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<mods:title id="BF30E0EAA95FC7E7AF93FE67C0AB5FE2">Synopsis of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae)</mods:title>
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<mods:namePart id="EF6DE8432A0E010B54DF1A351C05B779">Moore, Matthew R.</mods:namePart>
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<mods:namePart id="7FA4DF29074D5396C86D9DE332B5E68E">Cave, Ronald D.</mods:namePart>
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<taxonomicName id="5DB3FA9722B4C28C1AB4434A9A79E0E6" ID-CoL="35M7" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Augoderia Burmeister, 1847</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="DAD1690F98F884CB414383497AC7D953" pageId="44" pageNumber="45">Type species.</paragraph>
<paragraph id="257A1814607CF85784BB8264D82A56EF" pageId="44" pageNumber="45">
<taxonomicName id="BB4C34AF549CE21CA5BC706BCBC9AB98" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia nitidula" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="nitidula">Augoderia nitidula</taxonomicName>
Burmeister, 1847: 34, by monotypy.
</paragraph>
</subSubSection>
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<paragraph id="A5F62A01DA39A139D8CDE1A602DA7F85" pageId="44" pageNumber="45">Valid taxa.</paragraph>
<paragraph id="B6B3B23E513A537CBC7887725F6B4711" pageId="44" pageNumber="45">Five species and subspecies.</paragraph>
<paragraph id="B70569369E4F9771169301AEDE2C5250" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
The five species and subspecies of
<taxonomicName id="1F5CCE5D26DB2B4B35B0325CD9A10848" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
are distributed in Argentina, Bolivia, Brazil, French Guiana, Peru, and Venezuela (
<bibRefCitation id="72FAB33EF72802D9933D3C4FF747302F" author="Burmeister, H" journalOrPublisher="TCF Enslin, Berlin" pageId="66" pageNumber="67" title="Handbuch der Entomologie, volume 5." year="1847">Burmeister 1847</bibRefCitation>
,
<bibRefCitation id="3789ECDFA6ED37229ACCEC4547230DAC" pageId="44" pageNumber="45">Harold 1869b</bibRefCitation>
,
<bibRefCitation id="E169DDB9E6B2F646D1FC958EB7710750" pageId="44" pageNumber="45">Arrow 1937b</bibRefCitation>
,
<bibRefCitation id="33AE51F0689772107E80FF5656842296" author="Blackwelder, RE" journalOrPublisher="Bulletin of the United States National Museum" pageId="64" pageNumber="65" pagination="189 - 341" title="Checklist of coleopterous insects of Mexico, Central America, the West Indies, and South America, part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
,
<normalizedToken id="EA866384D3988AB5F274D03CBAF29A19" originalValue="Guimarães">Guimaraes</normalizedToken>
1944,
<bibRefCitation id="F42225192EC56A0EAA4472E1FD57456C" author="Martinez, A" journalOrPublisher="Neotropica" pageId="79" pageNumber="80" pagination="72 - 80" title="Algunos Dynastinae neotropicales nuevos o poco conocidos (Coleoptera)." volume="12" year="1966">
<normalizedToken id="E93AAFE4B7661BFF3678F6C6E7C62EA0" originalValue="Martínez">Martinez</normalizedToken>
1966
</bibRefCitation>
,
<bibRefCitation id="89808BE4766D7A62C6399916B636350B" author="Gibbs, PE" journalOrPublisher="Ciencia e Cultura" pageId="72" pageNumber="73" pagination="1437 - 1441" title="Floral biology of Talaumaovata St. Hil. (Magnoliaceae)." volume="29" year="1977">Gibbs et al. 1977</bibRefCitation>
,
<bibRefCitation id="E31A5FCE9E6908DBDF0D168D6F96378F" author="Endrodi, S" journalOrPublisher="Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden" pageId="70" pageNumber="71" pagination="1 - 460" title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil." volume="33" year="1966">
<normalizedToken id="2BEDC1CC2D6312A04979A127395BA5B4" originalValue="Endrődi">Endrodi</normalizedToken>
1966
</bibRefCitation>
,
<bibRefCitation id="CAD9F5D13A84E954A4BB8F2E4C0F9154" author="Endrodi, S" journalOrPublisher="Frey Tutzing bei Muenchen" pageId="70" pageNumber="71" pagination="406 - 411" title="Ergaenzungen zur Kenntnis der Cyclocephalini. Entomologische Arbeiten aus dem Museum G." volume="18" year="1967 a">1967a</bibRefCitation>
,
<bibRefCitation id="02A9491FC4631253C60FFA402074571E" author="Endrodi, S" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="71" pageNumber="72" pagination="197 - 202" title="Neue und seltene Dynastinen aus Suedamerika und eine synonymische Bemerkung (Coleoptera, Melolonthidae)." volume="73" year="1981">1981</bibRefCitation>
,
<bibRefCitation id="CE4482DC9B13FAED60FB700EF1124DFB" author="Endrodi, S" journalOrPublisher="Junk Publisher, Dordrecht" pageId="71" pageNumber="72" title="The Dynastinae of the World. Dr. W." year="1985 a">1985a</bibRefCitation>
,
<bibRefCitation id="CA6627DC423A7AF088E4DA057F1BD1EE" author="Riehs, PJ" journalOrPublisher="Ambiencia-Revista do Centro de Ciencias Agrarias e Ambientais" pageId="85" pageNumber="86" pagination="59 - 69" title="Similaridade entre comunidades de Dynastinae (Coleoptera, Scarabaeidae) do leste e centro-oeste do Parana: umaabordagem paleoclimatica." volume="1" year="2005">Riehs 2005</bibRefCitation>
,
<bibRefCitation id="325F756B11F1464E23C12E5A3728FC61" author="Ronqui, DC" journalOrPublisher="Iheringia, Serie Zoolgia" pageId="86" pageNumber="87" pagination="103 - 108" title="Composicao e diversidade de Scarabaeoidea (Coleoptera) atraidos por armadilha de luz em area rural norte do Parana, Brasil." url="https://doi.org/10.1590/S0073-47212006000100018" volume="96" year="2006">Ronqui and Lopes 2006</bibRefCitation>
, Ponchel 2009,
<bibRefCitation id="F50FB60FC8385CCE9F2B203611E58352" pageId="44" pageNumber="45">Grossi et al. 2011</bibRefCitation>
,
<bibRefCitation id="3A3F9758AD1907023E4A6D6037E60A50" author="Ratcliffe, BC" journalOrPublisher="Journal of the Kansas Entomogical Society" pageId="85" pageNumber="86" pagination="186 - 207" title="Beetles (Coleoptera) of Peru: A survey of families. Scarabaeoidea." url="https://doi.org/10.2317/kent-88-02-186-207.1" volume="88" year="2015">Ratcliffe et al. 2015</bibRefCitation>
) (Fig. 55).
<taxonomicName id="3966C9AF3A69397D01AA32717996DE30" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
species are similar to some
<taxonomicName id="1C173E517EA18CD796701F833DA2480F" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Cyclocephala</taxonomicName>
in overall appearance, although three taxa (
<taxonomicName id="5A979FDD737D30CA55F52E63E16D652E" lsidName="A. giuglarisi" pageId="44" pageNumber="45" rank="species" species="giuglarisi">A. giuglarisi</taxonomicName>
Ponchel,
<taxonomicName id="E262E1966D7F5EBD1E7CE6B140221F6D" lsidName="A. nitidula" pageId="44" pageNumber="45" rank="subspecies" species="nitidula" subspecies="nitidula">A. nitidula nitidula</taxonomicName>
, and
<taxonomicName id="04FC37AA4063F99FA595F0AAB09D2D8A" lsidName="A. nitidula" pageId="44" pageNumber="45" rank="subspecies" species="nitidula" subspecies="yungana">A. nitidula yungana</taxonomicName>
<normalizedToken id="175BAB6AF1FB4854B00A96383DD2CCB0" originalValue="Martínez">Martinez</normalizedToken>
) are notable for their metallic,
<pageBreakToken id="542F18658A2E3301402A79715BB3CE36" pageId="45" pageNumber="46" start="start">mother-of-pearl</pageBreakToken>
luster that reflects circularly polarized light, a cuticular trait that is rare in
<taxonomicName id="13220FB4F6C312ED2B3639817C6062C2" lsidName="" pageId="45" pageNumber="46" rank="subfamily" subfamily="Dynastinae">Dynastinae</taxonomicName>
(
<bibRefCitation id="FAD1675E588CB1F0A216B47A769245BB" author="Endrodi, S" journalOrPublisher="Frey Tutzing bei Muenchen" pageId="70" pageNumber="71" pagination="406 - 411" title="Ergaenzungen zur Kenntnis der Cyclocephalini. Entomologische Arbeiten aus dem Museum G." volume="18" year="1967 a">
<normalizedToken id="1D41C60AB01BC583BE3742A988990B9F" originalValue="Endrődi">Endrodi</normalizedToken>
1967a
</bibRefCitation>
,
<bibRefCitation id="2B82640998AA6963F8B7C84D21CB9270" author="Endrodi, S" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="71" pageNumber="72" pagination="197 - 202" title="Neue und seltene Dynastinen aus Suedamerika und eine synonymische Bemerkung (Coleoptera, Melolonthidae)." volume="73" year="1981">1981</bibRefCitation>
, Ponchel 2009,
<bibRefCitation id="7A56A3F3FEF7E109806C7993EA7721DA" author="Pye, JD" journalOrPublisher="Biological Journal of the Linnean Society" pageId="83" pageNumber="84" pagination="585 - 596" title="The distribution of circularly polarized light reflection in the Scarabaeoidea (Coleoptera)." url="https://doi.org/10.1111/j.1095-8312.2010.01449.x" volume="100" year="2010">Pye 2010</bibRefCitation>
). The biology of
<taxonomicName id="00ECE3300C8ADC1F909689F25A141500" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
species is completely unknown.
<bibRefCitation id="A30D095185180D423E536E4BCF47C500" author="Gibbs, PE" journalOrPublisher="Ciencia e Cultura" pageId="72" pageNumber="73" pagination="1437 - 1441" title="Floral biology of Talaumaovata St. Hil. (Magnoliaceae)." volume="29" year="1977">Gibbs et al. (1977)</bibRefCitation>
reported
<taxonomicName id="12A7323C5F97C1BD41089DF360BAF71D" lsidName="A. nitidula" pageId="45" pageNumber="46" rank="species" species="nitidula">A. nitidula</taxonomicName>
as a floral visitor of
<taxonomicName id="0BBC7BB5346C3B5DA8F0447AE4C70CB6" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Magnolia ovata" order="Magnoliales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="ovata">Magnolia ovata</taxonomicName>
, but this beetle was likely a misidentified
<taxonomicName id="14C37A93CD121C56C4DF7EDBE33B5B34" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Cyclocephala</taxonomicName>
species (see
<bibRefCitation id="ACE749FD8DD35FA4FDDFBCA0A03DFFA9" author="Gottsberger, G" journalOrPublisher="Flora-Morphology, Distribution, Functional Ecology of Plants" pageId="73" pageNumber="74" pagination="107 - 118" title="Pollination ecology of Magnoliaovata may explain the overall large flower size of the genus." url="https://doi.org/10.1016/j.flora.2011.11.003" volume="207" year="2012">Gottsberger et al. 2012</bibRefCitation>
,
<bibRefCitation id="094410E991E507C0FC98A21ECC491F35" author="Moore, MR" journalOrPublisher="Journal of Insect Science" pageId="80" pageNumber="81" pagination="1 - 43" title="Floral associations of cyclocephaline scarab beetles." url="https://doi.org/10.1673/031.013.10001" volume="13" year="2013">Moore and Jameson 2013</bibRefCitation>
). The immature stages are undescribed. Adults are attracted to lights at night (
<bibRefCitation id="15C93B7ED99B2E358903BCCE141D1FE6" author="Riehs, PJ" journalOrPublisher="Ambiencia-Revista do Centro de Ciencias Agrarias e Ambientais" pageId="85" pageNumber="86" pagination="59 - 69" title="Similaridade entre comunidades de Dynastinae (Coleoptera, Scarabaeidae) do leste e centro-oeste do Parana: umaabordagem paleoclimatica." volume="1" year="2005">Riehs 2005</bibRefCitation>
,
<bibRefCitation id="B9C9B2DF2BB1640872B603424115D078" author="Ronqui, DC" journalOrPublisher="Iheringia, Serie Zoolgia" pageId="86" pageNumber="87" pagination="103 - 108" title="Composicao e diversidade de Scarabaeoidea (Coleoptera) atraidos por armadilha de luz em area rural norte do Parana, Brasil." url="https://doi.org/10.1590/S0073-47212006000100018" volume="96" year="2006">Ronqui and Lopes 2006</bibRefCitation>
,
<bibRefCitation id="1190BE9B049D7B1BEF7F693F7C5EE511" pageId="45" pageNumber="46">Grossi et al. 2011</bibRefCitation>
).
</paragraph>
<caption id="A79B8E19D40D583A402A3C01DF23A857" pageId="45" pageNumber="46">
<paragraph id="A52F21F9C3F6A17055315B039CD44D39" pageId="45" pageNumber="46">
Figure 55. Country-level distribution of
<taxonomicName id="9B589E82BA5CE19634467651989B9A8C" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
taxa in South America. Numbers indicate taxa per country.
</paragraph>
</caption>
<paragraph id="E51EEA906BCFFC83C11D992B7C3CAF31" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<taxonomicName id="1C0DCCE777BEC300D6E58585C9E40CA5" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
, though maintained as a valid genus since
<bibRefCitation id="682274B2A90DD60CBD38A9F9DC19EC2D" author="Burmeister, H" journalOrPublisher="TCF Enslin, Berlin" pageId="66" pageNumber="67" title="Handbuch der Entomologie, volume 5." year="1847">Burmeister (1847)</bibRefCitation>
, is poorly defined and diagnosed in the literature. The irregularly spaced punctures of the elytra and the mother-of-pearl sheen of some taxa are the only characters historically used to separate
<taxonomicName id="48025A6DC9EF61539FD0D51791668491" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
from
<taxonomicName id="794984B1AD91AEA3F2424828572B048F" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Cyclocephala</taxonomicName>
. Thus, the genus has no clearly hypothesized synapomorphic characters. For example, many characters used to diagnose
<taxonomicName id="72AF98EBE444680314DE41A6461249AA" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
in
<bibRefCitation id="C18A7B4565DB0F6338F2F24167247777" author="Endrodi, S" journalOrPublisher="Junk Publisher, Dordrecht" pageId="71" pageNumber="72" title="The Dynastinae of the World. Dr. W." year="1985 a">
<normalizedToken id="0704C269F71499350BBC8A379905EA38" originalValue="Endrődis">Endrodi's</normalizedToken>
(1985a)
</bibRefCitation>
<taxonomicName id="D36F10B6C62A43592B8883CB8F2DEECE" lsidName="" pageId="45" pageNumber="46" rank="subfamily" subfamily="Dynastinae">Dynastinae</taxonomicName>
of the World are all variably present in
<taxonomicName id="2117A924C8B4190867304E46EB4DE682" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Cyclocephala</taxonomicName>
,
<taxonomicName id="98B3877CFF15406F48319256B64207F9" class="Insecta" family="Scarabaeidae" genus="Arriguttia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arriguttia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Arriguttia</taxonomicName>
, and
<taxonomicName id="1CBF20FDF3FF8A36A2789A32DF4D4D2C" class="Insecta" family="Scarabaeidae" genus="Aspidolea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspidolea" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Aspidolea</taxonomicName>
species: 1) body short, convex; 2) dorsal coloration yellow,
<pageBreakToken id="1F385CAF35268617D21360AD6AD676BF" pageId="46" pageNumber="47" start="start">with</pageBreakToken>
dark maculae, and with or without metallic reflections; 3) mandibles of males with small anterolateral tooth, lacking in females; 4) frontoclypeal suture complete; 5) 10-segmented antennae with a short club in both sexes; 6) large eyes; 7) males with thickened protarsi; and 8) protibia tridentate in both sexes.
</paragraph>
<paragraph id="B19FFA69E888716594D2C8F758FBE8BE" pageId="46" pageNumber="47">
The following combination of characters can be used to recognize
<taxonomicName id="E17F9B146F3F13AF55561D2C4CB22516" class="Insecta" family="Scarabaeidae" genus="Augoderia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Augoderia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Augoderia</taxonomicName>
species: 1) dorsal coloration yellowish or light brown, with or without elytral maculae, with or without metallic, mother-of-pearl sheen; 2) body not anteroposteriorly compressed or dorsoventrally flattened; 3) clypeal apex evenly rounded in dorsal view; 4) frons mesad of eyes with long, erect setae; 5) frontoclypeal suture complete; 6) males with anterolateral margin of mandibles weakly toothed; 7) mandibular molar area with rows of circular micropunctures; 8) apical margin of mentum weakly emarginated; 9) galea of the maxilla on inner surface with 3 fused basal teeth, a free median tooth, and 2 fused apical teeth (3-1-2 arrangement); 10) pronotum at base with incomplete or complete marginal bead; 11) pronotum on anterolateral portions with long, erect setae; 12) males and females with 3 protibial teeth, basal tooth reduced, removed from the apical 2 teeth, and oriented anteriorly; 13) protibial spur straight to weakly deflexed; 14) males with inner protarsal claw enlarged and narrowly cleft at apex; 15) mesocoxae widely separated; 16) metatibiae without distal, transverse carinae; 17) metacoxae with lateral edge perpendicular to ventral surface; 18) anterior edge of hindwing distal to apical hinge lacking setae and with produced, membranous border; 19) vein RA with 2 rows of pegs extending distally nearly to margin of apical hinge.
</paragraph>
</subSubSection>
</treatment>
</document>