5A1625FFB7C" box="[1231,1312,1215,1241]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="21" type="Museum">ZFMK</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F043B5A16579FB58" pageId="20" pageNumber="21">
1 ♂; “Kauka”;
<collectorName id="425F535E4D110442F44EB5FC6609FB59" box="[306,374,1250,1276]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<collectionCode id="89BBAE4D4D110442F4C9B5FD6579FB58" box="[437,518,1251,1277]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="21" type="Museum">ZFMK</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F761B5FC6345FB59" box="[541,1082,1250,1277]" collectionCode="MZLU" collectorName="Leder" country="Azerbaijan" pageId="20" pageNumber="21" specimenCode="MZLU 2020-001" specimenCount="1" specimenCount-female="1">
<specimenCount id="F9ACFD014D110442F761B5FC6537FB58" box="[541,584,1250,1277]" count="1" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
; “Caucasus”;
<collectorName id="425F535E4D110442F791B5FC644EFB59" box="[749,817,1250,1276]" pageId="20" pageNumber="21">Leder</collectorName>
leg.;
<specimenCode id="BF0C9EF34D110442F60DB5FD6345FB59" box="[881,1082,1250,1277]" collectionCode="MZLU" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="20" pageNumber="21">MZLU 2020-001</specimenCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F12DB5FC6671FA85" collectionCode="ZFMK" country="Azerbaijan" pageId="20" pageNumber="21" specimenCount="1">
<specimenCount id="F9ACFD014D110442F12DB5FC63A7FB59" box="[1105,1240,1250,1276]" count="1" pageId="20" pageNumber="21" type="generic">1 specimen</specimenCount>
; “Kaukasus”;
<collectionCode id="89BBAE4D4D110442F5C1B4186671FA85" box="[189,270,1286,1312]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="21" type="Museum">ZFMK</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F45BB418651DFA85" box="[295,610,1286,1312]" collectionCode="SMTD" country="Azerbaijan" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="2">
<specimenCount id="F9ACFD014D110442F45BB4186616FA85" box="[295,361,1286,1312]" count="2" pageId="20" pageNumber="21" type="female">2 ♀♀</specimenCount>
; “Caucasus”;
<collectionCode id="89BBAE4D4D110442F76DB418651DFA85" box="[529,610,1286,1312]" pageId="20" pageNumber="21">SMTD</collectionCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F700B418637DFA85" box="[636,1026,1286,1312]" pageId="20" pageNumber="21">
<specimenCount id="F9ACFD014D110442F700B418647AFA85" box="[636,773,1286,1312]" count="1" pageId="20" pageNumber="21" type="generic">1 specimen</specimenCount>
; “Kaukasus”;
<collectionCode id="89BBAE4D4D110442F6D7B418637DFA85" box="[939,1026,1286,1312]" pageId="20" pageNumber="21">MFNB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="EF1536884D110442F5C1B456662CFA68" blockId="20.[189,1399,1349,1485]" pageId="20" pageNumber="21">
<materialsCitation id="5FC23CD54D110442F5C1B4566232FAC7" box="[189,1357,1349,1378]" collectingDate="1970-04-29" collectorName="Wittmer &amp; Bothmer" country="Iran" location="Chalus-Polzoghal" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="2" stateProvince="Gilan">
<collectingCountry id="97BD76184D110442F5C1B456667AFAC7" box="[189,261,1352,1378]" name="Iran" pageId="20" pageNumber="21">IRAN</collectingCountry>
<emphasis id="DDDEEA9A4D110442F459B4596604FAC4" bold="true" box="[293,379,1351,1377]" pageId="20" pageNumber="21">
<collectingRegion id="2D6EF86A4D110442F459B4596604FAC4" box="[293,379,1351,1377]" country="Iran" name="Gilan" pageId="20" pageNumber="21">Guilan</collectingRegion>
</emphasis>
• 1 ♂,
<specimenCount id="F9ACFD014D110442F4ADB456656BFAC7" box="[465,532,1352,1378]" count="2" pageId="20" pageNumber="21" type="female">2 ♀♀</specimenCount>
;
<location id="EA7560534D110442F758B4596587FAC4" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D110442F758B4596587FAC4" box="[548,760,1351,1377]" country="Iran" name="Chalus-Polzoghal" pageId="20" pageNumber="21" stateProvince="Gilan">Chalus-Polzoghal</location>
;
<date id="9B1410484D110442F679B45664DCFAC7" box="[773,931,1352,1378]" pageId="20" pageNumber="21" value="1970-04-29">
<collectingDate id="8B50E9A04D110442F679B45664DCFAC7" box="[773,931,1352,1378]" pageId="20" pageNumber="21" value="1970-04-29">29 Apr. 1970</collectingDate>
</date>
;
<collectorName id="425F535E4D110442F6D3B459636FFAC4" box="[943,1040,1351,1377]" pageId="20" pageNumber="21">Wittmer</collectorName>
and
<collectorName id="425F535E4D110442F137B45963CDFAC4" box="[1099,1202,1351,1377]" pageId="20" pageNumber="21">Bothmer</collectorName>
leg.; MHNB
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F014B456620CFA20" collectingDate="1973-08-20" collectorName="S. Vit" country="Iran" location="Bandar Pahlavi" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="Gilan">
<specimenCount id="F9ACFD014D110442F014B45667ACFA23" count="1" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
;
<location id="EA7560534D110442F59EB47566EAFA20" LSID="urn:lsid:plazi:treatment:6703879E4D16044EF741B0CB6213FCBF:EA7560534D110442F59EB47566EAFA20" box="[226,405,1387,1413]" country="Iran" name="Bandar Pahlavi" pageId="20" pageNumber="21" stateProvince="Gilan">Bandar Pahlavi</location>
[modern Bandar-e Anzali];
<date id="9B1410484D110442F7A0B4756402FA20" box="[732,893,1387,1413]" pageId="20" pageNumber="21" value="1973-08-20">
<collectingDate id="8B50E9A04D110442F7A0B4756402FA20" box="[732,893,1387,1413]" pageId="20" pageNumber="21" value="1973-08-20">20 Aug. 1973</collectingDate>
</date>
;
<collectorName id="425F535E4D110442F6F4B47564B5FA20" box="[904,970,1387,1413]" pageId="20" pageNumber="21">S. Vit</collectorName>
leg.; “marais” [marshland];
<collectionCode id="89BBAE4D4D110442F069B475620CFA20" box="[1301,1395,1387,1413]" pageId="20" pageNumber="21">MHNG</collectionCode>
</materialsCitation>
.
<materialsCitation id="5FC23CD54D110442F5AFB49164C7FA0D" box="[211,952,1421,1449]" collectionCode="NHMB" collectorName="Dadu &amp; Unknown" country="Iran" location="Mazandaran" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="Mazandaran">
<emphasis id="DDDEEA9A4D110442F5AFB491660BFA0C" bold="true" box="[211,372,1423,1449]" pageId="20" pageNumber="21">
<collectingRegion id="2D6EF86A4D110442F5AFB491660BFA0C" box="[211,372,1423,1449]" country="Iran" name="Mazandaran" pageId="20" pageNumber="21">Mazandaran</collectingRegion>
</emphasis>
• 1 ♂,
<specimenCount id="F9ACFD014D110442F4B5B491668AFA0C" box="[457,501,1423,1449]" count="1" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
;
<collectorName id="425F535E4D110442F779B4916537FA0C" box="[517,584,1423,1449]" pageId="20" pageNumber="21">Dadu</collectorName>
;
<collectionCode id="89BBAE4D4D110442F728B49165D3FA0C" box="[596,684,1423,1449]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="20" pageNumber="21">NHMB</collectionCode>
<emphasis id="DDDEEA9A4D110442F7B6B49164C7FA0D" bold="true" box="[714,952,1422,1449]" pageId="20" pageNumber="21">
<collectorName id="425F535E4D110442F7B6B491643BFA0C" box="[714,836,1423,1449]" pageId="20" pageNumber="21">Unknown</collectorName>
localities
</emphasis>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F6AEB4916256FA0C" box="[978,1321,1421,1449]" collectingDate="1917-11" collectionCode="SDEI" country="Iran" pageId="20" pageNumber="21" specimenCode="SDEI 10793" specimenCount="1" stateProvince="Mazandaran">
1 ♂;
<date id="9B1410484D110442F172B49163F4FA0C" box="[1038,1163,1423,1449]" pageId="20" pageNumber="21" value="1917-11">
<collectingDate id="8B50E9A04D110442F172B49163F4FA0C" box="[1038,1163,1423,1449]" pageId="20" pageNumber="21" value="1917-11">Nov. 1917</collectingDate>
</date>
;
<specimenCode id="BF0C9EF34D110442F1E4B4916256FA0C" box="[1176,1321,1423,1449]" collectionCode="SDEI" pageId="20" pageNumber="21">SDEI 10793</specimenCode>
</materialsCitation>
<materialsCitation id="5FC23CD54D110442F03FB491662CFA68" pageId="20" pageNumber="21">
1 ♂,
<specimenCount id="F9ACFD014D110442F5C1B4AC6796FA68" box="[189,233,1458,1485]" count="1" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
;
<collectionCode id="89BBAE4D4D110442F584B4AD6630FA68" box="[248,335,1459,1485]" pageId="20" pageNumber="21">MFNB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="EF1536884D110442F5C1B4EA6622F9AB" blockId="20.[189,349,1524,1550]" box="[189,349,1524,1550]" pageId="20" pageNumber="21">
<heading id="B45D81E44D110442F5C1B4EA6622F9AB" bold="true" box="[189,349,1524,1550]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="DDDEEA9A4D110442F5C1B4EA6622F9AB" bold="true" box="[189,349,1524,1550]" pageId="20" pageNumber="21">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D110442F5C1B73D622BF998" blockId="20.[189,1364,1571,1597]" box="[189,1364,1571,1597]" pageId="20" pageNumber="21">
“Derbent, non loin de la mer Caspienne” [Derbent, not far from the Caspian Sea],
<collectingRegion id="2D6EF86A4D110442F1FEB73D638DF998" box="[1154,1266,1571,1597]" country="Russia" name="Dagestan" pageId="20" pageNumber="21">Dagestan</collectingRegion>
,
<collectingCountry id="97BD76184D110442F182B73D622FF998" box="[1278,1360,1571,1597]" name="Russia" pageId="20" pageNumber="21">Russia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D110443F5C1B77A65CFFDEF" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="EF1536884D110442F5C1B77A6631F9DB" blockId="20.[189,334,1636,1662]" box="[189,334,1636,1662]" pageId="20" pageNumber="21">
<heading id="B45D81E44D110442F5C1B77A6631F9DB" bold="true" box="[189,334,1636,1662]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="DDDEEA9A4D110442F5C1B77A6631F9DB" bold="true" box="[189,334,1636,1662]" pageId="20" pageNumber="21">Description</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D110442F5C1B78A65FBF826" blockId="20.[188,1400,1683,1923]" pageId="20" pageNumber="21">
Male as in
<figureCitation id="77912A0D4D110442F442B78A66D7F90B" box="[318,424,1684,1710]" captionStart="Fig" captionStartId="41.[189,232,1928,1954]" captionTargetBox="[321,1272,530,1820]" captionTargetId="figure-115@41.[311,1276,513,1878]" captionTargetPageId="41" captionText="Fig. 11. Male habitus, dorsal view. A. Cholovocera formicaria Victor, 1838. B. Ch. formiceticola (Rosenhauer, 1856). C. Ch. occulta sp. nov., paratype. D. Ch. punctata (Märkel, 1845). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10424537" httpUri="https://zenodo.org/record/10424537/files/figure.png" pageId="20" pageNumber="21">Fig. 11A</figureCitation>
. Body length:
<quantity id="28529B6D4D110442F72EB78A65C5F90B" box="[594,698,1684,1710]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="20" pageNumber="21" unit="mm" value="1.43">1.43 mm</quantity>
average, range
<quantity id="28529B6D4D110442F60FB78D6362F90B" box="[883,1053,1683,1710]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="20" pageNumber="21" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.301.50 mm</quantity>
(N = 41, males and females). Shape of body elliptical, with the lateral margins of the pronotum continuous with those of the elytra, i.e., without an indentation. Elytral apex markedly acute. Terminal antennomeres subtriangular, with round angles (
<figureCitation id="77912A0D4D110442F41DB7E066B2F8BD" box="[353,461,1790,1816]" captionStart="Fig" captionStartId="42.[189,232,1892,1918]" captionTargetBox="[341,1310,800,1791]" captionTargetId="figure-179@42.[221,1367,721,1840]" captionTargetPageId="42" captionText="Fig. 12. Terminal female antennomeres. A. Cholovocera afghana Johnson, 1977. B. Ch. attae (Kraatz, 1858). C. Ch. balcánica (Karaman, 1936). D. Ch. gallica (Schaufuss, 1876). E. Ch. formicaria Victor, 1838. F. Ch. formiceticola (Rosenhauer, 1856). G. Ch. occulta sp. nov. H. Ch. punctata (Märkel, 1845). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424539" httpUri="https://zenodo.org/record/10424539/files/figure.png" pageId="20" pageNumber="21">Figs 12E</figureCitation>
,
<figureCitation id="77912A0D4D110442F4A4B7E06571F8BD" box="[472,526,1790,1816]" captionStart="Fig" captionStartId="44.[189,232,1928,1954]" captionTargetBox="[247,1330,760,1834]" captionTargetId="figure-280@44.[217,1371,719,1875]" captionTargetPageId="44" captionText="Fig. 13. Terminal male antennomeres.A. Cholovocera attae (Kraatz, 1858). B. Ch. balcánica(Karaman, 1936). C. Ch. gallica (Schaufuss, 1876). D. Ch. formicaria Victor, 1838. E. Ch. formiceticola (Rosenhauer, 1856). F. Ch. occulta sp. nov. G. Ch. punctata (Märkel, 1845). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424541" httpUri="https://zenodo.org/record/10424541/files/figure.png" pageId="20" pageNumber="21">13D</figureCitation>
). Metatibiae as in
<figureCitation id="77912A0D4D110442F79DB7E064D3F8BD" box="[737,940,1790,1816]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[293,1322,589,1725]" captionTargetId="figure-210@46.[250,1337,543,1774]" captionTargetPageId="46" captionText="Fig. 14. Metatibiae. A. Cholovocera afghana Johnson, 1977, ♀. B. Ch. attae (Kraatz, 1858), ♀. C. Ch. attae (Kraatz, 1858), ♂. D. Ch. balcanica (Karaman, 1936), ♀. E. Ch. balcanica (Karaman, 1936), ♂. F. Ch. gallica (Schaufuss, 1876), ♀. G. Ch. gallica (Schaufuss, 1876), ♂. H. Ch. formicaria Victor, 1838, ♀. I. Ch. formicaria Victor, 1838, ♂. J. Ch. formiceticola (Rosenhauer, 1856), ♀. K. Ch. formiceticola (Rosenhauer, 1856), ♂. L. Ch. occulta sp. nov., ♀. M. Ch. occulta sp. nov., ♂. N. Ch. punctata (Märkel, 1845), ♀. O. Ch. punctata (Märkel, 1845), ♂. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424543" httpUri="https://zenodo.org/record/10424543/files/figure.png" pageId="20" pageNumber="21">Figs 14H and 14I</figureCitation>
, narrower in the proximal half and with sinuous margins, especially in the male. Prosternal process slightly keeled anteriorly, with a marked median constriction and rounded distally (
<figureCitation id="77912A0D4D110442F7D7B65B6478F8C5" box="[683,775,1861,1888]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[264,1318,1324,1816]" captionTargetId="figure-432@10.[187,1400,1277,1877]" captionTargetPageId="10" captionText="Fig. 4. Prosternal processes. A. Cholovocera afghana Johnson, 1977. B. Ch. attae (Kraatz, 1858). C. Ch. balcánica (Karaman, 1936). D. Ch. gallica (Schaufuss, 1876). E. Ch. formicaria Victor, 1838. F. Ch. formiceticola (Rosenhauer, 1856). G. Ch. occulta sp. nov. H. Ch. punctata (Märkel, 1845)." figureDoi="http://doi.org/10.5281/zenodo.10424520" httpUri="https://zenodo.org/record/10424520/files/figure.png" pageId="20" pageNumber="21">Fig. 4E</figureCitation>
). Male last visible ventrite with a slight emargination and bordered by a brush of short setae.
</paragraph>
<paragraph id="EF1536884D110443F5C1B6B563DFFECE" blockId="20.[189,1398,1963,2025]" lastBlockId="21.[189,1398,265,364]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
Median lobe of aedeagus sinuous and narrow from the first third of its length in ventral view, tapering and acutely pointed distally (
<figureCitation id="77912A0D4D110442F75EB6D065EBF84C" box="[546,660,1998,2025]" captionStart="Fig" captionStartId="53.[189,232,1056,1082]" captionTargetBox="[456,1134,285,992]" captionTargetId="figure-59@53.[422,1166,272,1017]" captionTargetPageId="53" captionText="Fig. 15. Aedeagus and paramere of Cholovocera formicaria Victor, 1838. Male from type locality (Derbent). AB. Ventral views. CD. Lateral views. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10424545" httpUri="https://zenodo.org/record/10424545/files/figure.png" pageId="20" pageNumber="21">Fig. 15A</figureCitation>
). Aedeagus in lateral view as in
<figureCitation id="77912A0D4D110442F152B6D063E3F84C" box="[1070,1180,1998,2025]" captionStart="Fig" captionStartId="53.[189,232,1056,1082]" captionTargetBox="[456,1134,285,992]" captionTargetId="figure-59@53.[422,1166,272,1017]" captionTargetPageId="53" captionText="Fig. 15. Aedeagus and paramere of Cholovocera formicaria Victor, 1838. Male from type locality (Derbent). AB. Ventral views. CD. Lateral views. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10424545" httpUri="https://zenodo.org/record/10424545/files/figure.png" pageId="20" pageNumber="21">Fig. 15C</figureCitation>
. Distal portion of paramere long, subcylindrical, sinuous, with a round apex bearing a brush of many setae (
<figureCitation id="77912A0D4D100443F07CB017678CFEED" captionStart="Fig" captionStartId="53.[189,232,1056,1082]" captionTargetBox="[456,1134,285,992]" captionTargetId="figure-59@53.[422,1166,272,1017]" captionTargetPageId="53" captionText="Fig. 15. Aedeagus and paramere of Cholovocera formicaria Victor, 1838. Male from type locality (Derbent). AB. Ventral views. CD. Lateral views. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10424545" httpUri="https://zenodo.org/record/10424545/files/figure.png" pageId="21" pageNumber="22">Figs 15B, 15D</figureCitation>
). Spermathecal duct and spermathecal reservoir short; ramus long and curved distally, cornu round and nodulus moderately developed, smaller than cornu and ramus together (
<figureCitation id="77912A0D4D100443F144B04F63ECFECE" box="[1080,1171,337,363]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[301,1319,679,1805]" captionTargetId="figure-184@13.[265,1322,663,1845]" captionTargetPageId="13" captionText="Fig. 7. Spermathecae, lateral view. A. Cholovocera afghana Johnson, 1977. B. Ch. balcánica (Karaman, 1936). C. Ch. occulta sp. nov. D. Ch. gallica (Schaufuss, 1876). E. Ch. formicaria Victor, 1838. F. Ch. formiceticola (Rosenhauer, 1856). G. Ch. punctata (Märkel, 1845). Abbreviations: See Material and methods. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424528" httpUri="https://zenodo.org/record/10424528/files/figure.png" pageId="21" pageNumber="22">Fig. 7E</figureCitation>
).
</paragraph>
<paragraph id="EF1536884D100443F5C1B08D6694FE08" blockId="21.[189,491,403,429]" box="[189,491,403,429]" pageId="21" pageNumber="22">
<heading id="B45D81E44D100443F5C1B08D6694FE08" bold="true" box="[189,491,403,429]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="DDDEEA9A4D100443F5C1B08D6694FE08" bold="true" box="[189,491,403,429]" pageId="21" pageNumber="22">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D100443F5C1B0DD65CFFDEF" blockId="21.[187,1399,451,586]" pageId="21" pageNumber="22">
The known distribution of
<taxonomicName id="28AA4D0B4D100443F487B0DD6468FE78" authorityName="Victor" authorityYear="1838" box="[507,791,451,477]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D100443F487B0DD6468FE78" box="[507,791,451,477]" italics="true" pageId="21" pageNumber="22">Cholovocera formicaria</emphasis>
</taxonomicName>
is on the Caucasus Mountains and the Caspian Sea coast, extending from eastern
<collectingCountry id="97BD76184D100443F769B0F96505FDA4" box="[533,634,487,513]" name="Turkey" pageId="21" pageNumber="22">Anatolia</collectingCountry>
to north-eastern
<collectingCountry id="97BD76184D100443F640B0F66414FDA7" box="[828,875,488,514]" name="Iran" pageId="21" pageNumber="22">Iran</collectingCountry>
(
<figureCitation id="77912A0D4D100443F606B0F964ABFDA4" box="[890,980,487,513]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[303,1293,291,1782]" captionTargetId="figure-70@18.[247,1340,265,1810]" captionTargetPageId="18" captionText="Fig. 9. Known geographical distribution of the species ofCholovocera Victor, 1838.A. Ch. formiceticola (Rosenhauer, 1856) (red dots) and Ch. gallica (Schaufuss, 1876) (yellow dots). B. Ch. punctata (Märkel, 1845) (red dots) and Ch. occulta sp. nov. (yellow dots). C. Ch. balcanica (Karaman, 1936) (red dots), Ch. attae (Kraatz, 1858) (yellow dots), Ch. formicaria (dark blue dots) and Ch. afghana Johnson, 1977 (orange dot)." figureDoi="http://doi.org/10.5281/zenodo.10424533" httpUri="https://zenodo.org/record/10424533/files/figure.png" pageId="21" pageNumber="22">Fig. 9C</figureCitation>
). There is a record from
<collectingCountry id="97BD76184D100443F197B0F96208FDA4" box="[1259,1399,487,513]" name="Switzerland" pageId="21" pageNumber="22">Switzerland</collectingCountry>
(L̂bl &amp;
<bibRefCitation id="8B3B4B794D100443F459B31566AFFD80" author="Smetana A." box="[293,464,523,549]" pageId="21" pageNumber="22" refId="ref40417" refString="L ˆ bl I. &amp; Smetana A. 2007. Catalogue of Palaearctic Coleoptera, vol. 4: Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea and Cucujoidea. Apollo Books, Stenstrup. https: // doi. org / 10.1163 / 9789004260894" type="book" year="2007">Smetana 2007</bibRefCitation>
;
<bibRefCitation id="8B3B4B794D100443F4A1B315659CFD80" author="Shockley F. W. &amp; Tomaszewska K. W. &amp; McHugh J. V." box="[477,739,523,549]" pageId="21" pageNumber="22" pagination="1 - 113" refId="ref42306" refString="Shockley F. W., Tomaszewska K. W. &amp; McHugh J. V. 2009 b. An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1 - 113. https: // doi. org / 10.11646 / zootaxa. 1999.1.1" type="journal article" year="2009">
Shockley
<emphasis id="DDDEEA9A4D100443F728B31265F1FD80" box="[596,654,523,549]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2009b
</bibRefCitation>
;
<bibRefCitation id="8B3B4B794D100443F78CB31564E7FD80" author="Rucker W. H." box="[752,920,523,549]" pageId="21" pageNumber="22" refId="ref41721" refString="Rucker W. H. 2011 b. Fauna Europaea: Coleoptera, Endomychidae: Merophysiinae. Fauna Europaea ver. 2017.06 http: // www. faunaeur. org" type="url" year="2011">Rücker 2011b</bibRefCitation>
,
<bibRefCitation id="8B3B4B794D100443F6D9B315649BFD80" author="Rucker W. H." box="[933,996,523,549]" pageId="21" pageNumber="22" pagination="1 - 40" refId="ref41772" refString="Rucker W. H. 2020. Checkliste der Latridiidae &amp; Merophysiidae der Welt. Latridiidae und Merophysiinae 22: 1 - 40." type="journal article" year="2020">2020</bibRefCitation>
), which needs to be reviewed, but we regard it as almost certainly erroneous.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D100443F5C1B36C6329FBC2" pageId="21" pageNumber="22" type="biology_ecology">
<paragraph id="EF1536884D100443F5C1B36C664CFD29" blockId="21.[189,307,626,652]" box="[189,307,626,652]" pageId="21" pageNumber="22">
<heading id="B45D81E44D100443F5C1B36C664CFD29" bold="true" box="[189,307,626,652]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="DDDEEA9A4D100443F5C1B36C664CFD29" bold="true" box="[189,307,626,652]" pageId="21" pageNumber="22">Host ants</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D100443F5C1B3BF6352FC31" blockId="21.[188,1399,673,916]" pageId="21" pageNumber="22">
Determining the identity of the species of
<vernacularName id="61A946A64D100443F7CDB3BC65AAFD19" authorityName="Latreille" authorityYear="1802" box="[689,725,674,700]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">ant</vernacularName>
hosting
<taxonomicName id="28AA4D0B4D100443F641B3BF6326FD1E" authorityName="Victor" authorityYear="1838" box="[829,1113,673,699]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D100443F641B3BF6326FD1E" box="[829,1113,673,699]" italics="true" pageId="21" pageNumber="22">Cholovocera formicaria</emphasis>
</taxonomicName>
is not an easy task. The main problem is that the name “
<taxonomicName id="28AA4D0B4D100443F73AB3DB6588FD45" authorityName="Victor" authorityYear="1838" box="[582,759,709,736]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D100443F73AB3DB6588FD45" box="[582,759,709,736]" italics="true" pageId="21" pageNumber="22">Ch. formicaria</emphasis>
</taxonomicName>
” has been incorrectly applied to almost all the other species of the genus for over 170 years (see Taxonomic history below). For example,
<bibRefCitation id="8B3B4B794D100443F1D3B3F4678AFC8D" author="Wasmann E." pageId="21" pageNumber="22" refId="ref43060" refString="Wasmann E. 1894. Kritisches Verzeichnis der myrmekophilen und termitophilen Arthropoden. Mit Angabe der Lebenweise und mit Beschreibung neuer Arten. Dames, Berlin." type="book" year="1894">Wasmann (1894: 133)</bibRefCitation>
,
<bibRefCitation id="8B3B4B794D100443F47DB2106692FC8D" author="Bernard F." box="[257,493,782,808]" pageId="21" pageNumber="22" refId="ref38498" refString="Bernard F. 1968. Les Fourmis (Hymenoptera, Formicidae) d'Europe occidentale et septentrionale. Masson, Paris." type="book" year="1968">Bernard (1968: 383)</bibRefCitation>
and
<bibRefCitation id="8B3B4B794D100443F758B2106474FC8C" author="Kistner D. H." box="[548,779,782,809]" pageId="21" pageNumber="22" pagination="124 - 127" refId="ref39834" refString="Kistner D. H. 1982. The social insects' bestiary. In: Hermann H. R. (ed.) Social Insects. Vol. 3: 124 - 127. Academic Press, New York." type="journal article" year="1982">Kistner (1982: 125)</bibRefCitation>
, among many others, cited species of
<taxonomicName id="28AA4D0B4D100443F1C2B211626BFC8D" authorityName="Forel" authorityYear="1890" box="[1214,1300,783,808]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D100443F1C2B211626BFC8D" box="[1214,1300,783,808]" italics="true" pageId="21" pageNumber="22">Messor</emphasis>
</taxonomicName>
as hosts of “
<taxonomicName id="28AA4D0B4D100443F595B22C66E9FCE9" authorityName="Victor" authorityYear="1838" box="[233,406,818,844]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D100443F595B22C66E9FCE9" box="[233,406,818,844]" italics="true" pageId="21" pageNumber="22">Ch. formicaria</emphasis>
</taxonomicName>
”, which are erroneous due to the misidentification of the beetles.
<bibRefCitation id="8B3B4B794D100443F1EEB22C6208FCE9" author="Rucker H. W." box="[1170,1399,818,844]" pageId="21" pageNumber="22" pagination="141 - 154" refId="ref41539" refString="Rucker H. W. 1980. Bestimmungstabelle der Merophysiidae des Mittelmeerraumes und der angrenzenden Gebiete (Coleoptera: Merophysiidae). Entomologische Blatter 75 (3): 141 - 154." type="journal article" year="1980">Rücker (1980: 143)</bibRefCitation>
goes even further and incorrectly associates “
<taxonomicName id="28AA4D0B4D100443F79DB24864EDFCD5" authorityName="Victor" authorityYear="1838" box="[737,914,854,880]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D100443F79DB24864EDFCD5" box="[737,914,854,880]" italics="true" pageId="21" pageNumber="22">Ch. formicaria</emphasis>
</taxonomicName>
” with
<taxonomicName id="28AA4D0B4D100443F694B24963CBFCD5" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1000,1204,854,880]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="DDDEEA9A4D100443F694B24963CBFCD5" box="[1000,1204,854,880]" italics="true" pageId="21" pageNumber="22">Messor barbarus</emphasis>
</taxonomicName>
,
<taxonomicName id="28AA4D0B4D100443F1B9B2486570FC31" authority="(Lucas, 1849)" baseAuthorityName="Lucas" baseAuthorityYear="1849" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="testaceopilosa">
<emphasis id="DDDEEA9A4D100443F1B9B248661AFC31" italics="true" pageId="21" pageNumber="22">Aphaenogaster testaceopilosa</emphasis>
(
<bibRefCitation id="8B3B4B794D100443F408B2646578FC31" author="Lucas H." box="[372,519,890,916]" pageId="21" pageNumber="22" refId="ref40515" refString="Lucas H. 1849. Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842. Zoologie: Animaux articules: Vol. 2: Insectes. Imprimerie Nationale, Paris." type="journal volume" year="1849">Lucas, 1849</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="28AA4D0B4D100443F734B2646356FC31" authority="(Fabricius, 1793)" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[584,1065,890,916]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="megacephala">
<emphasis id="DDDEEA9A4D100443F734B264642BFC31" box="[584,852,890,916]" italics="true" pageId="21" pageNumber="22">Pheidole megacephala</emphasis>
(Fabricius, 1793)
</taxonomicName>
.
</paragraph>
<paragraph id="EF1536884D100443F5C1B2A36329FBC2" blockId="21.[189,1399,957,1128]" pageId="21" pageNumber="22">
From literature reports and our own examination of specimens associated with
<vernacularName id="61A946A64D100443F10AB2A363D7FC72" authorityName="Latreille" authorityYear="1802" box="[1142,1192,957,983]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">ants</vernacularName>
, we were able to recognise two
<vernacularName id="61A946A64D100443F416B2FF66F1FC5E" authorityName="Latreille" authorityYear="1802" box="[362,398,993,1019]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">ant</vernacularName>
taxa correctly associated with
<taxonomicName id="28AA4D0B4D100443F789B2FF64DCFC5E" authorityName="Victor" authorityYear="1838" box="[757,931,993,1019]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D100443F789B2FF64DCFC5E" box="[757,931,993,1019]" italics="true" pageId="21" pageNumber="22">Ch. formicaria</emphasis>
</taxonomicName>
in its true geographic distribution: (1) a species of
<taxonomicName id="28AA4D0B4D100443F440B51866ACFBBA" authorityName="Mayr" authorityYear="1855" box="[316,467,1030,1055]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D100443F440B51866ACFBBA" box="[316,467,1030,1055]" italics="true" pageId="21" pageNumber="22">Tetramorium</emphasis>
</taxonomicName>
, based on the identification of
<specimenCount id="F9ACFD014D100443F63FB51B64B4FBBA" box="[835,971,1029,1055]" count="2" pageId="21" pageNumber="22" type="worker">two worker</specimenCount>
<vernacularName id="61A946A64D100443F6AFB51B637DFBBA" authorityName="Latreille" authorityYear="1802" box="[979,1026,1029,1055]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">ants</vernacularName>
collected with the
<typeStatus id="3011882A4D100443F199B51B622AFBBA" box="[1253,1365,1029,1055]" pageId="21" pageNumber="22" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="28AA4D0B4D100443F5C1B5376540FBE6" authorityName="Reitter" authorityYear="1887" box="[189,575,1065,1092]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="variety" species="formicaria" variety="major">
<emphasis id="DDDEEA9A4D100443F5C1B53766C1FBE6" box="[189,446,1065,1091]" italics="true" pageId="21" pageNumber="22">Coluocera formicaria</emphasis>
var.
<emphasis id="DDDEEA9A4D100443F485B5346540FBE6" box="[505,575,1066,1091]" italics="true" pageId="21" pageNumber="22">major</emphasis>
</taxonomicName>
in the Talish Region,
<collectingCountry id="97BD76184D100443F63DB53764BDFBE6" box="[833,962,1065,1091]" name="Azerbaijan" pageId="21" pageNumber="22">Azerbaijan</collectingCountry>
(see above), and (2)
<taxonomicName id="28AA4D0B4D100443F1C0B5346602FBC2" authority="(Latreille, 1798)" authorityName="P. Werner" baseAuthorityName="Latreille" baseAuthorityYear="1798" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="structor">
<emphasis id="DDDEEA9A4D100443F1C0B5346208FBE6" box="[1212,1399,1066,1091]" italics="true" pageId="21" pageNumber="22">Messor structor</emphasis>
(Latreille, 1798)
</taxonomicName>
reported by
<bibRefCitation id="8B3B4B794D100443F76DB55364DDFBCD" author="Arakelian G. &amp; Kalashian M." box="[529,930,1101,1128]" pageId="21" pageNumber="22" pagination="50 - 52" refId="ref37976" refString="Arakelian G. &amp; Kalashian M. 1993. Fauna of Armenia: the beetles (Coleoptera) living in association with ants (Hymenoptera, Formicidae). Biologicheski Zhurnal Armenii 46 (2): 50 - 52 [In Russian]." type="journal article" year="1993">Arakelian &amp; Kalashian (1993: 51)</bibRefCitation>
from
<collectingCountry id="97BD76184D100443F694B553632CFBC2" box="[1000,1107,1101,1127]" name="Armenia" pageId="21" pageNumber="22">Armenia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D100443F5C1B5916471FB7C" pageId="21" pageNumber="22" type="synonymic_list">
<paragraph id="EF1536884D100443F5C1B59166ECFB0F" blockId="21.[189,403,1167,1194]" box="[189,403,1167,1194]" pageId="21" pageNumber="22">
<heading id="B45D81E44D100443F5C1B59166ECFB0F" bold="true" box="[189,403,1167,1194]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="DDDEEA9A4D100443F5C1B59166ECFB0F" bold="true" box="[189,403,1167,1194]" pageId="21" pageNumber="22">Junior synonyms</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D100443F596B5A06471FB7C" blockId="21.[189,1399,1214,1458]" box="[234,782,1214,1241]" pageId="21" pageNumber="22">
<taxonomicName id="28AA4D0B4D100443F596B5A06471FB7C" authority="Motchoulsky, 1845" authorityName="Motchoulsky" authorityYear="1845" box="[234,782,1214,1241]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="DDDEEA9A4D100443F596B5A06471FB7C" bold="true" box="[234,782,1214,1241]" pageId="21" pageNumber="22">
<emphasis id="DDDEEA9A4D100443F596B5A06563FB7C" bold="true" box="[234,540,1214,1241]" italics="true" pageId="21" pageNumber="22">Cholovocera subterranea</emphasis>
<bibRefCitation id="8B3B4B794D100443F75FB5A16471FB7C" author="Motchoulsky V. de" box="[547,782,1215,1241]" pageId="21" pageNumber="22" pagination="3 - 127" refId="ref40872" refString="Motchoulsky V. de 1845. Remarques sur la collection de coleopteres russes. Bulletin de la Societe imperiale des Naturalistes de Moscou 18 (1): 3 - 127." type="journal article" year="1845">Motchoulsky, 1845</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D100443F5C1B5FD6674F923" pageId="21" pageNumber="22" type="description">
<paragraph id="EF1536884D100443F5C1B5FD6432FA17" blockId="21.[189,1399,1214,1458]" pageId="21" pageNumber="22">
<bibRefCitation id="8B3B4B794D100443F5C1B5FD6683FB58" author="Motchoulsky V. de" box="[189,508,1251,1278]" pageId="21" pageNumber="22" pagination="3 - 127" refId="ref40872" refString="Motchoulsky V. de 1845. Remarques sur la collection de coleopteres russes. Bulletin de la Societe imperiale des Naturalistes de Moscou 18 (1): 3 - 127." type="journal article" year="1845">Motchoulsky (1845: 111)</bibRefCitation>
described
<taxonomicName id="28AA4D0B4D100443F7F3B5FD6427FB58" authorityName="Motchoulsky" authorityYear="1845" box="[655,856,1251,1277]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="DDDEEA9A4D100443F7F3B5FD6427FB58" box="[655,856,1251,1277]" italics="true" pageId="21" pageNumber="22">Ch. subterranea</emphasis>
</taxonomicName>
from “
<collectingRegion id="2D6EF86A4D100443F6C3B5FD633DFB58" box="[959,1090,1251,1277]" country="Russia" name="Dagestan" pageId="21" pageNumber="22">Daghestan</collectingRegion>
”, distinguishing it from
<taxonomicName id="28AA4D0B4D100443F5C1B419660EFA84" authorityName="Victor" authorityYear="1838" box="[189,369,1287,1313]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D100443F5C1B419660EFA84" box="[189,369,1287,1313]" italics="true" pageId="21" pageNumber="22">Ch. formicaria</emphasis>
</taxonomicName>
by being slightly smaller, darker and brighter.
<bibRefCitation id="8B3B4B794D100443F6B5B4196209FA87" author="Gemminger M. &amp; Harold B. de" box="[969,1398,1287,1314]" pageId="21" pageNumber="22" refId="ref39469" refString="Gemminger M. &amp; Harold B. de 1868. Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. III. Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydidae, Rhysodidae, Cucujidae, Cryptophagidae, Derodontidae, Latrididae, Othnidae, Mycetophagidae, Thorictidae, Dermestidae, Byrrhidae, Georyssidae, Parnidae, Heteroceridae, Lucanidae. Sumptu E. H. Gummi, Monachii, Paris, London." type="book" year="1868">Gemminger &amp; Harold (1868: 905)</bibRefCitation>
and
<bibRefCitation id="8B3B4B794D100443F58EB435656FFAE3" author="Schaufuss L. W." box="[242,528,1323,1350]" pageId="21" pageNumber="22" pagination="412 - 414" refId="ref42080" refString="Schaufuss L. W. 1876 b. Nachtrag zu Merophysia etc. Nunquam otiosus 2: 412 - 414." type="journal article" year="1876">Schaufuss (1876b: 413)</bibRefCitation>
accepted
<taxonomicName id="28AA4D0B4D100443F7F1B4356430FAE0" authorityName="Motchoulsky" authorityYear="1845" box="[653,847,1323,1349]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="DDDEEA9A4D100443F7F1B4356430FAE0" box="[653,847,1323,1349]" italics="true" pageId="21" pageNumber="22">Ch. subterranea</emphasis>
</taxonomicName>
as a good species, without further comment. However,
<bibRefCitation id="8B3B4B794D100443F44BB4516683FACF" author="Reitter E." box="[311,508,1359,1386]" pageId="21" pageNumber="22" pagination="1 - 6" refId="ref41226" refString="Reitter E. 1877. Ueber Merophysia Luc., Coluocera Motsch. und Reitteria Leder. Mittheilungen des Munchener Entomologischen Vereins 1877: 1 - 6." type="journal article" year="1877">Reitter (1877: 5)</bibRefCitation>
found that the differences given by
<bibRefCitation id="8B3B4B794D100443F6D6B45163E3FACF" author="Motchoulsky V. de" box="[938,1180,1359,1386]" pageId="21" pageNumber="22" pagination="3 - 127" refId="ref40872" refString="Motchoulsky V. de 1845. Remarques sur la collection de coleopteres russes. Bulletin de la Societe imperiale des Naturalistes de Moscou 18 (1): 3 - 127." type="journal article" year="1845">Motchoulsky (1845)</bibRefCitation>
did not justify the recognition of
<taxonomicName id="28AA4D0B4D100443F411B46A6552FA2B" authorityName="Motchoulsky" authorityYear="1845" box="[365,557,1396,1422]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="DDDEEA9A4D100443F411B46A6552FA2B" box="[365,557,1396,1422]" italics="true" pageId="21" pageNumber="22">Ch. subterranea</emphasis>
</taxonomicName>
as a different, valid species. We agree with
<bibRefCitation id="8B3B4B794D100443F144B46A63A0FA2B" author="Reitter E." box="[1080,1247,1396,1422]" pageId="21" pageNumber="22" pagination="1 - 6" refId="ref41226" refString="Reitter E. 1877. Ueber Merophysia Luc., Coluocera Motsch. und Reitteria Leder. Mittheilungen des Munchener Entomologischen Vereins 1877: 1 - 6." type="journal article" year="1877">Reitter (1877)</bibRefCitation>
in regarding
<taxonomicName id="28AA4D0B4D100443F5C2B4866602FA17" authorityName="Motchoulsky" authorityYear="1845" box="[190,381,1432,1458]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="DDDEEA9A4D100443F5C2B4866602FA17" box="[190,381,1432,1458]" italics="true" pageId="21" pageNumber="22">Ch. subterranea</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="28AA4D0B4D100443F7EBB486643AFA17" authorityName="Victor" authorityYear="1838" box="[663,837,1432,1458]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D100443F7EBB486643AFA17" box="[663,837,1432,1458]" italics="true" pageId="21" pageNumber="22">Ch. formicaria</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EF1536884D100443F5C1B4C56674F923" blockId="21.[188,1399,1498,1670]" pageId="21" pageNumber="22">
<bibRefCitation id="8B3B4B794D100443F5C1B4C566BBFA51" author="Wasmann E." box="[189,452,1498,1525]" pageId="21" pageNumber="22" refId="ref43060" refString="Wasmann E. 1894. Kritisches Verzeichnis der myrmekophilen und termitophilen Arthropoden. Mit Angabe der Lebenweise und mit Beschreibung neuer Arten. Dames, Berlin." type="book" year="1894">Wasmann (1894: 133)</bibRefCitation>
, still using the name
<taxonomicName id="28AA4D0B4D100443F7C3B4C46402FA51" authorityName="Motchoulsky" authorityYear="1845" box="[703,893,1498,1524]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="DDDEEA9A4D100443F7C3B4C46402FA51" box="[703,893,1498,1524]" italics="true" pageId="21" pageNumber="22">Ch. subterranea</emphasis>
</taxonomicName>
, made a doubtful association of this beetle with a species of the
<vernacularName id="61A946A64D100443F4B8B4E16697F9BC" authorityName="Latreille" authorityYear="1802" box="[452,488,1535,1561]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">ant</vernacularName>
genus
<taxonomicName id="28AA4D0B4D100443F73DB4E064F5F9BC" authority="Mayr, 1853" authorityName="Mayr" authorityYear="1853" box="[577,906,1534,1561]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D100443F73DB4E0658CF9BD" box="[577,755,1534,1560]" italics="true" pageId="21" pageNumber="22">Aphaenogaster</emphasis>
Mayr, 1853
</taxonomicName>
, without giving any details or reference. More recently,
<bibRefCitation id="8B3B4B794D100443F40DB73C65C9F998" author="Shockley F. W. &amp; Tomaszewska K. W. &amp; McHugh J. V." box="[369,694,1570,1597]" pageId="21" pageNumber="22" pagination="1 - 113" refId="ref42306" refString="Shockley F. W., Tomaszewska K. W. &amp; McHugh J. V. 2009 b. An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1 - 113. https: // doi. org / 10.11646 / zootaxa. 1999.1.1" type="journal article" year="2009">
Shockley
<emphasis id="DDDEEA9A4D100443F494B73D655BF999" box="[488,548,1570,1596]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2009b: 65)
</bibRefCitation>
and
<bibRefCitation id="8B3B4B794D100443F78DB73C64BCF998" author="Rucker W. H." box="[753,963,1570,1597]" pageId="21" pageNumber="22" pagination="1 - 15" refId="ref41629" refString="Rucker W. H. 2009. Checklist Latridiidae &amp; Merophysiinae of the World. Latridiidae und Merophysiinae 9: 1 - 15." type="journal article" year="2009">Rücker (2009: 14</bibRefCitation>
;
<bibRefCitation id="8B3B4B794D100443F6ADB73D6343F998" author="Rucker W. H." box="[977,1084,1571,1597]" pageId="21" pageNumber="22" pagination="1 - 40" refId="ref41772" refString="Rucker W. H. 2020. Checkliste der Latridiidae &amp; Merophysiidae der Welt. Latridiidae und Merophysiinae 22: 1 - 40." type="journal article" year="2020">2020: 34</bibRefCitation>
) accepted
<bibRefCitation id="8B3B4B794D100443F1C7B73C6208F998" author="Reitter E." box="[1211,1399,1570,1597]" pageId="21" pageNumber="22" pagination="1 - 6" refId="ref41226" refString="Reitter E. 1877. Ueber Merophysia Luc., Coluocera Motsch. und Reitteria Leder. Mittheilungen des Munchener Entomologischen Vereins 1877: 1 - 6." type="journal article" year="1877">Reitters (1877)</bibRefCitation>
synonymy but, without explanation, they erroneously name it as “
<taxonomicName id="28AA4D0B4D100443F6C9B7596783F923" authority="Motschulsky, 1845" authorityName="Motschulsky" authorityYear="1845" class="Insecta" family="Endomychidae" genus="Merophysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="DDDEEA9A4D100443F6C9B75963ABF9C4" box="[949,1236,1607,1633]" italics="true" pageId="21" pageNumber="22">Merophysia subterranea</emphasis>
Motschulsky, 1845
</taxonomicName>
”.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D100443F596B7B36464F962" box="[234,795,1709,1736]" pageId="21" pageNumber="22" type="synonymic_list">
<paragraph id="EF1536884D100443F596B7B36464F962" blockId="21.[188,1399,1709,2025]" box="[234,795,1709,1736]" pageId="21" pageNumber="22">
<taxonomicName id="28AA4D0B4D100443F596B7B36464F962" authority="Reitter, 1887" authorityName="Reitter" authorityYear="1887" box="[234,795,1709,1736]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="variety" species="formicaria" variety="major">
<emphasis id="DDDEEA9A4D100443F596B7B36464F962" bold="true" box="[234,795,1709,1736]" pageId="21" pageNumber="22">
<emphasis id="DDDEEA9A4D100443F596B7B3668FF962" bold="true" box="[234,496,1709,1736]" italics="true" pageId="21" pageNumber="22">Coluocera formicaria</emphasis>
var.
<emphasis id="DDDEEA9A4D100443F752B7B36509F962" bold="true" box="[558,630,1709,1735]" italics="true" pageId="21" pageNumber="22">major</emphasis>
<bibRefCitation id="8B3B4B794D100443F701B7B36464F962" author="Reitter E." box="[637,795,1709,1735]" pageId="21" pageNumber="22" refId="ref41271" refString="Reitter E. 1887. Bestimmungs-Tabellen der europaischen Coleopteren. III Helft. II Auflage. Enthaltend die Familien: Scaphidiidae, Lathridiidae und Dermestidae. E. Reitter, M ˆ dling." type="book" year="1887">Reitter, 1887</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D100440F5C1B7CF64EEFBEE" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="EF1536884D100443F5C1B7CF63F3F84D" blockId="21.[188,1399,1709,2025]" pageId="21" pageNumber="22">
<bibRefCitation id="8B3B4B794D100443F5C1B7CF66DCF94E" author="Reitter E." box="[189,419,1745,1771]" pageId="21" pageNumber="22" refId="ref41271" refString="Reitter E. 1887. Bestimmungs-Tabellen der europaischen Coleopteren. III Helft. II Auflage. Enthaltend die Familien: Scaphidiidae, Lathridiidae und Dermestidae. E. Reitter, M ˆ dling." type="book" year="1887">Reitters (1887: 10)</bibRefCitation>
description of
<taxonomicName id="28AA4D0B4D100443F72CB7CF64E1F94E" authorityName="Reitter" authorityYear="1887" box="[592,926,1745,1771]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="formicaria" subSpecies="major">
<emphasis id="DDDEEA9A4D100443F72CB7CF64E1F94E" box="[592,926,1745,1771]" italics="true" pageId="21" pageNumber="22">Coluocera formicaria major</emphasis>
</taxonomicName>
is very brief, only distinguishing it from the nominate subspecies by size.
<bibRefCitation id="8B3B4B794D100443F73CB7EB646AF8B5" author="Leder H." box="[576,789,1781,1808]" pageId="21" pageNumber="22" pagination="89 - 235" refId="ref40376" refString="Leder H. 1886. Die Coleopteren des Talysch-Gebietes, 1. In: Radde G. (ed.) Die Fauna und Flora des sudwestlichen Caspi-Gebietes: 89 - 235. F. A. Brockhaus, Leipzig." type="book chapter" year="1886">Leder (1886: 133)</bibRefCitation>
included this taxon in his catalogue, qualifying it as identical to
<taxonomicName id="28AA4D0B4D100443F43AB604668BF891" authorityName="Victor" authorityYear="1838" box="[326,500,1818,1844]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D100443F43AB604668BF891" box="[326,500,1818,1844]" italics="true" pageId="21" pageNumber="22">Ch. formicaria</emphasis>
</taxonomicName>
, but larger.
<bibRefCitation id="8B3B4B794D100443F7FCB60464E0F891" author="Heyden L. von &amp; Reitter E. &amp; Weise J." box="[640,927,1818,1844]" pageId="21" pageNumber="22" refId="ref39594" refString="Heyden L. von, Reitter E. &amp; Weise J. 1883. Catalogus coleopterorum Europae et Caucasi. 3 th ed. Libraria Nicolai, Berlin." type="book" year="1883">
Heyden
<emphasis id="DDDEEA9A4D100443F79EB6046462F891" box="[738,797,1818,1844]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(1883: 80)
</bibRefCitation>
and
<bibRefCitation id="8B3B4B794D100443F6AAB60463A5F891" author="Wasmann E." box="[982,1242,1818,1844]" pageId="21" pageNumber="22" refId="ref43060" refString="Wasmann E. 1894. Kritisches Verzeichnis der myrmekophilen und termitophilen Arthropoden. Mit Angabe der Lebenweise und mit Beschreibung neuer Arten. Dames, Berlin." type="book" year="1894">Wasmann (1894: 133)</bibRefCitation>
included
<taxonomicName id="28AA4D0B4D100443F031B60466F6F8FD" authorityName="Reitter" authorityYear="1887" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="formicaria" subSpecies="major">
<emphasis id="DDDEEA9A4D100443F031B60466F6F8FD" italics="true" pageId="21" pageNumber="22">Co. formicaria major</emphasis>
</taxonomicName>
in their catalogues, indicating its location, but without an
<vernacularName id="61A946A64D100443F138B6206317F8FD" authorityName="Latreille" authorityYear="1802" box="[1092,1128,1854,1880]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">ant</vernacularName>
association. Escherich (1897) and
<bibRefCitation id="8B3B4B794D100443F439B67C6551F8D9" author="Seidlitz G." box="[325,558,1890,1916]" pageId="21" pageNumber="22" pagination="68 - 320" refId="ref42170" refString="Seidlitz G. 1898. Bericht uber die wissenschaftlichen Leistungen im Gebiete der Entomologie wahrend des Jahres 1897, Coleoptera. Archiv fur Naturgeschichte 64 (2): 68 - 320." type="journal article" year="1898">Seidlitz (1898: 197)</bibRefCitation>
associated this subspecies with the
<vernacularName id="61A946A64D100443F6B3B67C648CF8D9" authorityName="Latreille" authorityYear="1802" box="[975,1011,1890,1916]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">ant</vernacularName>
<taxonomicName id="28AA4D0B4D100443F68BB67D63CEF8D9" authorityName="P. Werner" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[1015,1201,1891,1916]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="structor">
<emphasis id="DDDEEA9A4D100443F68BB67D63CEF8D9" box="[1015,1201,1891,1916]" italics="true" pageId="21" pageNumber="22">Messor structor</emphasis>
</taxonomicName>
in his catalogue, but the correct identity of the beetles would have been
<taxonomicName id="28AA4D0B4D100443F65EB69864BAF805" box="[802,965,1926,1952]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D100443F65EB69864BAF805" box="[802,965,1926,1952]" italics="true" pageId="21" pageNumber="22">Ch. balcanica</emphasis>
</taxonomicName>
. In his key to species of
<taxonomicName id="28AA4D0B4D100443F1A9B69866E1F861" authority=", Rucker (1980: 144" authorityName=", Rucker" authorityYear="1980" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D100443F1A9B6986212F805" box="[1237,1389,1926,1952]" italics="true" pageId="21" pageNumber="22">Cholovocera</emphasis>
,
<bibRefCitation id="8B3B4B794D100443F5C1B6B466E1F861" author="Rucker H. W." box="[189,414,1962,1988]" pageId="21" pageNumber="22" pagination="141 - 154" refId="ref41539" refString="Rucker H. W. 1980. Bestimmungstabelle der Merophysiidae des Mittelmeerraumes und der angrenzenden Gebiete (Coleoptera: Merophysiidae). Entomologische Blatter 75 (3): 141 - 154." type="journal article" year="1980">Rücker (1980: 144</bibRefCitation>
</taxonomicName>
, fig. 26) elevated this taxon to full species, but his interpretation of its
<typeStatus id="3011882A4D100443F198B6B56268F860" box="[1252,1303,1963,1989]" pageId="21" pageNumber="22">type</typeStatus>
locality was erroneous, placing the Talish Region in “
<collectingRegion id="2D6EF86A4D100443F7ACB6D06450F84D" box="[720,815,1998,2024]" country="Turkey" name="Ankara" pageId="21" pageNumber="22">Angora</collectingRegion>
”, which is actually in
<collectingCountry id="97BD76184D100443F14EB6D063F7F84D" box="[1074,1160,1998,2024]" name="Turkey" pageId="21" pageNumber="22">
<vernacularName id="61A946A64D100443F14EB6D063F7F84D" authorityName="Linnaeus" authorityYear="1758" box="[1074,1160,1998,2024]" canonicalName="Meleagris gallopavo" class="Aves" family="Phasianidae" genus="Meleagris" kingdom="Animalia" language="eng" order="Galliformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="gallopavo">Turkey</vernacularName>
</collectingCountry>
.
</paragraph>
<paragraph id="EF1536884D130440F5C1B017645AFDB9" blockId="22.[187,1400,265,540]" pageId="22" pageNumber="23">
<bibRefCitation id="8B3B4B794D130440F5C1B0176529FE81" author="Arakelian G. &amp; Kalashian M." box="[189,598,265,292]" pageId="22" pageNumber="23" pagination="50 - 52" refId="ref37976" refString="Arakelian G. &amp; Kalashian M. 1993. Fauna of Armenia: the beetles (Coleoptera) living in association with ants (Hymenoptera, Formicidae). Biologicheski Zhurnal Armenii 46 (2): 50 - 52 [In Russian]." type="journal article" year="1993">Arakelian &amp; Kalashian (1993: 51)</bibRefCitation>
cited
<emphasis id="DDDEEA9A4D130440F7D8B0176449FE86" box="[676,822,265,291]" italics="true" pageId="22" pageNumber="23">
Co. f.
<taxonomicName id="28AA4D0B4D130440F78CB0146449FE86" box="[752,822,266,291]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="major">major</taxonomicName>
</emphasis>
from
<collectingCountry id="97BD76184D130440F602B0176497FE86" box="[894,1000,265,291]" name="Armenia" pageId="22" pageNumber="23">Armenia</collectingCountry>
, around the city of Noubarashen, reporting 10 to 15 individuals inside nests of
<taxonomicName id="28AA4D0B4D130440F7BDB0336405FEE3" authorityName="P. Werner" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[705,890,301,326]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="structor">
<emphasis id="DDDEEA9A4D130440F7BDB0336405FEE3" box="[705,890,301,326]" italics="true" pageId="22" pageNumber="23">Messor structor</emphasis>
</taxonomicName>
from March to June. We have not examined these beetles, which are deposited in the Entomology Museum of the National Academy of Sciences of
<collectingCountry id="97BD76184D130440F5C1B06A665BFE2B" box="[189,292,372,398]" name="Armenia" pageId="22" pageNumber="23">Armenia</collectingCountry>
in Yereban,
<collectingCountry id="97BD76184D130440F4C8B06A6564FE2B" box="[436,539,372,398]" name="Armenia" pageId="22" pageNumber="23">Armenia</collectingCountry>
(G. Arakelian &amp; M. Kalashian pers. comm. 2020). Several checklists and catalogues (i.e., L̂bl &amp;
<bibRefCitation id="8B3B4B794D130440F4A5B08965C0FE17" author="Smetana A." box="[473,703,407,434]" pageId="22" pageNumber="23" refId="ref40417" refString="L ˆ bl I. &amp; Smetana A. 2007. Catalogue of Palaearctic Coleoptera, vol. 4: Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea and Cucujoidea. Apollo Books, Stenstrup. https: // doi. org / 10.1163 / 9789004260894" type="book" year="2007">Smetana 2007: 557</bibRefCitation>
;
<bibRefCitation id="8B3B4B794D130440F7B7B0896483FE17" author="Shockley F. W. &amp; Tomaszewska K. W. &amp; McHugh J. V." box="[715,1020,407,434]" pageId="22" pageNumber="23" pagination="1 - 113" refId="ref42306" refString="Shockley F. W., Tomaszewska K. W. &amp; McHugh J. V. 2009 b. An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1 - 113. https: // doi. org / 10.11646 / zootaxa. 1999.1.1" type="journal article" year="2009">
Shockley
<emphasis id="DDDEEA9A4D130440F63DB0866404FE14" box="[833,891,407,433]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2009b: 65
</bibRefCitation>
;
<bibRefCitation id="8B3B4B794D130440F174B08963B0FE14" author="Rucker W. H." box="[1032,1231,407,433]" pageId="22" pageNumber="23" pagination="1 - 15" refId="ref41629" refString="Rucker W. H. 2009. Checklist Latridiidae &amp; Merophysiinae of the World. Latridiidae und Merophysiinae 9: 1 - 15." type="journal article" year="2009">Rücker 2009: 14</bibRefCitation>
;
<bibRefCitation id="8B3B4B794D130440F1A0B0896258FE14" author="Rucker W. H." box="[1244,1319,407,433]" pageId="22" pageNumber="23" refId="ref41721" refString="Rucker W. H. 2011 b. Fauna Europaea: Coleoptera, Endomychidae: Merophysiinae. Fauna Europaea ver. 2017.06 http: // www. faunaeur. org" type="url" year="2011">2011b</bibRefCitation>
;
<bibRefCitation id="8B3B4B794D130440F048B08967A3FE70" author="Rucker W. H." pageId="22" pageNumber="23" pagination="1 - 40" refId="ref41772" refString="Rucker W. H. 2020. Checkliste der Latridiidae &amp; Merophysiidae der Welt. Latridiidae und Merophysiinae 22: 1 - 40." type="journal article" year="2020">2020: 34</bibRefCitation>
) still regarded
<emphasis id="DDDEEA9A4D130440F4ECB0A5655FFE70" box="[400,544,443,469]" italics="true" pageId="22" pageNumber="23">
Co. f.
<taxonomicName id="28AA4D0B4D130440F4A6B0A2655FFE70" box="[474,544,444,469]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="major">major</taxonomicName>
</emphasis>
as a full species and gave an erroneous geographical distribution, either by citing
<collectingCountry id="97BD76184D130440F451B0C066FEFE5D" box="[301,385,478,504]" name="Turkey" pageId="22" pageNumber="23">
<vernacularName id="61A946A64D130440F451B0C066FEFE5D" authorityName="Linnaeus" authorityYear="1758" box="[301,385,478,504]" canonicalName="Meleagris gallopavo" class="Aves" family="Phasianidae" genus="Meleagris" kingdom="Animalia" language="eng" order="Galliformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="gallopavo">Turkey</vernacularName>
</collectingCountry>
or by including several countries where this taxon does not occur, probably confusing it with the distribution of
<taxonomicName id="28AA4D0B4D130440F496B31C65F0FDB9" box="[490,655,514,540]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D130440F496B31C65F0FDB9" box="[490,655,514,540]" italics="true" pageId="22" pageNumber="23">Ch. balcanica</emphasis>
</taxonomicName>
(see below).
</paragraph>
<paragraph id="EF1536884D130440F5C1B35A63CBFD48" blockId="22.[187,1398,580,749]" pageId="22" pageNumber="23">
<bibRefCitation id="8B3B4B794D130440F5C1B35A66DDFDFA" author="Rucker W. H." box="[189,418,580,607]" pageId="22" pageNumber="23" refId="ref41751" refString="Rucker W. H. 2018. Latridiidae und Merophysiidae der West-Palaarktis. Selbstverlag Wolfgang H. Rucker, Neuwied." type="book" year="2018">Rücker (2018: 576</bibRefCitation>
, figs 11791180) described in detail what he believed to be “
<taxonomicName id="28AA4D0B4D130440F1F0B35A6279FDFB" box="[1164,1286,580,606]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="major">
<emphasis id="DDDEEA9A4D130440F1F0B35A6279FDFB" box="[1164,1286,580,606]" italics="true" pageId="22" pageNumber="23">Ch. major</emphasis>
</taxonomicName>
”, but his figure of the aedeagus clearly shows that it was
<taxonomicName id="28AA4D0B4D130440F795B37964F1FD24" box="[745,910,615,641]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D130440F795B37964F1FD24" box="[745,910,615,641]" italics="true" pageId="22" pageNumber="23">Ch. balcanica</emphasis>
</taxonomicName>
, associated with
<taxonomicName id="28AA4D0B4D130440F129B3766260FD24" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1109,1311,615,641]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="DDDEEA9A4D130440F129B3766260FD24" box="[1109,1311,615,641]" italics="true" pageId="22" pageNumber="23">Messor barbarus</emphasis>
</taxonomicName>
and
<taxonomicName id="28AA4D0B4D130440F02AB3766665FD00" authorityName="P. Werner" baseAuthorityName="Latreille" baseAuthorityYear="1798" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="structor">
<emphasis id="DDDEEA9A4D130440F02AB3766665FD00" italics="true" pageId="22" pageNumber="23">M. structor</emphasis>
</taxonomicName>
.
<bibRefCitation id="8B3B4B794D130440F455B3956577FD00" author="Stalling T." box="[297,520,651,677]" pageId="22" pageNumber="23" pagination="11 - 13" refId="ref42492" refString="Stalling T. 2019. Aktuelle funde einiger Merophysiinae und eine neue Wirtsameise von Reitteria escherichi Wasmann, 1896 von der insel Kos, Griechenland (Coleoptera: Merophysiidae). Folia Latridiidae et Merophysiidae 2: 11 - 13." type="journal article" year="2019">Stalling (2019: 13)</bibRefCitation>
also misidentified material of
<taxonomicName id="28AA4D0B4D130440F60FB3956367FD00" box="[883,1048,651,677]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D130440F60FB3956367FD00" box="[883,1048,651,677]" italics="true" pageId="22" pageNumber="23">Ch. balcanica</emphasis>
</taxonomicName>
from the
<collectingCountry id="97BD76184D130440F1F1B3956264FD00" box="[1165,1307,651,677]" name="Greece" pageId="22" pageNumber="23">Dodecanese</collectingCountry>
Islands (
<collectingCountry id="97BD76184D130440F5BAB3B16662FD6C" box="[198,285,687,713]" name="Greece" pageId="22" pageNumber="23">Greece</collectingCountry>
) as “
<taxonomicName id="28AA4D0B4D130440F428B3B066B5FD6D" box="[340,458,686,712]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="major">
<emphasis id="DDDEEA9A4D130440F428B3B066B5FD6D" box="[340,458,686,712]" italics="true" pageId="22" pageNumber="23">Ch. major</emphasis>
</taxonomicName>
”, an error repeated by
<bibRefCitation id="8B3B4B794D130440F7ACB3B06304FD6D" author="Lapeva-Gjonova A. &amp; Rucker W. H." box="[720,1147,686,713]" pageId="22" pageNumber="23" pagination="5 - 8" refId="ref40190" refString="Lapeva-Gjonova A. &amp; Rucker W. H. 2011. Latridiidae and Endomychidae beetles (Coleoptera) from ant nests in Bulgaria. Latridiidae 8: 5 - 8." type="journal article" year="2011">Lapeva-Gjonova &amp; Rücker (2011: 6)</bibRefCitation>
and
<bibRefCitation id="8B3B4B794D130440F1CDB3B06657FD49" author="Lapeva-Gjonova, A." pageId="22" pageNumber="23" pagination="1 - 14" refId="ref40149" refString="Lapeva-Gjonova, A. 2013. Ant-associated beetle fauna in Bulgaria: A review and new data. Psyche 2013: 1 - 14. Article ID 242037. https: // doi. org / 10.1155 / 2013 / 242037" type="journal article" year="2013">Lapeva-Gjonova (2013: 9)</bibRefCitation>
with specimens of
<taxonomicName id="28AA4D0B4D130440F771B3CC65CDFD49" box="[525,690,722,748]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D130440F771B3CC65CDFD49" box="[525,690,722,748]" italics="true" pageId="22" pageNumber="23">Ch. balcanica</emphasis>
</taxonomicName>
from
<collectingCountry id="97BD76184D130440F786B3CC641DFD49" box="[762,866,722,748]" name="Bulgaria" pageId="22" pageNumber="23">Bulgaria</collectingCountry>
, associated with
<taxonomicName id="28AA4D0B4D130440F155B3CD63D2FD49" authorityName="P. Werner" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[1065,1197,723,748]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="structor">
<emphasis id="DDDEEA9A4D130440F155B3CD63D2FD49" box="[1065,1197,723,748]" italics="true" pageId="22" pageNumber="23">M. structor</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EF1536884D130440F5C1B20A64EEFBEE" blockId="22.[189,1399,788,1099]" pageId="22" pageNumber="23">
We have examined the
<typeStatus id="3011882A4D130440F4AFB20A6545FC8B" box="[467,570,788,814]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
male and
<specimenCount id="F9ACFD014D130440F7CBB20A6423FC8B" box="[695,860,788,814]" count="2" pageId="22" pageNumber="23" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
(male and female) of
<emphasis id="DDDEEA9A4D130440F114B20A6388FC8B" box="[1128,1271,788,814]" italics="true" pageId="22" pageNumber="23">
Co. f.
<taxonomicName id="28AA4D0B4D130440F1CDB20B6388FC8B" box="[1201,1271,789,814]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="major">major</taxonomicName>
</emphasis>
, with type locality in Rasano (Talish Mountains), as well as over
<specimenCount id="F9ACFD014D130440F638B22964D1FCF4" box="[836,942,823,849]" count="20" pageId="22" pageNumber="23" type="male">20 males</specimenCount>
of
<taxonomicName id="28AA4D0B4D130440F6AAB22963FAFCF4" authorityName="Victor" authorityYear="1838" box="[982,1157,823,849]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F6AAB22963FAFCF4" box="[982,1157,823,849]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
. The body length of the
<typeStatus id="3011882A4D130440F596B245662EFCD0" box="[234,337,859,885]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
falls within the range measured for males of
<taxonomicName id="28AA4D0B4D130440F616B2456367FCD0" authorityName="Victor" authorityYear="1838" box="[874,1048,859,885]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F616B2456367FCD0" box="[874,1048,859,885]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
. Also, we have compared the morphology of the aedeagus of the
<typeStatus id="3011882A4D130440F720B26065BCFC3D" box="[604,707,894,920]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
with those of
<taxonomicName id="28AA4D0B4D130440F616B2606368FC3C" authorityName="Victor" authorityYear="1838" box="[874,1047,894,921]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F616B2606368FC3C" box="[874,1047,894,921]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
. Although the distal tip of the median lobe of the aedeagus and the paramere are missing in the
<typeStatus id="3011882A4D130440F6C0B2BC635CFC19" box="[956,1059,930,956]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(
<figureCitation id="77912A0D4D130440F14FB2BC63BFFC19" box="[1075,1216,930,956]" captionStart="Fig" captionStartId="60.[189,232,1787,1813]" captionTargetBox="[257,1331,228,1700]" captionTargetId="figure-1@60.[257,1331,225,1744]" captionTargetPageId="60" captionText="Fig. 24. A. A syntype of Cholovocera formicaria Victor, 1838, dorsal view (courtesy of K.V. Makarov). B. Same specimen, ventral view (courtesy of K.V. Makarov). C. Holotype of Ch. brevicornis Johnson, 1977 (= Displotera beloni (Wasmann, 1899)), dorsal view. D. Same specimen, ocular area, lateral view. E. Ocular area of Ch. formiceticola (Rosenhauer, 1856) (Lisbon), lateral view. F. Aedeagus of Ch. formicaria major, lectotype, ventral view. G. Same specimen, lateral view. H. Aedeagus of Co punctata sardoa, lectotype, ventral view. I. Same specimen, lateral view. Scale bars: AC = 0.5 mm; DI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424576" httpUri="https://zenodo.org/record/10424576/files/figure.png" pageId="22" pageNumber="23">Fig. 24FG</figureCitation>
), the remaining parts are identical to those of the many males of
<taxonomicName id="28AA4D0B4D130440F780B2D864D5FC45" authorityName="Victor" authorityYear="1838" box="[764,938,966,992]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F780B2D864D5FC45" box="[764,938,966,992]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
, which we have studied, also collected in the Talish Region (
<figureCitation id="77912A0D4D130440F4BBB2F76522FBA6" box="[455,605,1001,1028]" captionStart="Fig" captionStartId="53.[189,232,1056,1082]" captionTargetBox="[456,1134,285,992]" captionTargetId="figure-59@53.[422,1166,272,1017]" captionTargetPageId="53" captionText="Fig. 15. Aedeagus and paramere of Cholovocera formicaria Victor, 1838. Male from type locality (Derbent). AB. Ventral views. CD. Lateral views. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10424545" httpUri="https://zenodo.org/record/10424545/files/figure.png" pageId="22" pageNumber="23">Fig. 15A, C</figureCitation>
). Furthermore, Rasano is located approximately
<quantity id="28529B6D4D130440F1D1B2F76276FBA6" box="[1197,1289,1001,1027]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="23" unit="km" value="400.0">400 km</quantity>
south of Derbent, the type locality of
<taxonomicName id="28AA4D0B4D130440F762B51365B0FB82" authorityName="Victor" authorityYear="1838" box="[542,719,1037,1063]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F762B51365B0FB82" box="[542,719,1037,1063]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
. Therefore, we have no hesitation in placing
<taxonomicName id="28AA4D0B4D130440F187B51366F7FBEF" authorityName="Reitter" authorityYear="1887" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="formicaria" subSpecies="major">
<emphasis id="DDDEEA9A4D130440F187B51366F7FBEF" italics="true" pageId="22" pageNumber="23">Coluocera formicaria major</emphasis>
</taxonomicName>
as a new junior synonym of
<taxonomicName id="28AA4D0B4D130440F7A0B52E64F6FBEF" authorityName="Victor" authorityYear="1838" box="[732,905,1072,1098]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F7A0B52E64F6FBEF" box="[732,905,1072,1098]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D130440F5C1B56C652BFB1E" pageId="22" pageNumber="23" type="synonymic_list">
<paragraph id="EF1536884D130440F5C1B56C66A6FB29" blockId="22.[189,473,1138,1164]" box="[189,473,1138,1164]" pageId="22" pageNumber="23">
<heading id="B45D81E44D130440F5C1B56C66A6FB29" bold="true" box="[189,473,1138,1164]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="DDDEEA9A4D130440F5C1B56C66A6FB29" bold="true" box="[189,473,1138,1164]" pageId="22" pageNumber="23">Erroneous synonymies</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D130440F596B5BF652BFB1E" blockId="22.[189,1399,1184,1354]" box="[234,596,1184,1211]" pageId="22" pageNumber="23">
<emphasis id="DDDEEA9A4D130440F596B5BF652BFB1E" bold="true" box="[234,596,1184,1211]" pageId="22" pageNumber="23">
<taxonomicName id="28AA4D0B4D130440F585B5BE6539FB1E" authority="Dejean" authorityName="Dejean" box="[249,582,1184,1211]" class="Insecta" family="Cerylonidae" genus="Cerylon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="lapidarium">
<emphasis id="DDDEEA9A4D130440F585B5BE6699FB1E" bold="true" box="[249,486,1184,1211]" italics="true" pageId="22" pageNumber="23">Cerylon lapidarium</emphasis>
Dejean
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D130440F5C1B5DA622CFAEF" pageId="22" pageNumber="23" type="description">
<paragraph id="EF1536884D130440F5C1B5DA622CFAEF" blockId="22.[189,1399,1184,1354]" pageId="22" pageNumber="23">
This species has not been formally published, therefore it is not taxonomically available, i.e., it is a nomen nudum (A. Slipinski, pers. com. 2021). However,
<bibRefCitation id="8B3B4B794D130440F615B5F6633BFAA7" author="Belon M. - J." box="[873,1092,1256,1282]" pageId="22" pageNumber="23" pagination="157 - 365" refId="ref38155" refString="Belon M. - J. 1879. Famille des Lathridiens, 1 re partie. In: Mulsant E. (ed.) Histoire naturelle des coleopteres de France. Annales de la Societe linneenne de Lyon, n. s. 26: 157 - 365." type="book chapter" year="1879">Belon (1879: 194)</bibRefCitation>
regarded it as a synonym of
<taxonomicName id="28AA4D0B4D130440F5A2B41266F2FA83" authorityName="Victor" authorityYear="1838" box="[222,397,1292,1318]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F5A2B41266F2FA83" box="[222,397,1292,1318]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
. Considering the morphology of species of
<taxonomicName id="28AA4D0B4D130440F6E4B41263D3FA83" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[920,1196,1292,1318]" class="Insecta" family="Cerylonidae" genus="Cerylon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D130440F6E4B4126488FA83" box="[920,1015,1292,1318]" italics="true" pageId="22" pageNumber="23">Cerylon</emphasis>
Latreille, 1802
</taxonomicName>
(
<taxonomicName id="28AA4D0B4D130440F1C0B412622DFA83" authorityName="Billberg" authorityYear="1820" box="[1212,1362,1292,1318]" class="Insecta" family="Cerylonidae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="family">Cerylonidae</taxonomicName>
), it is possible that the material studied by
<bibRefCitation id="8B3B4B794D130440F7FBB431645DFAEC" author="Belon M. - J." box="[647,802,1327,1353]" pageId="22" pageNumber="23" pagination="157 - 365" refId="ref38155" refString="Belon M. - J. 1879. Famille des Lathridiens, 1 re partie. In: Mulsant E. (ed.) Histoire naturelle des coleopteres de France. Annales de la Societe linneenne de Lyon, n. s. 26: 157 - 365." type="book chapter" year="1879">Belon (1879)</bibRefCitation>
belonged to a species of the genus
<taxonomicName id="28AA4D0B4D130440F1BDB4316233FAEC" authorityName="Lucas" authorityYear="1852" box="[1217,1356,1327,1353]" class="Insecta" family="Endomychidae" genus="Merophysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D130440F1BDB4316233FAEC" box="[1217,1356,1327,1353]" italics="true" pageId="22" pageNumber="23">Merophysia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D130440F596B46F6505FA2E" box="[234,634,1393,1420]" pageId="22" pageNumber="23" type="synonymic_list">
<paragraph id="EF1536884D130440F596B46F6505FA2E" blockId="22.[187,1400,1393,1597]" box="[234,634,1393,1420]" pageId="22" pageNumber="23">
<taxonomicName id="28AA4D0B4D130440F596B46F6505FA2E" authority="Belon, 1895" authorityName="Belon" authorityYear="1895" box="[234,634,1393,1420]" class="Insecta" family="Endomychidae" genus="Merophysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ragusae">
<emphasis id="DDDEEA9A4D130440F596B46F6505FA2E" bold="true" box="[234,634,1393,1420]" pageId="22" pageNumber="23">
<emphasis id="DDDEEA9A4D130440F596B46F669EFA2E" bold="true" box="[234,481,1393,1420]" italics="true" pageId="22" pageNumber="23">Merophysia ragusae</emphasis>
<bibRefCitation id="8B3B4B794D130440F495B46F6505FA2E" author="Belon M. - J." box="[489,634,1393,1419]" pageId="22" pageNumber="23" pagination="75 - 105" refId="ref38354" refString="Belon M. - J. 1895. Nouvelle contribution a l'etude des Lathridiens. Annales de la Societe entomologique de Belgique 39: 75 - 105." type="journal article" year="1895">Belon, 1895</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D130440F5C1B48B64A2F998" pageId="22" pageNumber="23" type="description">
<paragraph id="EF1536884D130440F5C1B48B64A2F998" blockId="22.[187,1400,1393,1597]" pageId="22" pageNumber="23">
<bibRefCitation id="8B3B4B794D130440F5C1B48B66E3FA0A" author="Rucker W. H." box="[189,412,1429,1455]" pageId="22" pageNumber="23" pagination="1 - 40" refId="ref41772" refString="Rucker W. H. 2020. Checkliste der Latridiidae &amp; Merophysiidae der Welt. Latridiidae und Merophysiinae 22: 1 - 40." type="journal article" year="2020">Rücker (2020: 34)</bibRefCitation>
regards
<taxonomicName id="28AA4D0B4D130440F777B48B647EFA0B" authorityName="Belon" authorityYear="1895" box="[523,769,1429,1455]" class="Insecta" family="Endomychidae" genus="Merophysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ragusae">
<emphasis id="DDDEEA9A4D130440F777B48B647EFA0B" box="[523,769,1429,1455]" italics="true" pageId="22" pageNumber="23">Merophysia ragusae</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="28AA4D0B4D130440F14AB48B6397FA0A" authorityName="Victor" authorityYear="1838" box="[1078,1256,1429,1455]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F14AB48B6397FA0A" box="[1078,1256,1429,1455]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
. This is an error considering that
<bibRefCitation id="8B3B4B794D130440F4BAB4A6651BFA76" author="Belon M. - J." box="[454,612,1464,1491]" pageId="22" pageNumber="23" pagination="75 - 105" refId="ref38354" refString="Belon M. - J. 1895. Nouvelle contribution a l'etude des Lathridiens. Annales de la Societe entomologique de Belgique 39: 75 - 105." type="journal article" year="1895">Belon (1895)</bibRefCitation>
clearly places
<taxonomicName id="28AA4D0B4D130440F665B4A764DEFA77" authorityName="Belon" authorityYear="1895" box="[793,929,1465,1490]" class="Insecta" family="Endomychidae" genus="Merophysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ragusae">
<emphasis id="DDDEEA9A4D130440F665B4A764DEFA77" box="[793,929,1465,1490]" italics="true" pageId="22" pageNumber="23">M. ragusae</emphasis>
</taxonomicName>
from
<collectingRegion id="2D6EF86A4D130440F696B4A66351FA77" box="[1002,1070,1464,1490]" country="Italy" name="Sicilia" pageId="22" pageNumber="23">Sicily</collectingRegion>
close to
<taxonomicName id="28AA4D0B4D130440F1E2B4A666DFFA53" authority="Kiesenwetter, 1872" authorityName="Kiesenwetter" authorityYear="1872" class="Insecta" family="Endomychidae" genus="Merophysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sicula">
<emphasis id="DDDEEA9A4D130440F1E2B4A66207FA77" box="[1182,1400,1464,1490]" italics="true" pageId="22" pageNumber="23">Merophysia sicula</emphasis>
Kiesenwetter, 1872
</taxonomicName>
, and not to any species of
<taxonomicName id="28AA4D0B4D130440F7BBB4C26420FA53" authorityName=", Rucker" authorityYear="1980" box="[711,863,1500,1526]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D130440F7BBB4C26420FA53" box="[711,863,1500,1526]" italics="true" pageId="22" pageNumber="23">Cholovocera</emphasis>
</taxonomicName>
. Furthermore,
<bibRefCitation id="8B3B4B794D130440F170B4C26394FA53" author="Rucker W. H." box="[1036,1259,1500,1526]" pageId="22" pageNumber="23" pagination="11 - 20" refId="ref41653" refString="Rucker W. H. 2011 a. Ergebnisse von Typenstudien der Gattungen Metophthalmus Motschulsky, 1850 (Coleoptera: Latridiidae), Cholovocera Motschulsky, 1838, Reitteria Leder, 1872, Merophysia Lucas, 1852 (Endomychidae: Merophysiinae) und Holoparamecus Curtis, 1833 (Endomychidae: Holoparamecinae). Latridiidae - Mitteilungsblatt fur Systematik und Taxonomie der Latridiidae 8: 11 - 20." type="journal article" year="2011">Rücker (2011a: 18)</bibRefCitation>
had already placed
<taxonomicName id="28AA4D0B4D130440F46EB4E1657BF9BC" authorityName="Belon" authorityYear="1895" box="[274,516,1535,1561]" class="Insecta" family="Endomychidae" genus="Merophysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ragusae">
<emphasis id="DDDEEA9A4D130440F46EB4E1657BF9BC" box="[274,516,1535,1561]" italics="true" pageId="22" pageNumber="23">Merophysia ragusae</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="28AA4D0B4D130440F658B4E163AFF9BF" authority="Lucas, 1852" authorityName="Lucas" authorityYear="1852" box="[804,1232,1535,1562]" class="Insecta" family="Endomychidae" genus="Merophysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F658B4E1634BF9BC" box="[804,1076,1535,1561]" italics="true" pageId="22" pageNumber="23">Merophysia formicaria</emphasis>
Lucas, 1852
</taxonomicName>
. It is apparent that
<bibRefCitation id="8B3B4B794D130440F58DB73D66B7F998" author="Rucker W. H." box="[241,456,1571,1597]" pageId="22" pageNumber="23" pagination="1 - 40" refId="ref41772" refString="Rucker W. H. 2020. Checkliste der Latridiidae &amp; Merophysiidae der Welt. Latridiidae und Merophysiinae 22: 1 - 40." type="journal article" year="2020">Rücker (2020: 34)</bibRefCitation>
<vernacularName id="61A946A64D130440F4B3B73D6546F998" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[463,569,1571,1597]" canonicalName="Apamea furva" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" language="eng" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="furva">confused</vernacularName>
<taxonomicName id="28AA4D0B4D130440F73CB73D6591F998" authorityName="Lucas" authorityYear="1852" box="[576,750,1571,1597]" class="Insecta" family="Endomychidae" genus="Merophysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F73CB73D6591F998" box="[576,750,1571,1597]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
with
<taxonomicName id="28AA4D0B4D130440F64DB73A64AAF998" authorityName="Lucas" authorityYear="1852" box="[817,981,1571,1597]" class="Insecta" family="Endomychidae" genus="Merophysia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F64DB73A64AAF998" box="[817,981,1571,1597]" italics="true" pageId="22" pageNumber="23">M. formicaria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A7B065034D13044EF5C1B77A6213FCBF" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="discussion">
<paragraph id="EF1536884D130440F5C1B77A6532F9DA" blockId="22.[189,589,1636,1663]" box="[189,589,1636,1663]" pageId="22" pageNumber="23">
<heading id="B45D81E44D130440F5C1B77A6532F9DA" bold="true" box="[189,589,1636,1663]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="DDDEEA9A4D130440F5C1B77A6532F9DA" bold="true" box="[189,589,1636,1663]" pageId="22" pageNumber="23">Taxonomic history and remarks</emphasis>
</heading>
</paragraph>
<paragraph id="EF1536884D130440F5C1B78D63EEF899" blockId="22.[189,1399,1683,1852]" pageId="22" pageNumber="23">
Although
<bibRefCitation id="8B3B4B794D130440F44AB78D6693F90B" author="Victor T." box="[310,492,1683,1710]" pageId="22" pageNumber="23" pagination="175 - 188" refId="ref42813" refString="Victor T. 1838. Coleopteres du Caucase et des Provinces transcaucasiennes recueillis et decrits. Bulletin de la Societe imperiale des Naturalistes de Moscou 11 (2): 175 - 188." type="journal article" year="1838">Victors (1838)</bibRefCitation>
description of
<taxonomicName id="28AA4D0B4D130440F7D9B78D64BBF90B" authorityName="Victor" authorityYear="1838" box="[677,964,1683,1710]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F7D9B78D64BBF90B" box="[677,964,1683,1710]" italics="true" pageId="22" pageNumber="23">Cholovocera formicaria</emphasis>
</taxonomicName>
is very brief, his illustrations allow an accurate identification of the taxon. However, as it will be discussed below, most of the records of
<taxonomicName id="28AA4D0B4D130440F5A3B7C566F0F950" authorityName="Victor" authorityYear="1838" box="[223,399,1755,1781]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F5A3B7C566F0F950" box="[223,399,1755,1781]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
published until recently proved to be incorrect, as well as geographic distributions and names of host
<vernacularName id="61A946A64D130440F4E3B7E166B1F8BC" authorityName="Latreille" authorityYear="1802" box="[415,462,1791,1817]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="family">ants</vernacularName>
given for
<taxonomicName id="28AA4D0B4D130440F734B7E06589F8BD" authorityName="Victor" authorityYear="1838" box="[584,758,1790,1816]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F734B7E06589F8BD" box="[584,758,1790,1816]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
in several checklists. Although there are many papers dealing with
<taxonomicName id="28AA4D0B4D130440F42AB63C657BF899" authorityName="Victor" authorityYear="1838" box="[342,516,1826,1852]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F42AB63C657BF899" box="[342,516,1826,1852]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
, there is a great deal of incorrect data repeated in them.
</paragraph>
<paragraph id="EF1536884D130441F5C1B67A657AFDC3" blockId="22.[189,1398,1891,2025]" lastBlockId="23.[189,1399,265,614]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<bibRefCitation id="8B3B4B794D130440F5C1B67A66D9F8DB" author="Markel F." box="[189,422,1891,1918]" pageId="22" pageNumber="23" pagination="193 - 271" refId="ref40745" refString="Markel F. 1845. Beitrage zur Kenntniss der unter Ameisen lebenden Insekten. Zeitschrift fur die Entomologie 5: 193 - 271 [1844]." type="journal article" year="1845">Märkel (1845: 255)</bibRefCitation>
, while describing
<taxonomicName id="28AA4D0B4D130440F701B67A646BF8D8" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[637,788,1892,1918]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D130440F701B67A646BF8D8" box="[637,788,1892,1918]" italics="true" pageId="22" pageNumber="23">Ch. punctata</emphasis>
</taxonomicName>
, studied a specimen of
<taxonomicName id="28AA4D0B4D130440F156B67A63A7F8DB" authorityName="Victor" authorityYear="1838" box="[1066,1240,1892,1918]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F156B67A63A7F8DB" box="[1066,1240,1892,1918]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
from
<collectingCountry id="97BD76184D130440F05DB67A620DF8DB" box="[1313,1394,1892,1918]" name="Russia" pageId="22" pageNumber="23">Russia</collectingCountry>
, sent to him by Motschulsky (see Material examined, above).
<bibRefCitation id="8B3B4B794D130440F608B696633AF807" author="Lucas H." box="[884,1093,1927,1954]" pageId="22" pageNumber="23" refId="ref40515" refString="Lucas H. 1849. Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842. Zoologie: Animaux articules: Vol. 2: Insectes. Imprimerie Nationale, Paris." type="journal volume" year="1849">Lucas (1849: 553)</bibRefCitation>
reported “
<taxonomicName id="28AA4D0B4D130440F1C2B6996216F804" authorityName="Victor" authorityYear="1838" box="[1214,1385,1927,1953]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F1C2B6996216F804" box="[1214,1385,1927,1953]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
” from the margins of Lake
<collectingCountry id="97BD76184D130440F49FB6B56553F860" box="[483,556,1963,1989]" name="Tonga" pageId="22" pageNumber="23">Tonga</collectingCountry>
(
<collectingCountry id="97BD76184D130440F745B6B565E8F860" box="[569,663,1963,1989]" name="Algeria" pageId="22" pageNumber="23">Algeria</collectingCountry>
), on the border with
<collectingCountry id="97BD76184D130440F602B6B564A7F860" box="[894,984,1963,1989]" name="Tunisia" pageId="22" pageNumber="23">Tunisia</collectingCountry>
, but it was most likely
<taxonomicName id="28AA4D0B4D130440F1A4B6B56212F860" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[1240,1389,1963,1989]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D130440F1A4B6B56212F860" box="[1240,1389,1963,1989]" italics="true" pageId="22" pageNumber="23">Ch. punctata</emphasis>
</taxonomicName>
.
<bibRefCitation id="8B3B4B794D130440F5C1B6D0669AF84C" author="Rosenhauer W. G." box="[189,485,1998,2025]" pageId="22" pageNumber="23" refId="ref41502" refString="Rosenhauer W. G. 1856. Die Thiere Andalusiens nach dem Resultaten einer Reise zusammengestellt nebst den Beschreibungen von 249 neuen oder bis jetzt noch unbeshcreibenen Gattungen und Arten. T. Blaesing, Erlangen." type="book" year="1856">Rosenhauer (1856: 355)</bibRefCitation>
, while describing
<taxonomicName id="28AA4D0B4D130440F7B6B6D064E3F84D" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Rosenhauer" baseAuthorityYear="1856" box="[714,924,1998,2024]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D130440F7B6B6D064E3F84D" box="[714,924,1998,2024]" italics="true" pageId="22" pageNumber="23">Ch. formiceticola</emphasis>
</taxonomicName>
, compared it with
<taxonomicName id="28AA4D0B4D130440F1F1B6D06240F84D" authorityName="Victor" authorityYear="1838" box="[1165,1343,1998,2024]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D130440F1F1B6D06240F84D" box="[1165,1343,1998,2024]" italics="true" pageId="22" pageNumber="23">Ch. formicaria</emphasis>
</taxonomicName>
and
<taxonomicName id="28AA4D0B4D120441F5C1B0176629FE86" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[189,342,265,291]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F5C1B0176629FE86" box="[189,342,265,291]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
.
<bibRefCitation id="8B3B4B794D120441F41AB01765E9FE86" author="Redtenbacher L." box="[358,662,265,292]" pageId="23" pageNumber="24" refId="ref41171" refString="Redtenbacher L. 1858. Fauna Austriaca. Die Kafer. Carl Herold's Son, Wien." type="book" year="1858">Redtenbacher (1858: 380</bibRefCitation>
;
<bibRefCitation id="8B3B4B794D120441F7DFB0176465FE86" author="Redtenbacher L." box="[675,794,265,291]" pageId="23" pageNumber="24" refId="ref41188" refString="Redtenbacher L. 1874. Fauna Austriaca. Die Kafer, Vol 1. Carl Herold's Son, Wien." type="book" year="1874">1874: 411</bibRefCitation>
) discussed the differences between
<taxonomicName id="28AA4D0B4D120441F1B4B0176208FE86" authorityName="Victor" authorityYear="1838" box="[1224,1399,265,291]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F1B4B0176208FE86" box="[1224,1399,265,291]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
and
<taxonomicName id="28AA4D0B4D120441F593B03366F8FEE2" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[239,391,301,327]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F593B03366F8FEE2" box="[239,391,301,327]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
, in particular their elytral punctuation.
<bibRefCitation id="8B3B4B794D120441F62AB0336326FEE2" author="Fairmaire L." box="[854,1113,301,328]" pageId="23" pageNumber="24" pagination="266 - 269" refId="ref39294" refString="Fairmaire L. 1859. Liste des insectes recoltes en Corse par Ph. Lareynie. Annales de la Societe entomologique de France 7: 266 - 269" type="journal article" year="1859">Fairmaire (1859: 267)</bibRefCitation>
and
<bibRefCitation id="8B3B4B794D120441F1EFB033620EFEE2" author="Dieck G." box="[1171,1393,301,327]" pageId="23" pageNumber="24" pagination="397 - 404" refId="ref38861" refString="Dieck G. 1870 a. Ein entomologischer Ausflug in die Berge Sud-Corsica's. Berliner Entomologische Zeitschrift 14: 397 - 404." type="journal article" year="1870">Dieck (1870a: 399</bibRefCitation>
;
<bibRefCitation id="8B3B4B794D120441F5C1B04F6649FECE" author="Dieck G." box="[189,310,337,363]" pageId="23" pageNumber="24" pagination="201 - 204" refId="ref38906" refString="Dieck G. 1871. Coleotteri raccolti in Corsica. Bullettino della Societa entomologica Italiana 3 (2): 201 - 204." type="journal article" year="1871">1871: 202</bibRefCitation>
) recorded “
<taxonomicName id="28AA4D0B4D120441F4BCB04F6511FECE" authorityName="Victor" authorityYear="1838" box="[448,622,337,363]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F4BCB04F6511FECE" box="[448,622,337,363]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
” from
<collectingCountry id="97BD76184D120441F7B8B04F645EFECE" box="[708,801,337,363]" name="France" pageId="23" pageNumber="24">Corsica</collectingCountry>
, but these were misidentifications of
<taxonomicName id="28AA4D0B4D120441F1ABB04F6210FECF" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[1239,1391,337,363]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F1ABB04F6210FECF" box="[1239,1391,337,363]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
.
<bibRefCitation id="8B3B4B794D120441F5C1B06A66E1FE2A" author="Saulcy F. de" box="[189,414,372,399]" pageId="23" pageNumber="24" pagination="281 - 291" refId="ref41993" refString="Saulcy F. de 1862. Observations sur les genres Choleva, Catops et Catopsimorphus et remarques sur le nouveau catalogue de M. Schaum suivies de la description de deux nouveaux genres et de quatre nouvelles especes de coleopteres propres a la faune francaise. Annales de la Societe entomologique de France 2: 281 - 291." type="journal article" year="1862">Saulcy (1862: 291)</bibRefCitation>
reported “
<taxonomicName id="28AA4D0B4D120441F761B06A65B4FE2A" authorityName="Victor" authorityYear="1838" box="[541,715,372,399]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F761B06A65B4FE2A" box="[541,715,372,399]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
” from Banyuls (Southern
<collectingCountry id="97BD76184D120441F179B06B6327FE2A" box="[1029,1112,373,399]" name="France" pageId="23" pageNumber="24">France</collectingCountry>
, associated with
<taxonomicName id="28AA4D0B4D120441F05CB06B6682FE17" authority="(Latreille, 1798))" authorityName="" baseAuthorityName="Latreille" baseAuthorityYear="1798" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="capitatus">
<emphasis id="DDDEEA9A4D120441F05CB06B6656FE17" italics="true" pageId="23" pageNumber="24">Messor capitatus</emphasis>
(Latreille, 1798))
</taxonomicName>
but the correct identity of that material is uncertain because there are four species which occur in Southern
<collectingCountry id="97BD76184D120441F734B0A365E3FE72" box="[584,668,445,471]" name="France" pageId="23" pageNumber="24">France</collectingCountry>
:
<taxonomicName id="28AA4D0B4D120441F7D4B0A263CDFE73" box="[680,1202,444,471]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="variety" species="punctata" subSpecies="formiceticola" variety="gallica">
<emphasis id="DDDEEA9A4D120441F7D4B0A2643FFE73" box="[680,832,444,470]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
,
<emphasis id="DDDEEA9A4D120441F62CB0A26361FE73" box="[848,1054,444,470]" italics="true" pageId="23" pageNumber="24">Ch. formiceticola</emphasis>
,
<emphasis id="DDDEEA9A4D120441F153B0A263CDFE73" box="[1071,1202,444,470]" italics="true" pageId="23" pageNumber="24">Ch. gallica</emphasis>
</taxonomicName>
and
<taxonomicName id="28AA4D0B4D120441F193B0A26208FE73" authority="Delgado &amp; Palma, 2023" authorityName="Delgado &amp; Palma" authorityYear="2023" box="[1263,1399,444,470]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="DDDEEA9A4D120441F193B0A26208FE73" box="[1263,1399,444,470]" italics="true" pageId="23" pageNumber="24">Ch. occulta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6ED57E14D120441F5C1B0FF6666FE5E" box="[189,281,481,507]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
In their catalogue,
<bibRefCitation id="8B3B4B794D120441F77FB0FE64E1FE5E" author="Gemminger M. &amp; Harold B. de" box="[515,926,480,507]" pageId="23" pageNumber="24" refId="ref39469" refString="Gemminger M. &amp; Harold B. de 1868. Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. III. Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydidae, Rhysodidae, Cucujidae, Cryptophagidae, Derodontidae, Latrididae, Othnidae, Mycetophagidae, Thorictidae, Dermestidae, Byrrhidae, Georyssidae, Parnidae, Heteroceridae, Lucanidae. Sumptu E. H. Gummi, Monachii, Paris, London." type="book" year="1868">Gemminger &amp; Harold (1868: 905)</bibRefCitation>
gave the distribution of
<taxonomicName id="28AA4D0B4D120441F1B4B0FE6209FE5F" authorityName="Victor" authorityYear="1838" box="[1224,1398,480,506]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F1B4B0FE6209FE5F" box="[1224,1398,480,506]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
in “Grusia” (
<collectingCountry id="97BD76184D120441F421B31A66BDFDBB" box="[349,450,516,542]" name="Georgia" pageId="23" pageNumber="24">Georgia</collectingCountry>
).
<bibRefCitation id="8B3B4B794D120441F4A6B31A65B1FDBB" author="Piccioli F." box="[474,718,516,542]" pageId="23" pageNumber="24" pagination="301 - 305" refId="ref40989" refString="Piccioli F. 1871. Rivista dei coleotteri spettanti alla fauna sotterranea recentemente scoperti in Italia e descrizione di due nuove specie anottalme. Bullettino della Societa entomologica italiana 2 (4): 301 - 305." type="journal article" year="1871">Piccioli (1871: 304)</bibRefCitation>
recorded “
<taxonomicName id="28AA4D0B4D120441F626B31A6372FDBB" authorityName="Victor" authorityYear="1838" box="[858,1037,516,542]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F626B31A6372FDBB" box="[858,1037,516,542]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
” from Florence and
<collectingCountry id="97BD76184D120441F069B31A620DFDBB" box="[1301,1394,516,542]" name="France" pageId="23" pageNumber="24">Corsica</collectingCountry>
, and
<bibRefCitation id="8B3B4B794D120441F58DB3366695FDE7" author="Bargagli P." box="[241,490,552,578]" pageId="23" pageNumber="24" pagination="97 - 104" refId="ref38090" refString="Bargagli P. 1872. Materiali per la fauna entomologica dell'isola di Sardegna, coleotteri. Bullettino della Societa entomologica italiana 4 (1): 97 - 104." type="journal article" year="1872">Bargagli (1872: 100)</bibRefCitation>
from Sardinia; however, these records are likely to be misidentifications of
<taxonomicName id="28AA4D0B4D120441F5C1B3556640FDC0" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Schaufuss" baseAuthorityYear="1876" box="[189,319,587,613]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gallica">
<emphasis id="DDDEEA9A4D120441F5C1B3556640FDC0" box="[189,319,587,613]" italics="true" pageId="23" pageNumber="24">Ch. gallica</emphasis>
</taxonomicName>
or
<taxonomicName id="28AA4D0B4D120441F41AB3556681FDC0" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[358,510,587,613]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F41AB3556681FDC0" box="[358,510,587,613]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EF1536884D120441F5C1B39363E5FBDC" blockId="23.[186,1400,653,1146]" pageId="23" pageNumber="24">
<bibRefCitation id="8B3B4B794D120441F5C1B39366BBFD02" author="Schaufuss L. W." box="[189,452,653,679]" pageId="23" pageNumber="24" pagination="393 - 400" refId="ref42052" refString="Schaufuss L. W. 1876 a. Ueber Merophysia Luc., Coluocera Mtsch. und Reitteria Led. Nunquam otiosus 2: 393 - 400." type="journal article" year="1876">Schaufuss (1876a:396</bibRefCitation>
, 400) commented about the morphology of
<taxonomicName id="28AA4D0B4D120441F6D0B3936328FD0D" authorityName="Victor" authorityYear="1838" box="[940,1111,653,680]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F6D0B3936328FD0D" box="[940,1111,653,680]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
and incorrectly regarded it as an endemic of
<collectingCountry id="97BD76184D120441F4DDB3AF6687FD6E" box="[417,504,689,715]" name="Greece" pageId="23" pageNumber="24">Greece</collectingCountry>
.
<bibRefCitation id="8B3B4B794D120441F77DB3AF65A2FD6E" author="Andre E." box="[513,733,689,715]" pageId="23" pageNumber="24" pagination="152 - 235" refId="ref37904" refString="Andre E. 1874. Description des fourmis d'Europe pour servir a l'etude des insectes myrmecophiles. Revue et Magasin de Zoologie pure et appliquee 3 (2): 152 - 235." type="journal article" year="1874">André (1874: 226)</bibRefCitation>
associated
<taxonomicName id="28AA4D0B4D120441F615B3AF6368FD6E" authorityName="Victor" authorityYear="1838" box="[873,1047,689,715]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F615B3AF6368FD6E" box="[873,1047,689,715]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
with
<taxonomicName id="28AA4D0B4D120441F127B3AC625BFD6E" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1115,1316,689,715]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="DDDEEA9A4D120441F127B3AC625BFD6E" box="[1115,1316,689,715]" italics="true" pageId="23" pageNumber="24">Messor barbarus</emphasis>
</taxonomicName>
, citing
<bibRefCitation id="8B3B4B794D120441F5C1B3CB6617FD4A" author="Saulcy F. de" box="[189,360,725,751]" pageId="23" pageNumber="24" pagination="281 - 291" refId="ref41993" refString="Saulcy F. de 1862. Observations sur les genres Choleva, Catops et Catopsimorphus et remarques sur le nouveau catalogue de M. Schaum suivies de la description de deux nouveaux genres et de quatre nouvelles especes de coleopteres propres a la faune francaise. Annales de la Societe entomologique de France 2: 281 - 291." type="journal article" year="1862">Saulcy (1862)</bibRefCitation>
, who actually cited
<taxonomicName id="28AA4D0B4D120441F721B3C86456FD4A" authorityName="" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[605,809,726,751]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="capitatus">
<emphasis id="DDDEEA9A4D120441F721B3C86456FD4A" box="[605,809,726,751]" italics="true" pageId="23" pageNumber="24">Messor capitatus</emphasis>
</taxonomicName>
(see above).
<bibRefCitation id="8B3B4B794D120441F6B6B3CB63EEFD55" author="Reitter E." box="[970,1169,725,752]" pageId="23" pageNumber="24" pagination="1 - 6" refId="ref41226" refString="Reitter E. 1877. Ueber Merophysia Luc., Coluocera Motsch. und Reitteria Leder. Mittheilungen des Munchener Entomologischen Vereins 1877: 1 - 6." type="journal article" year="1877">Reitter (1877: 5)</bibRefCitation>
placed
<taxonomicName id="28AA4D0B4D120441F188B3CB6207FD4A" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Schaufuss" baseAuthorityYear="1876" box="[1268,1400,725,751]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gallica">
<emphasis id="DDDEEA9A4D120441F188B3CB6207FD4A" box="[1268,1400,725,751]" italics="true" pageId="23" pageNumber="24">Ch. gallica</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="28AA4D0B4D120441F49FB3E765EBFCB6" authorityName="Victor" authorityYear="1838" box="[483,660,761,787]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F49FB3E765EBFCB6" box="[483,660,761,787]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
, a status that we do not agree with.
<bibRefCitation id="8B3B4B794D120441F125B3E76247FCB6" author="Belon M. - J." box="[1113,1336,761,787]" pageId="23" pageNumber="24" pagination="157 - 365" refId="ref38155" refString="Belon M. - J. 1879. Famille des Lathridiens, 1 re partie. In: Mulsant E. (ed.) Histoire naturelle des coleopteres de France. Annales de la Societe linneenne de Lyon, n. s. 26: 157 - 365." type="book chapter" year="1879">Belon (1879: 191)</bibRefCitation>
took a radical view, placing all five species of
<taxonomicName id="28AA4D0B4D120441F7BDB2036426FC92" authorityName=", Rucker" authorityYear="1980" box="[705,857,797,823]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D120441F7BDB2036426FC92" box="[705,857,797,823]" italics="true" pageId="23" pageNumber="24">Cholovocera</emphasis>
</taxonomicName>
described until then, as junior synonyms of
<taxonomicName id="28AA4D0B4D120441F5C1B25F6611FCFE" authorityName="Victor" authorityYear="1838" box="[189,366,833,859]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F5C1B25F6611FCFE" box="[189,366,833,859]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
; however,
<bibRefCitation id="8B3B4B794D120441F48BB25F65A0FCFE" author="Reitter E." box="[503,735,833,859]" pageId="23" pageNumber="24" pagination="161 - 165" refId="ref41254" refString="Reitter E. 1882. Neuere literatur. Deutsche entomologische Zeitschrift 26: 161 - 165." type="journal article" year="1882">Reitter (1882: 161)</bibRefCitation>
rejected such action claiming that
<taxonomicName id="28AA4D0B4D120441F1F0B25F6258FCFF" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[1164,1319,833,859]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F1F0B25F6258FCFF" box="[1164,1319,833,859]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
was a different species from
<taxonomicName id="28AA4D0B4D120441F4B1B27A6501FCDA" authorityName="Victor" authorityYear="1838" box="[461,638,868,895]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F4B1B27A6501FCDA" box="[461,638,868,895]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
because of its obvious punctuation, a position which was later accepted by
<bibRefCitation id="8B3B4B794D120441F433B2966571FC07" author="Belon M. - J." box="[335,526,904,930]" pageId="23" pageNumber="24" pagination="1 - 17" refId="ref38205" refString="Belon M. - J. 1884 a. Supplement a la monographie des Lathridiens de France. In: Mulsant E. (ed.) Histoire naturelle des Coleopteres de France. Famille des Lathridiens 10: 1 - 17. Pritat Aine. Lyon." type="journal article" year="1884">Belon (1884a: 2</bibRefCitation>
;
<bibRefCitation id="8B3B4B794D120441F767B29665E8FC07" author="Belon M. - J." box="[539,663,904,930]" pageId="23" pageNumber="24" pagination="215 - 229" refId="ref38289" refString="Belon M. - J. 1887. Lathridiens nouveaux ou peu connus. Revue d'Entomologie 6: 215 - 229." type="journal article" year="1887">1887: 216</bibRefCitation>
).
<bibRefCitation id="8B3B4B794D120441F7D6B29664B6FC07" author="Heyden L. von &amp; Reitter E. &amp; Weise J." box="[682,969,904,930]" pageId="23" pageNumber="24" refId="ref39594" refString="Heyden L. von, Reitter E. &amp; Weise J. 1883. Catalogus coleopterorum Europae et Caucasi. 3 th ed. Libraria Nicolai, Berlin." type="book" year="1883">
Heyden
<emphasis id="DDDEEA9A4D120441F670B2976438FC07" box="[780,839,904,930]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
(1883: 80)
</bibRefCitation>
incorrectly placed
<taxonomicName id="28AA4D0B4D120441F1D6B2966227FC07" authorityName="Victor" authorityYear="1838" box="[1194,1368,904,930]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F1D6B2966227FC07" box="[1194,1368,904,930]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
in southern Europe in their catalogue.
<bibRefCitation id="8B3B4B794D120441F715B2B26438FC62" author="Bolivar I." box="[617,839,940,967]" pageId="23" pageNumber="24" pagination="48 - 55" refId="ref38547" refString="Bolivar I. 1886. Nota sobre los coleopteros recogidos en Espana y Norte de Africa. Actas de la Sociedad Espanola de Historia Natural 15: 48 - 55." type="journal article" year="1886">Bolívar (1886: 51)</bibRefCitation>
recorded samples from Alicante and Algeciras (
<collectingCountry id="97BD76184D120441F5BBB2CE6670FC4F" box="[199,271,976,1002]" name="Spain" pageId="23" pageNumber="24">Spain</collectingCountry>
),
<collectingCountry id="97BD76184D120441F45FB2CF66F2FC4E" box="[291,397,977,1003]" name="Morocco" pageId="23" pageNumber="24">Morocco</collectingCountry>
and Blidah (
<collectingCountry id="97BD76184D120441F75CB2CE6503FC4F" box="[544,636,976,1002]" name="Algeria" pageId="23" pageNumber="24">Algeria</collectingCountry>
) as “
<taxonomicName id="28AA4D0B4D120441F7C8B2CE641EFC4F" authorityName="Victor" authorityYear="1838" box="[692,865,976,1002]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F7C8B2CE641EFC4F" box="[692,865,976,1002]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
”; we have studied a great part of this material which includes
<taxonomicName id="28AA4D0B4D120441F40CB2EA6544FBAB" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Rosenhauer" baseAuthorityYear="1856" box="[368,571,1012,1038]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D120441F40CB2EA6544FBAB" box="[368,571,1012,1038]" italics="true" pageId="23" pageNumber="24">Ch. formiceticola</emphasis>
</taxonomicName>
(specimens from
<collectingCountry id="97BD76184D120441F676B2EA6432FBAB" box="[778,845,1012,1038]" name="Spain" pageId="23" pageNumber="24">Spain</collectingCountry>
and
<collectingCountry id="97BD76184D120441F6FEB2EA6491FBAB" box="[898,1006,1012,1038]" name="Morocco" pageId="23" pageNumber="24">Morocco</collectingCountry>
) and
<taxonomicName id="28AA4D0B4D120441F156B2EA63BFFBAB" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[1066,1216,1012,1038]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F156B2EA63BFFBAB" box="[1066,1216,1012,1038]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
(from
<collectingCountry id="97BD76184D120441F071B2EA6215FBAB" box="[1293,1386,1012,1038]" name="Algeria" pageId="23" pageNumber="24">Algeria</collectingCountry>
). Similarly,
<bibRefCitation id="8B3B4B794D120441F445B506654FFB97" author="Oertzen E. von" box="[313,560,1048,1074]" pageId="23" pageNumber="24" pagination="189 - 291" refId="ref40904" refString="Oertzen E. von. 1886. Verzeichniss der Coleopteren Griechenlands und Cretas. Berliner Entomologische Zeitschrift 30 (2): 189 - 291." type="journal article" year="1886">Oertzen (1886: 201)</bibRefCitation>
recorded “
<taxonomicName id="28AA4D0B4D120441F7BCB506640CFB97" authorityName="Victor" authorityYear="1838" box="[704,883,1048,1074]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F7BCB506640CFB97" box="[704,883,1048,1074]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
” from Naxos (
<collectingCountry id="97BD76184D120441F14EB50663F4FB97" box="[1074,1163,1048,1074]" name="Greece" pageId="23" pageNumber="24">Greece</collectingCountry>
), which were most likely
<taxonomicName id="28AA4D0B4D120441F474B52566D2FBF0" box="[264,429,1083,1109]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D120441F474B52566D2FBF0" box="[264,429,1083,1109]" italics="true" pageId="23" pageNumber="24">Ch. balcanica</emphasis>
</taxonomicName>
or
<taxonomicName id="28AA4D0B4D120441F4A9B525644AFBF3" authority=", Walker (1889: 377)" authorityName=", Walker" authorityPageNumber="377" authorityYear="1889" box="[469,821,1083,1110]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="attae">
<emphasis id="DDDEEA9A4D120441F4A9B525653FFBF0" box="[469,576,1083,1109]" italics="true" pageId="23" pageNumber="24">Ch. attae</emphasis>
,
<bibRefCitation id="8B3B4B794D120441F733B525644AFBF3" author="Walker J. J." box="[591,821,1083,1110]" pageId="23" pageNumber="24" pagination="374 - 378" refId="ref42974" refString="Walker J. J. 1889. Notes on ant's-nest beetles at Gibraltar and Tangier, with special reference to the Histeridae. Entomologist's Monthly Magazine 25: 374 - 378." type="journal article" year="1889">
<vernacularName id="61A946A64D120441F733B52565DCFBF0" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[591,675,1083,1109]" canonicalName="Polyphylla fullo" class="Insecta" family="Scarabaeidae" genus="Polyphylla" kingdom="Animalia" language="deu" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fullo">Walker</vernacularName>
(1889: 377)
</bibRefCitation>
</taxonomicName>
reported it from
<collectingCountry id="97BD76184D120441F683B5256316FBF0" box="[1023,1129,1083,1109]" name="Gibraltar" pageId="23" pageNumber="24">Gibraltar</collectingCountry>
and
<collectingCountry id="97BD76184D120441F1DEB5256384FBF0" box="[1186,1275,1083,1109]" name="Morocco" pageId="23" pageNumber="24">Tangier</collectingCountry>
actually
<taxonomicName id="28AA4D0B4D120441F5C1B54166F5FBDC" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Rosenhauer" baseAuthorityYear="1856" box="[189,394,1119,1145]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D120441F5C1B54166F5FBDC" box="[189,394,1119,1145]" italics="true" pageId="23" pageNumber="24">Ch. formiceticola</emphasis>
</taxonomicName>
and
<bibRefCitation id="8B3B4B794D120441F4A5B54165CFFBDC" author="Gallois M. J." box="[473,688,1119,1145]" pageId="23" pageNumber="24" pagination="17 - 19" refId="ref39380" refString="Gallois M. J. 1893. Catalogue des Coleopteres de Maine-et-Loire. Bulletin de la Societe des Sciences naturelles de l'Ouest de la France 3 (2): 17 - 19." type="journal article" year="1893">Gallois (1893: 18)</bibRefCitation>
cited it incorrectly from Nantes (
<collectingCountry id="97BD76184D120441F144B54163F0FBDC" box="[1080,1167,1119,1145]" name="France" pageId="23" pageNumber="24">France</collectingCountry>
).
</paragraph>
<paragraph id="EF1536884D120441F5C1B5BC6209FAEE" blockId="23.[188,1399,1185,1355]" pageId="23" pageNumber="24">
<bibRefCitation id="8B3B4B794D120441F5C1B5BC66BFFB19" author="Wasmann E." box="[189,448,1186,1212]" pageId="23" pageNumber="24" refId="ref43060" refString="Wasmann E. 1894. Kritisches Verzeichnis der myrmekophilen und termitophilen Arthropoden. Mit Angabe der Lebenweise und mit Beschreibung neuer Arten. Dames, Berlin." type="book" year="1894">Wasmann (1894: 133)</bibRefCitation>
summarised published associations of what he regarded as “
<taxonomicName id="28AA4D0B4D120441F1FDB5BF6251FB1E" authorityName="Victor" authorityYear="1838" box="[1153,1326,1185,1211]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F1FDB5BF6251FB1E" box="[1153,1326,1185,1211]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
” with several species of
<vernacularName id="61A946A64D120441F4E1B5D866AEFB45" authorityName="Latreille" authorityYear="1802" box="[413,465,1222,1248]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">ants</vernacularName>
: with
<taxonomicName id="28AA4D0B4D120441F763B5D86592FB7A" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[543,749,1221,1247]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="DDDEEA9A4D120441F763B5D86592FB7A" box="[543,749,1221,1247]" italics="true" pageId="23" pageNumber="24">Messor barbarus</emphasis>
</taxonomicName>
in eastern Pyrenees (from
<bibRefCitation id="8B3B4B794D120441F145B5DB63ACFB7A" author="Saulcy F. de" box="[1081,1235,1221,1247]" pageId="23" pageNumber="24" pagination="281 - 291" refId="ref41993" refString="Saulcy F. de 1862. Observations sur les genres Choleva, Catops et Catopsimorphus et remarques sur le nouveau catalogue de M. Schaum suivies de la description de deux nouveaux genres et de quatre nouvelles especes de coleopteres propres a la faune francaise. Annales de la Societe entomologique de France 2: 281 - 291." type="journal article" year="1862">Saulcy 1862</bibRefCitation>
), in
<collectingCountry id="97BD76184D120441F071B5DB6208FB7A" box="[1293,1399,1221,1247]" name="Gibraltar" pageId="23" pageNumber="24">Gibraltar</collectingCountry>
and
<collectingCountry id="97BD76184D120441F58CB5F76636FAA6" box="[240,329,1257,1283]" name="Morocco" pageId="23" pageNumber="24">Tangier</collectingCountry>
(from
<bibRefCitation id="8B3B4B794D120441F4E2B5F76548FAA6" author="Walker J. J." box="[414,567,1257,1283]" pageId="23" pageNumber="24" pagination="175 - 187" refId="ref42952" refString="Walker J. J. 1888. A year's insect-hunting at Gibraltar. Entomologist's Monthly Magazine 24: 175 - 187." type="journal article" year="1888">
<vernacularName id="61A946A64D120441F4E2B5F7668DFAA6" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[414,498,1257,1283]" canonicalName="Polyphylla fullo" class="Insecta" family="Scarabaeidae" genus="Polyphylla" kingdom="Animalia" language="deu" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fullo">Walker</vernacularName>
1888
</bibRefCitation>
) and in
<collectingCountry id="97BD76184D120441F7E0B5F7658CFAA6" box="[668,755,1257,1283]" name="Tunisia" pageId="23" pageNumber="24">Tunisia</collectingCountry>
(from
<bibRefCitation id="8B3B4B794D120441F634B5F4637BFAA6" author="Wasmann E." box="[840,1028,1257,1284]" pageId="23" pageNumber="24" pagination="297 - 304" refId="ref43029" refString="Wasmann E. 1890. Verzeichniss der von Dr. August Forel (Zurich) in Sud-Tunesien und Ost-Algerien gesammelten Ameisen-Gaste. Deutsche Entomologische Zeitschrift 2: 297 - 304." type="journal article" year="1890">Wasmann 1890</bibRefCitation>
); with a species of
<taxonomicName id="28AA4D0B4D120441F18CB5F76227FAA6" authorityName="Westwood" authorityYear="1839" box="[1264,1368,1257,1283]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D120441F18CB5F76227FAA6" box="[1264,1368,1257,1283]" italics="true" pageId="23" pageNumber="24">Pheidole</emphasis>
</taxonomicName>
in
<collectingRegion id="2D6EF86A4D120441F5C1B4136648FA82" box="[189,311,1293,1319]" country="Spain" name="Andalucia" pageId="23" pageNumber="24">Andalusia</collectingRegion>
,
<collectingCountry id="97BD76184D120441F438B41366F8FA82" box="[324,391,1293,1319]" name="Spain" pageId="23" pageNumber="24">Spain</collectingCountry>
(from
<bibRefCitation id="8B3B4B794D120441F4A7B41365D3FA8D" author="Rosenhauer W. G." box="[475,684,1293,1320]" pageId="23" pageNumber="24" refId="ref41502" refString="Rosenhauer W. G. 1856. Die Thiere Andalusiens nach dem Resultaten einer Reise zusammengestellt nebst den Beschreibungen von 249 neuen oder bis jetzt noch unbeshcreibenen Gattungen und Arten. T. Blaesing, Erlangen." type="book" year="1856">Rosenhauer 1856</bibRefCitation>
), and with
<taxonomicName id="28AA4D0B4D120441F64FB41363EAFA82" baseAuthorityName="Lucas" baseAuthorityYear="1849" box="[819,1173,1293,1319]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="testaceopilosa">
<emphasis id="DDDEEA9A4D120441F64FB41363EAFA82" box="[819,1173,1293,1319]" italics="true" pageId="23" pageNumber="24">Aphaenogaster testaceopilosa</emphasis>
</taxonomicName>
in Menton,
<collectingCountry id="97BD76184D120441F05BB4136208FA82" box="[1319,1399,1293,1319]" name="France" pageId="23" pageNumber="24">France</collectingCountry>
(from
<bibRefCitation id="8B3B4B794D120441F47AB42F66E0FAEE" author="Walker J. J." box="[262,415,1329,1355]" pageId="23" pageNumber="24" pagination="175 - 187" refId="ref42952" refString="Walker J. J. 1888. A year's insect-hunting at Gibraltar. Entomologist's Monthly Magazine 24: 175 - 187." type="journal article" year="1888">
<vernacularName id="61A946A64D120441F47AB42F6625FAEE" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[262,346,1329,1355]" canonicalName="Polyphylla fullo" class="Insecta" family="Scarabaeidae" genus="Polyphylla" kingdom="Animalia" language="deu" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fullo">Walker</vernacularName>
1888
</bibRefCitation>
). Even if the
<vernacularName id="61A946A64D120441F749B42F6526FAEE" authorityName="Latreille" authorityYear="1802" box="[565,601,1329,1355]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">ant</vernacularName>
identities were correct, the beetles were certainly not
<taxonomicName id="28AA4D0B4D120441F1BDB42F6212FAEE" authorityName="Victor" authorityYear="1838" box="[1217,1389,1329,1355]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F1BDB42F6212FAEE" box="[1217,1389,1329,1355]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EF1536884D120441F5C1B46D6490FA28" blockId="23.[189,1007,1395,1421]" box="[189,1007,1395,1421]" pageId="23" pageNumber="24">
Other reports of beetle samples misidentified as “
<taxonomicName id="28AA4D0B4D120441F67CB46D64D1FA28" authorityName="Victor" authorityYear="1838" box="[768,942,1395,1421]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F67CB46D64D1FA28" box="[768,942,1395,1421]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
” are:
</paragraph>
<paragraph id="EF1536884D12044EF5C1B4AB66CEFE12" blockId="23.[187,1400,1461,2025]" lastBlockId="24.[189,1399,265,439]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<bibRefCitation id="8B3B4B794D120441F5C1B4AB6680FA6A" author="Escherich K. &amp; Emery C." box="[189,511,1461,1487]" pageId="23" pageNumber="24" pagination="229 - 239" refId="ref39158" refString="Escherich K. &amp; Emery C. 1897. Zur kenntniss der myrmecophilen kleinasiens. I. Coleopteren. Wiener Entomologische Zeitung 16 (9): 229 - 239." type="journal article" year="1897">Escherich &amp; Emery (1897)</bibRefCitation>
cited it from “Brussa” (
<collectingRegion id="2D6EF86A4D120441F65CB4AB6415FA6A" box="[800,874,1461,1487]" country="Turkey" name="Bursa" pageId="23" pageNumber="24">Bursa</collectingRegion>
), associated with
<taxonomicName id="28AA4D0B4D120441F143B4A86385FA6A" authorityName="P. Werner" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[1087,1274,1462,1487]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="structor">
<emphasis id="DDDEEA9A4D120441F143B4A86385FA6A" box="[1087,1274,1462,1487]" italics="true" pageId="23" pageNumber="24">Messor structor</emphasis>
</taxonomicName>
, and from
<collectingRegion id="2D6EF86A4D120441F5C3B4C76668FA56" box="[191,279,1497,1523]" country="Turkey" name="Ankara" pageId="23" pageNumber="24">Angora</collectingRegion>
(both localities in
<collectingCountry id="97BD76184D120441F480B4C7651BFA56" box="[508,612,1497,1523]" name="Turkey" pageId="23" pageNumber="24">Anatolia</collectingCountry>
,
<collectingCountry id="97BD76184D120441F70FB4C765B3FA56" box="[627,716,1497,1523]" name="Turkey" pageId="23" pageNumber="24">
<vernacularName id="61A946A64D120441F70FB4C765B3FA56" authorityName="Linnaeus" authorityYear="1758" box="[627,716,1497,1523]" canonicalName="Meleagris gallopavo" class="Aves" family="Phasianidae" genus="Meleagris" kingdom="Animalia" language="eng" order="Galliformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="gallopavo">Turkey</vernacularName>
</collectingCountry>
), but the correct identity of the beetles would have been
<taxonomicName id="28AA4D0B4D120441F5C1B4E2661AF9B3" box="[189,357,1532,1558]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="balcanica">
<emphasis id="DDDEEA9A4D120441F5C1B4E2661AF9B3" box="[189,357,1532,1558]" italics="true" pageId="23" pageNumber="24">Ch. balcanica</emphasis>
</taxonomicName>
; Escherich (1897) reported it from
<collectingCountry id="97BD76184D120441F666B4E26400F9B3" box="[794,895,1532,1558]" name="Turkey" pageId="23" pageNumber="24">Anatolia</collectingCountry>
again, but as “
<taxonomicName id="28AA4D0B4D120441F140B4E2667CF99F" authorityName="Reitter" authorityYear="1887" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="variety" species="formicaria" variety="major">
<emphasis id="DDDEEA9A4D120441F140B4E2623FF9B2" box="[1084,1344,1532,1559]" italics="true" pageId="23" pageNumber="24">Coluocera formicaria</emphasis>
var.
<emphasis id="DDDEEA9A4D120441F5C1B73F667CF99F" box="[189,259,1569,1594]" italics="true" pageId="23" pageNumber="24">major</emphasis>
</taxonomicName>
” and associated with
<taxonomicName id="28AA4D0B4D120441F480B73F65BBF99F" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[508,708,1568,1594]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="DDDEEA9A4D120441F480B73F65BBF99F" box="[508,708,1568,1594]" italics="true" pageId="23" pageNumber="24">Messor barbarus</emphasis>
</taxonomicName>
;
<bibRefCitation id="8B3B4B794D120441F7AFB73E64C5F99F" author="Sahlberg J." box="[723,954,1568,1594]" pageId="23" pageNumber="24" pagination="1 - 70" refId="ref41845" refString="Sahlberg J. 1903. Coleoptera Numido-Punica mensibus Martio, Aprili et Majo 1899 in Tunisia et Algeria orientali collecta. Ofversigt af Finska vetenskaps-societetens forhandlingar 45 (19): 1 - 70." type="journal article" year="1903">Sahlberg (1903: 31)</bibRefCitation>
recorded a great number of specimens near
<collectingRegion id="2D6EF86A4D120441F58AB75A66FDF9FB" box="[246,386,1604,1630]" country="Algeria" name="Constantine" pageId="23" pageNumber="24">Constantine</collectingRegion>
(
<collectingCountry id="97BD76184D120441F4EDB75A6692F9FB" box="[401,493,1604,1630]" name="Algeria" pageId="23" pageNumber="24">Algeria</collectingCountry>
) associated with
<taxonomicName id="28AA4D0B4D120441F7C7B75B64FBF9FB" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[699,900,1604,1630]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="barbarus">
<emphasis id="DDDEEA9A4D120441F7C7B75B64FBF9FB" box="[699,900,1604,1630]" italics="true" pageId="23" pageNumber="24">Messor barbarus</emphasis>
</taxonomicName>
, but we have examined material from this locality, which is
<taxonomicName id="28AA4D0B4D120441F4ECB7766557F927" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[400,552,1640,1666]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F4ECB7766557F927" box="[400,552,1640,1666]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
;
<bibRefCitation id="8B3B4B794D120441F745B77664DAF927" author="Luigioni P. &amp; Adelchi T." box="[569,933,1640,1666]" pageId="23" pageNumber="24" pagination="43 - 93" refId="ref40614" refString="Luigioni P. &amp; Adelchi T. 1910. Coleotteri del Lazio non citati come tali nel´catalogo dei coleotteri d'Italia &quot;, Bollettino della Societa entomologica italiana 42 (1 - 4): 43 - 93." type="journal article" year="1910">Luigioni &amp; Adelchi (1910: 69)</bibRefCitation>
recorded about
<specimenCount id="F9ACFD014D120441F11BB776626AF927" box="[1127,1301,1640,1666]" count="100" pageId="23" pageNumber="24" type="generic">100 specimens</specimenCount>
under a rock in
<collectingRegion id="2D6EF86A4D120441F46BB7926625F903" box="[279,346,1676,1702]" country="Italy" name="Lazio" pageId="23" pageNumber="24">Lazio</collectingRegion>
(
<collectingCountry id="97BD76184D120441F415B79266D6F903" box="[361,425,1676,1702]" name="Italy" pageId="23" pageNumber="24">Italia</collectingCountry>
) in February, which we believe were
<taxonomicName id="28AA4D0B4D120441F621B792648AF903" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[861,1013,1676,1702]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F621B792648AF903" box="[861,1013,1676,1702]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
(see Material examined, below);
<bibRefCitation id="8B3B4B794D120441F5C1B7AE66BAF96F" author="Donisthorpe H." box="[189,453,1712,1738]" pageId="23" pageNumber="24" pagination="161 - 165" refId="ref38998" refString="Donisthorpe H. 1927. The Ants (Formicidae), and some Myrmecophiles, of Sicily. The Entomologist's Record 38: 161 - 165." type="journal article" year="1927">Donisthorpe (1927: 9)</bibRefCitation>
collected specimens in
<collectingRegion id="2D6EF86A4D120441F79AB7AE6453F96F" box="[742,812,1712,1738]" country="Italy" name="Sicilia" pageId="23" pageNumber="24">Sicily</collectingRegion>
, associated with “
<taxonomicName id="28AA4D0B4D120441F17BB7AE662AF94B" authority="Emery, 1921" authorityName="Emery" authorityYear="1921" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="atlantis" subSpecies="nylanderi">
<emphasis id="DDDEEA9A4D120441F17BB7AE6209F96F" box="[1031,1398,1712,1738]" italics="true" pageId="23" pageNumber="24">Camponotus atlantis nylanderi</emphasis>
Emery, 1921
</taxonomicName>
”, although this
<vernacularName id="61A946A64D120441F776B7CA6551F94B" authorityName="Latreille" authorityYear="1802" box="[522,558,1748,1774]" canonicalName="Formicidae" class="Insecta" family="Formicidae" kingdom="Animalia" language="eng" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">ant</vernacularName>
does occur in
<collectingRegion id="2D6EF86A4D120441F7AAB7CD6463F948" box="[726,796,1747,1773]" country="Italy" name="Sicilia" pageId="23" pageNumber="24">Sicily</collectingRegion>
, the beetles were either
<taxonomicName id="28AA4D0B4D120441F14CB7CD63CDF948" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Schaufuss" baseAuthorityYear="1876" box="[1072,1202,1747,1773]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gallica">
<emphasis id="DDDEEA9A4D120441F14CB7CD63CDF948" box="[1072,1202,1747,1773]" italics="true" pageId="23" pageNumber="24">Ch. gallica</emphasis>
</taxonomicName>
or
<taxonomicName id="28AA4D0B4D120441F1ABB7CD6210F948" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[1239,1391,1747,1773]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F1ABB7CD6210F948" box="[1239,1391,1747,1773]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
;
<bibRefCitation id="8B3B4B794D120441F5C1B7E966CFF8B7" author="Luigioni P." box="[189,432,1783,1810]" pageId="23" pageNumber="24" pagination="1 - 1159" refId="ref40583" refString="Luigioni P. 1929. I Coleotteri d'Italia. Catalogo sinonimico-topografico-bibliografico. Memorie della Pontificia Accademia delle Scienze, I Nuovi Lincei 13 (2): 1 - 1159." type="journal article" year="1929">Luigioni (1929: 528)</bibRefCitation>
reported material from
<collectingRegion id="2D6EF86A4D120441F7B8B7E96462F8B4" box="[708,797,1783,1809]" country="Italy" name="Liguria" pageId="23" pageNumber="24">Liguria</collectingRegion>
,
<collectingRegion id="2D6EF86A4D120441F65BB7E664F5F8B7" box="[807,906,1784,1810]" country="Italy" name="Toscana" pageId="23" pageNumber="24">Toscana</collectingRegion>
,
<collectingRegion id="2D6EF86A4D120441F6E8B7E9648EF8B4" box="[916,1009,1783,1809]" country="Italy" name="Umbria" pageId="23" pageNumber="24">Umbria</collectingRegion>
,
<collectingRegion id="2D6EF86A4D120441F687B7E9633EF8B4" box="[1019,1089,1783,1809]" country="Italy" name="Lazio" pageId="23" pageNumber="24">Lazio</collectingRegion>
,
<collectingRegion id="2D6EF86A4D120441F137B7E963E6F8B4" box="[1099,1177,1783,1809]" country="Italy" name="Puglia" pageId="23" pageNumber="24">Puglia</collectingRegion>
,
<collectingRegion id="2D6EF86A4D120441F1D8B7E96275F8B4" box="[1188,1290,1783,1809]" country="Italy" name="Calabria" pageId="23" pageNumber="24">Calabria</collectingRegion>
,
<collectingRegion id="2D6EF86A4D120441F069B7E96208F8B4" box="[1301,1399,1783,1809]" country="Italy" name="Sardegna" pageId="23" pageNumber="24">Sardinia</collectingRegion>
and
<collectingRegion id="2D6EF86A4D120441F58CB605664BF890" box="[240,308,1819,1845]" country="Italy" name="Sicilia" pageId="23" pageNumber="24">Sicily</collectingRegion>
(
<collectingCountry id="97BD76184D120441F43BB60566FDF890" box="[327,386,1819,1845]" name="Italy" pageId="23" pageNumber="24">Italy</collectingCountry>
), but the correct identity would have been
<taxonomicName id="28AA4D0B4D120441F6F1B605636EF890" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Schaufuss" baseAuthorityYear="1876" box="[909,1041,1819,1845]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gallica">
<emphasis id="DDDEEA9A4D120441F6F1B605636EF890" box="[909,1041,1819,1845]" italics="true" pageId="23" pageNumber="24">Ch. gallica</emphasis>
</taxonomicName>
or
<taxonomicName id="28AA4D0B4D120441F142B60563AAF890" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[1086,1237,1819,1845]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F142B60563AAF890" box="[1086,1237,1819,1845]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
;
<bibRefCitation id="8B3B4B794D120441F19BB60566A9F8FC" author="Martinez de la Escalera M." pageId="23" pageNumber="24" pagination="1 - 553" refId="ref40772" refString="Martinez de la Escalera M. 1914. Los Coleopteros de Marruecos. Trabajos del Museo Nacional de Ciencias Naturales. Serie Zoologica 2: 1 - 553." type="journal article" year="1914">Martínez de la Escalera (1914: 123)</bibRefCitation>
collected beetles from several localities in northern
<collectingCountry id="97BD76184D120441F137B65E63C7F8FF" box="[1099,1208,1856,1882]" name="Morocco" pageId="23" pageNumber="24">Morocco</collectingCountry>
, which we have examined and identified as
<taxonomicName id="28AA4D0B4D120441F486B67D65B9F8D8" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Rosenhauer" baseAuthorityYear="1856" box="[506,710,1891,1917]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D120441F486B67D65B9F8D8" box="[506,710,1891,1917]" italics="true" pageId="23" pageNumber="24">Ch. formiceticola</emphasis>
</taxonomicName>
;
<bibRefCitation id="8B3B4B794D120441F7AFB67D633FF8D8" author="Angelini F. &amp; Rucker H. W." box="[723,1088,1891,1917]" pageId="23" pageNumber="24" pagination="213 - 240" refId="ref37938" refString="Angelini F. &amp; Rucker H. W. 1999. Contributo alla conoscenza dei Merophysiidae e Lathridiidae dell'Italia meridionale e della Sicilia (Coleoptera). Memorie della Societa Entomologica Italiana 77: 213 - 240." type="journal article" year="1999">Angelini &amp; Rücker (1999: 218)</bibRefCitation>
reported “
<taxonomicName id="28AA4D0B4D120441F1C0B67D6216F8D8" authorityName="Victor" authorityYear="1838" box="[1212,1385,1891,1917]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D120441F1C0B67D6216F8D8" box="[1212,1385,1891,1917]" italics="true" pageId="23" pageNumber="24">Ch. formicaria</emphasis>
</taxonomicName>
” from Puglia, Calabria and Sicily, but it was most likely
<taxonomicName id="28AA4D0B4D120441F6FFB699635EF805" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[899,1057,1927,1953]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D120441F6FFB699635EF805" box="[899,1057,1927,1953]" italics="true" pageId="23" pageNumber="24">Ch. punctata</emphasis>
</taxonomicName>
or
<taxonomicName id="28AA4D0B4D120441F128B69963A3F804" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Schaufuss" baseAuthorityYear="1876" box="[1108,1244,1927,1953]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gallica">
<emphasis id="DDDEEA9A4D120441F128B69963A3F804" box="[1108,1244,1927,1953]" italics="true" pageId="23" pageNumber="24">Ch. gallica</emphasis>
</taxonomicName>
, equally for
<bibRefCitation id="8B3B4B794D120441F5C1B6B46530F860" author="Sabella G. &amp; Sparacio I." box="[189,591,1962,1989]" pageId="23" pageNumber="24" pagination="477 - 508" refId="ref41796" refString="Sabella G. &amp; Sparacio I. 2004. Il ruolo dei parchi siciliani nella conservazione di taxa di insetti di particolare interesse naturalistico (Insecta Coleoptera et Lepidoptera Rhopalocera). Il Naturalista Siciliano, n. s. 28 (1): 477 - 508." type="journal article" year="2004">Sabella &amp; Sparacios (2004: 498)</bibRefCitation>
report from Sicily, and for the listed record in
<bibRefCitation id="8B3B4B794D120441F1ECB6B4679BF84D" author="Fadda S. &amp; Orgeas J. &amp; Ponel P. &amp; Henry F. &amp; Buisson E. &amp; Dutoit T." pageId="23" pageNumber="24" pagination="53 - 71" refId="ref39233" refString="Fadda S., Orgeas J., Ponel P., Henry F., Buisson E. &amp; Dutoit T. 2007. Consequences de l'arret d'un paturage seculaire sur les communautes de coleopteres terricoles de pelouses seches. Ecologia Mediterranea 33: 53 - 71. https: // doi. org / 10.3406 / ecmed. 2007.1408" type="journal article" year="2007">
Fadda
<emphasis id="DDDEEA9A4D120441F19EB6B5625FF861" box="[1250,1312,1962,1988]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
(2007: 70)
</bibRefCitation>
from Provence (
<collectingCountry id="97BD76184D120441F4CEB6D06577F84D" box="[434,520,1998,2024]" name="France" pageId="23" pageNumber="24">France</collectingCountry>
), which may also refer to
<taxonomicName id="28AA4D0B4D120441F630B6D064A9F84D" authority="Delgado &amp; Palma, 2023" authorityName="Delgado &amp; Palma" authorityYear="2023" box="[844,982,1998,2024]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="DDDEEA9A4D120441F630B6D064A9F84D" box="[844,982,1998,2024]" italics="true" pageId="23" pageNumber="24">Ch. occulta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6ED57E14D120441F69DB6D16340F84C" box="[993,1087,1999,2025]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
and/or
<taxonomicName id="28AA4D0B4D120441F1E3B6D06210F84D" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Rosenhauer" baseAuthorityYear="1856" box="[1183,1391,1998,2024]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D120441F1E3B6D06210F84D" box="[1183,1391,1998,2024]" italics="true" pageId="23" pageNumber="24">Ch. formiceticola</emphasis>
</taxonomicName>
.
<bibRefCitation id="8B3B4B794D1D044EF5C1B01766EDFE81" author="Ponel P." box="[189,402,265,292]" pageId="24" pageNumber="25" pagination="245 - 256" refId="ref41027" refString="Ponel P. 2011. Chapitre 5. Analyses entomologiques (coleopteres). In: Excoffon P. (ed.) Ville et Campagne de Frejus romaine. Bibliotheque d'Archeologie mediterraneenne et africaine: 245 - 256. Errance, Aixen-Provence. https: // doi. org / 10.4000 / books. pccj. 1205" type="book chapter" year="2011">Ponel (2011: 254)</bibRefCitation>
reported “
<taxonomicName id="28AA4D0B4D1D044EF76EB01765BEFE86" authorityName="Victor" authorityYear="1838" box="[530,705,265,291]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D1D044EF76EB01765BEFE86" box="[530,705,265,291]" italics="true" pageId="24" pageNumber="25">Ch. formicaria</emphasis>
</taxonomicName>
” from Villeneuve à Fréjus, near Nice (
<collectingCountry id="97BD76184D1D044EF1EEB0176397FE86" box="[1170,1256,265,291]" name="France" pageId="24" pageNumber="25">France</collectingCountry>
), but it may have been
<taxonomicName id="28AA4D0B4D1D044EF43DB0306511FEED" box="[321,622,302,328]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="variety" species="punctata" variety="gallica">
<emphasis id="DDDEEA9A4D1D044EF43DB03066A7FEED" box="[321,472,302,328]" italics="true" pageId="24" pageNumber="25">Ch. punctata</emphasis>
,
<emphasis id="DDDEEA9A4D1D044EF496B0306511FEED" box="[490,622,302,328]" italics="true" pageId="24" pageNumber="25">Ch. gallica</emphasis>
</taxonomicName>
or
<taxonomicName id="28AA4D0B4D1D044EF7E0B030645AFEED" authority="Delgado &amp; Palma, 2023" authorityName="Delgado &amp; Palma" authorityYear="2023" box="[668,805,302,328]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="DDDEEA9A4D1D044EF7E0B030645AFEED" box="[668,805,302,328]" italics="true" pageId="24" pageNumber="25">Ch. occulta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6ED57E14D1D044EF651B03064F5FEED" box="[813,906,302,328]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
<bibRefCitation id="8B3B4B794D1D044EF6EFB0306279FEED" author="Prieto-Manzanares M." box="[915,1286,302,328]" pageId="24" pageNumber="25" pagination="407 - 482" refId="ref41085" refString="Prieto-Manzanares M. 2018. Els coleopters del delta del Llobregat. Aproximacio historica i noves aportacions. In: Germain i Otzet J. &amp; Pino i Vilalta J. (eds) Els sistemes naturals del delta del Llobregat. Treballs de la Institucio Catalana d'Historia Natural, 19: 407 - 482. Institut d'Estudis Catalans, Barcelona." type="journal article" year="2018">Prieto-Manzanares (2018: 464)</bibRefCitation>
recorded “
<taxonomicName id="28AA4D0B4D1D044EF5B6B04D6605FEC8" authorityName="Victor" authorityYear="1838" box="[202,378,339,365]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D1D044EF5B6B04D6605FEC8" box="[202,378,339,365]" italics="true" pageId="24" pageNumber="25">Ch. formicaria</emphasis>
</taxonomicName>
” in Barcelona (
<collectingCountry id="97BD76184D1D044EF746B04D65FDFEC8" box="[570,642,339,365]" name="Spain" pageId="24" pageNumber="25">Spain</collectingCountry>
), but it was either
<taxonomicName id="28AA4D0B4D1D044EF614B04D6348FEC8" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Rosenhauer" baseAuthorityYear="1856" box="[872,1079,339,365]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D1D044EF614B04D6348FEC8" box="[872,1079,339,365]" italics="true" pageId="24" pageNumber="25">Ch. formiceticola</emphasis>
</taxonomicName>
or
<taxonomicName id="28AA4D0B4D1D044EF11FB04D639AFEC8" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Schaufuss" baseAuthorityYear="1876" box="[1123,1253,339,365]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gallica">
<emphasis id="DDDEEA9A4D1D044EF11FB04D639AFEC8" box="[1123,1253,339,365]" italics="true" pageId="24" pageNumber="25">Ch. gallica</emphasis>
</taxonomicName>
.
<bibRefCitation id="8B3B4B794D1D044EF18AB04D66F8FE37" author="Parmentier T. &amp; Gaju-Ricart M. &amp; Wenseleer T. &amp; Molero-Baltanas R." pageId="24" pageNumber="25" pagination="583 - 593" refId="ref40930" refString="Parmentier T., Gaju-Ricart M., Wenseleer T. &amp; Molero-Baltanas R. 2020. Strategies of the beetle Oochrotus unicolor (Tenebrionidae) thriving in the waste dumps of seed-harvesting Messor ants (Formicidae). Ecological Entomology 45: 583 - 593. https: // doi. org / 10.1111 / een. 12832" type="journal article" year="2020">
Parmentier
<emphasis id="DDDEEA9A4D1D044EF5C3B0666785FE37" box="[191,250,376,402]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
(2020: 589)
</bibRefCitation>
reported “
<taxonomicName id="28AA4D0B4D1D044EF778B06665CDFE37" authorityName="Victor" authorityYear="1838" box="[516,690,376,402]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D1D044EF778B06665CDFE37" box="[516,690,376,402]" italics="true" pageId="24" pageNumber="25">Ch. formicaria</emphasis>
</taxonomicName>
” from Córdoba (
<collectingCountry id="97BD76184D1D044EF607B06664BEFE37" box="[891,961,376,402]" name="Spain" pageId="24" pageNumber="25">Spain</collectingCountry>
) but, again, it was a misidentification of
<taxonomicName id="28AA4D0B4D1D044EF5A1B08266D5FE13" authorityName="Delgado &amp; Palma" authorityYear="2023" baseAuthorityName="Rosenhauer" baseAuthorityYear="1856" box="[221,426,412,438]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="formiceticola">
<emphasis id="DDDEEA9A4D1D044EF5A1B08266D5FE13" box="[221,426,412,438]" italics="true" pageId="24" pageNumber="25">Ch. formiceticola</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EF1536884D1D044EF5C1B0FE645EFDCD" blockId="24.[189,1399,480,617]" pageId="24" pageNumber="25">
<bibRefCitation id="8B3B4B794D1D044EF5C1B0FE66D5FE5F" author="Rucker H. W." box="[189,426,480,506]" pageId="24" pageNumber="25" pagination="141 - 154" refId="ref41539" refString="Rucker H. W. 1980. Bestimmungstabelle der Merophysiidae des Mittelmeerraumes und der angrenzenden Gebiete (Coleoptera: Merophysiidae). Entomologische Blatter 75 (3): 141 - 154." type="journal article" year="1980">Rücker (1980: 143)</bibRefCitation>
published a key for the identification of the six species which he recognised as belonging to
<taxonomicName id="28AA4D0B4D1D044EF42BB31A6690FDBB" authorityName=", Rucker" authorityYear="1980" box="[343,495,516,542]" class="Insecta" family="Endomychidae" genus="Cholovocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="DDDEEA9A4D1D044EF42BB31A6690FDBB" box="[343,495,516,542]" italics="true" pageId="24" pageNumber="25">Cholovocera</emphasis>
</taxonomicName>
, illustrating the median lobe of the aedeagus of all the species (
<bibRefCitation id="8B3B4B794D1D044EF1AAB31A6794FDE1" author="Rucker H. W." pageId="24" pageNumber="25" pagination="141 - 154" refId="ref41539" refString="Rucker H. W. 1980. Bestimmungstabelle der Merophysiidae des Mittelmeerraumes und der angrenzenden Gebiete (Coleoptera: Merophysiidae). Entomologische Blatter 75 (3): 141 - 154." type="journal article" year="1980">Rücker 1980: 145</bibRefCitation>
, figs 2126), including that of
<taxonomicName id="28AA4D0B4D1D044EF724B3376479FDE6" authorityName="Victor" authorityYear="1838" box="[600,774,553,579]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D1D044EF724B3376479FDE6" box="[600,774,553,579]" italics="true" pageId="24" pageNumber="25">Ch. formicaria</emphasis>
</taxonomicName>
; however, the geographic distribution of this species is erroneous, and was repeated by
<bibRefCitation id="8B3B4B794D1D044EF72DB3506463FDCD" author="Rucker W. H." box="[593,796,590,616]" pageId="24" pageNumber="25" pagination="1 - 73" refId="ref41572" refString="Rucker W. H. 1983. Kul ˆ nb ˆ zi csapu bogarak VI. - Diversicornia VI. Bunkoscsapu bogarak VII. - Clavicornia VII. In: Zoltan K. (ed.) Magyarorszag Allatvilaga (Fauna Hungariae) 8 (3): 1 - 73. Akademiai Kiado, Budapest." type="journal article" year="1983">Rücker (1983: 3)</bibRefCitation>
.
</paragraph>
<paragraph id="EF1536884D1D044EF5C1B38F6213FCBF" blockId="24.[188,1399,657,795]" pageId="24" pageNumber="25">
Several recent checklists included
<taxonomicName id="28AA4D0B4D1D044EF712B38F645DFD09" authorityName="Victor" authorityYear="1838" box="[622,802,657,684]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D1D044EF712B38F645DFD09" box="[622,802,657,684]" italics="true" pageId="24" pageNumber="25">Ch. formicaria</emphasis>
</taxonomicName>
, but repeated incorrect geographic distributions published previously (
<bibRefCitation id="8B3B4B794D1D044EF4C3B3A865FBFD75" author="Rucker W. H." box="[447,644,694,720]" pageId="24" pageNumber="25" pagination="1 - 15" refId="ref41629" refString="Rucker W. H. 2009. Checklist Latridiidae &amp; Merophysiinae of the World. Latridiidae und Merophysiinae 9: 1 - 15." type="journal article" year="2009">Rücker 2009: 14</bibRefCitation>
, 2008: 576, 2020: 34;
<bibRefCitation id="8B3B4B794D1D044EF6F9B3A863CFFD74" author="Shockley F. W. &amp; Tomaszewska K. W. &amp; McHugh J. V." box="[901,1200,694,721]" pageId="24" pageNumber="25" pagination="1 - 113" refId="ref42306" refString="Shockley F. W., Tomaszewska K. W. &amp; McHugh J. V. 2009 b. An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1 - 113. https: // doi. org / 10.11646 / zootaxa. 1999.1.1" type="journal article" year="2009">
Shockley
<emphasis id="DDDEEA9A4D1D044EF68BB3A9634FFD75" box="[1015,1072,694,720]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2009b: 65
</bibRefCitation>
). Finally,
<bibRefCitation id="8B3B4B794D1D044EF05EB3A8663DFD53" author="Rucker W. H." pageId="24" pageNumber="25" refId="ref41751" refString="Rucker W. H. 2018. Latridiidae und Merophysiidae der West-Palaarktis. Selbstverlag Wolfgang H. Rucker, Neuwied." type="book" year="2018">Rücker (2018: 576</bibRefCitation>
, figs 11831184) gave a detailed description of
<taxonomicName id="28AA4D0B4D1D044EF6E9B3C56339FD50" authorityName="Victor" authorityYear="1838" box="[917,1094,731,757]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="formicaria">
<emphasis id="DDDEEA9A4D1D044EF6E9B3C56339FD50" box="[917,1094,731,757]" italics="true" pageId="24" pageNumber="25">Ch. formicaria</emphasis>
</taxonomicName>
, including figures of the aedeagus; however, we believe these figures actually represent the aedeagus of
<taxonomicName id="28AA4D0B4D1D044EF11CB21E6388FCBF" baseAuthorityName="Markel" baseAuthorityYear="1845" box="[1120,1271,768,794]" class="Insecta" family="Endomychidae" genus="Coluocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="DDDEEA9A4D1D044EF11CB21E6388FCBF" box="[1120,1271,768,794]" italics="true" pageId="24" pageNumber="25">Ch. punctata</emphasis>
</taxonomicName>
(
<figureCitation id="77912A0D4D1D044EF07BB21E6220FCBF" box="[1287,1375,768,794]" captionStart="Fig" captionStartId="53.[189,232,1963,1989]" captionTargetBox="[461,1136,1200,1890]" captionTargetId="figure-61@53.[393,1194,1169,1928]" captionTargetPageId="53" captionText="Fig. 16. Aedeagus and paramere of Cholovocera punctata (Märkel, 1845). Lectotype (Sicily). A B. Ventral views. C. Lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10424547" httpUri="https://zenodo.org/record/10424547/files/figure.png" pageId="24" pageNumber="25">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>