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<document id="B4D0433F207B48F630049BB5BB11E0C3" ID-DOI="10.5281/zenodo.184042" ID-GBIF-Dataset="a926e1cf-ec51-4e1d-a2a1-f8a0562033f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="184042" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459939599139" checkinUser="plazi" docAuthor="Rodrigues, Miguel Trefaut &amp; Santos, Ednilza Maranhão Dos" docDate="2008" docId="664A2532FFB8C70037D75503FCDF0991" docLanguage="en" docName="zt01873p060.pdf" docOrigin="Zootaxa 1873" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Scriptosaura catimbau Rodrigues &amp; Santos, 2008, sp. nov." docType="treatment" docVersion="7" lastPageNumber="57" masterDocId="9A735D4AFFBAC7083740535CFF950863" masterDocTitle="A new genus and species of eyelid-less and limb reduced gymnophthalmid lizard from northeastern Brazil (Squamata, Gymnophthalmidae)" masterLastPageNumber="60" masterPageNumber="50" pageNumber="52" updateTime="1698230350253" updateUser="plazi">
<mods:mods id="091531D3C942D7631470105604E08AC1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8634362350765832B9D6FB925E32F128">
<mods:title id="6688C400ED81EAD0E77C796ECA810A6B">A new genus and species of eyelid-less and limb reduced gymnophthalmid lizard from northeastern Brazil (Squamata, Gymnophthalmidae)</mods:title>
</mods:titleInfo>
<mods:name id="D39676FE5C8E1EB6FAFF182054088EF3" type="personal">
<mods:role id="9349750DFD9EDBEB4A2B866DF2F6B4C6">
<mods:roleTerm id="9C1B19B487F4B385F495922820E2710F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2B39500BF38A297617BEEC16EAD44509">Rodrigues, Miguel Trefaut</mods:namePart>
</mods:name>
<mods:name id="80FEF1BC0573936135ADDA9D936F0A0D" type="personal">
<mods:role id="D95D39B0DF5A3A43E34500F0BE0026C5">
<mods:roleTerm id="8F1FD91B0D82872680C6A4B4AA7715CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="99F58619D1F4252AD439683F41A82C39">Santos, Ednilza Maranhão Dos</mods:namePart>
</mods:name>
<mods:typeOfResource id="C0E1EB1ED6C52D9BDAC543F990401CC7">text</mods:typeOfResource>
<mods:relatedItem id="EB60B6C704A4A4E60B1CA57CD65DCACB" type="host">
<mods:titleInfo id="F55C2BE6F606C99A9086FF6E80AA9DF7">
<mods:title id="4ADDF9E262B6F8D0BB37C152376EAEAD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="59E755A5EA9E35A2FFFBA2A6AF00D071">
<mods:date id="9A1943C6D9F7B389666269220C72B6A0">2008</mods:date>
<mods:detail id="82EBC4ADDC561DD031EAD5EC807F2D97" type="volume">
<mods:number id="4A981BCC172DB878E7EC32063204A8DA">1873</mods:number>
</mods:detail>
<mods:extent id="B11DBB3ECFBAE13091340667B08A34F3" unit="page">
<mods:start id="6BE6755D9387E93E3B8EE85DEEB63214">50</mods:start>
<mods:end id="FFF6761BD5E2BB854CB8E74A33991AB5">60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F9AE2E4417FF43FF2D7514D4893AFFB8">journal article</mods:classification>
<mods:identifier id="7A91CADBFCAA185E30C3D502EDD951D3" type="DOI">10.5281/zenodo.184042</mods:identifier>
<mods:identifier id="06920B491BAD7398605F1FFEF4B90639" type="GBIF-Dataset">a926e1cf-ec51-4e1d-a2a1-f8a0562033f2</mods:identifier>
<mods:identifier id="53D34909046230A5051EA367927E22D4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A0E54C0BA20F12B356CFBD6688624486" type="Zenodo-Dep">184042</mods:identifier>
</mods:mods>
<treatment id="664A2532FFB8C70037D75503FCDF0991" ID-DOI="http://doi.org/10.5281/zenodo.6232847" ID-GBIF-Taxon="119375788" ID-Zenodo-Dep="6232847" LSID="urn:lsid:plazi:treatment:664A2532FFB8C70037D75503FCDF0991" httpUri="http://treatment.plazi.org/id/664A2532FFB8C70037D75503FCDF0991" lastPageId="8" lastPageNumber="57" pageId="2" pageNumber="52">
<subSubSection id="A6F9C7AFFFB8C70A37D75503FEA00EFF" pageId="2" pageNumber="52" type="nomenclature">
<paragraph id="EE5C9424FFB8C70A37D75503FD800E1A" blockId="2.[151,533,1631,1692]" box="[151,533,1631,1657]" pageId="2" pageNumber="52">
<heading id="B5142348FFB8C70A37D75503FD800E1A" bold="true" box="[151,533,1631,1657]" fontSize="11" level="1" pageId="2" pageNumber="52" reason="1">
<emphasis id="DC974836FFB8C70A37D75503FD800E1A" bold="true" box="[151,533,1631,1657]" pageId="2" pageNumber="52">
<taxonomicName id="29E3EFA7FFB8C70A37D75503FE3C0E1A" box="[151,425,1631,1657]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="52" phylum="Chordata" rank="species" species="catimbau" status="sp. nov.">
<emphasis id="DC974836FFB8C70A37D75503FE3C0E1A" bold="true" box="[151,425,1631,1657]" italics="true" pageId="2" pageNumber="52">Scriptosaura catimbau</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C7A4F54DFFB8C70A36F85503FD800E1A" box="[440,533,1631,1657]" pageId="2" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="EE5C9424FFB8C70A37D755DEFEA00EFF" blockId="2.[151,533,1631,1692]" box="[151,309,1666,1692]" pageId="2" pageNumber="52">
(
<figureCitation id="76D888A1FFB8C70A37DF55DEFF640EFF" box="[159,241,1666,1692]" captionStart="FIGURE 1" captionStartId="3.[151,255,1652,1676]" captionTargetBox="[265,1318,211,1617]" captionTargetId="figure@3.[265,1329,198,1621]" captionTargetPageId="3" captionText="FIGURE 1. Lateral, ventral, and dorsal views of the head of the holotype of Scriptosaura catimbau (MZUSP 98059)." httpUri="https://zenodo.org/record/184043/files/figure.png" pageId="2" pageNumber="52">Figs. 1</figureCitation>
,
<figureCitation id="76D888A1FFB8C70A37BF55DEFE850EFF" box="[255,272,1666,1692]" captionStart="FIGURE 2" captionStartId="4.[151,255,744,768]" captionTargetBox="[188,1411,205,710]" captionTargetId="figure@4.[168,1419,194,720]" captionTargetPageId="4" captionText="FIGURE 2. Ventral view of the hindlimbs and preanal region, and a schematic view of the midbody scalation of the holotype of Scriptosaura catimbau (MZUSP 98059)." httpUri="https://zenodo.org/record/184044/files/figure.png" pageId="2" pageNumber="52">2</figureCitation>
,
<figureCitation id="76D888A1FFB8C70A365C55DEFEBE0EFF" box="[284,299,1666,1692]" captionStart="FIGURE 3" captionStartId="4.[151,255,1735,1759]" captionTargetBox="[151,1435,856,1710]" captionTargetId="figure@4.[151,1436,856,1711]" captionTargetPageId="4" captionText="FIGURE 3. Living specimen of Scriptosaura catimbau from Parque Nacional do Catimbau." httpUri="https://zenodo.org/record/184045/files/figure.png" pageId="2" pageNumber="52">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A6F9C7AFFFB8C70A37D75589FC130FDE" pageId="2" pageNumber="52" type="materials_examined">
<paragraph id="EE5C9424FFB8C70A37D75589FBF10F21" blockId="2.[151,1436,1746,2021]" pageId="2" pageNumber="52">
<typeStatus id="31582A86FFB8C70A37D75589FE9C0E8C" box="[151,265,1749,1775]" pageId="2" pageNumber="52" type="holotype">
<emphasis id="DC974836FFB8C70A37D75589FE9C0E8C" bold="true" box="[151,265,1749,1775]" pageId="2" pageNumber="52">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="88F20CE1FFB8C70A36585589FEEF0E8C" box="[280,378,1749,1775]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="52">MZUSP</collectionCode>
98059, an adult male from Fazenda Porto Seguro (
<geoCoordinate id="8BD7F2E3FFB8C70A34925589FBF50E8C" box="[978,1120,1746,1775]" direction="south" orientation="latitude" pageId="2" pageNumber="52" precision="15" value="-8.486945">08o29’13’’S</geoCoordinate>
,
<geoCoordinate id="8BD7F2E3FFB8C70A332E5589FA920E8C" box="[1134,1287,1746,1775]" direction="west" orientation="longitude" pageId="2" pageNumber="52" precision="15" value="-37.281113">37o16’52’’W</geoCoordinate>
), municipality of Buíque, Parque Nacional do
<taxonomicName id="29E3EFA7FFB8C70A357D545CFD210F79" box="[573,692,1792,1818]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="58" phylum="Chordata" rank="species" species="catimbau">Catimbau</taxonomicName>
: State of Pernambuco:
<collectingCountry id="96F4D4B4FFB8C70A348B545CFB830F79" box="[971,1046,1792,1818]" name="Brazil" pageId="2" pageNumber="52">Brazil</collectingCountry>
; collected on
<date id="9A5DB2E4FFB8C70A33FF545CFAE00F79" box="[1215,1397,1789,1818]" pageId="2" pageNumber="52" value="2008-03-07">7th March 2008</date>
by Miguel T. Rodrigues and Ednilza Maranhão dos Santos, field number MTR 15401.
</paragraph>
<paragraph id="EE5C9424FFB8C70A3785540CFC130FDE" blockId="2.[151,1436,1746,2021]" pageId="2" pageNumber="52">
<typeStatus id="31582A86FFB8C70A3785540CFED70F09" box="[197,322,1872,1898]" pageId="2" pageNumber="52" type="paratype">
<emphasis id="DC974836FFB8C70A3785540CFED70F09" bold="true" box="[197,322,1872,1898]" pageId="2" pageNumber="52">Paratypes</emphasis>
</typeStatus>
: Males:
<collectionCode id="88F20CE1FFB8C70A36F1540CFD860F09" box="[433,531,1872,1898]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="52">MZUSP</collectionCode>
98060, 98061, 98063, 98065, 98068–71, 98075, 98076; females:
<collectionCode id="88F20CE1FFB8C70A327A540CFA090F09" box="[1338,1436,1872,1898]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="52">MZUSP</collectionCode>
98062, 98064, 98066, 98067, 98072–4; collected between
<date id="9A5DB2E4FFB8C70A34225427FBAD0FF6" box="[866,1080,1912,1941]" pageId="2" pageNumber="52" value="2008-03-07" valueMax="2008-03-08" valueMin="2008-03-07">7–8th March 2008</date>
by Miguel T. Rodrigues and Ednilza Maranhão dos Santos; all other data as for the
<typeStatus id="31582A86FFB8C70A345854FFFC170FDE" box="[792,898,1955,1981]" pageId="2" pageNumber="52" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="A6F9C7AFFFB8C70A37855497FAD60F86" box="[197,1347,1995,2021]" pageId="2" pageNumber="52" type="etymology">
<paragraph id="EE5C9424FFB8C70A37855497FAD60F86" blockId="2.[151,1436,1746,2021]" box="[197,1347,1995,2021]" pageId="2" pageNumber="52">
<emphasis id="DC974836FFB8C70A37855497FED90F86" bold="true" box="[197,332,1995,2021]" pageId="2" pageNumber="52">Etymology</emphasis>
: A noun in apposition, in reference to the
<typeStatus id="31582A86FFB8C70A347D5497FCE50F86" box="[829,880,1995,2021]" pageId="2" pageNumber="52">type</typeStatus>
locality, Parque Nacional do
<taxonomicName id="29E3EFA7FFB8C70A338A5497FAD50F86" box="[1226,1344,1995,2021]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="58" phylum="Chordata" rank="species" species="catimbau">Catimbau</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="BA9CC4ACFFB9C70B37D75528FA140EE8" box="[151,1409,1652,1676]" httpUri="https://zenodo.org/record/184043/files/figure.png" pageId="3" pageNumber="53" targetBox="[265,1318,211,1617]" targetPageId="3">
<paragraph id="EE5C9424FFB9C70B37D75528FA140EE8" blockId="3.[151,1409,1652,1676]" box="[151,1409,1652,1676]" pageId="3" pageNumber="53">
<emphasis id="DC974836FFB9C70B37D75528FE8C0EEF" bold="true" box="[151,281,1652,1676]" pageId="3" pageNumber="53">FIGURE 1.</emphasis>
Lateral, ventral, and dorsal views of the head of the holotype of
<taxonomicName id="29E3EFA7FFB9C70B348C5528FB2A0EE8" box="[972,1215,1652,1675]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="53" phylum="Chordata" rank="species" species="catimbau">
<emphasis id="DC974836FFB9C70B348C5528FB2A0EE8" box="[972,1215,1652,1675]" italics="true" pageId="3" pageNumber="53">Scriptosaura catimbau</emphasis>
</taxonomicName>
(MZUSP 98059).
</paragraph>
</caption>
<subSubSection id="A6F9C7AFFFB9C70B378555E5FA120F8F" pageId="3" pageNumber="53" type="diagnosis">
<paragraph id="EE5C9424FFB9C70B378555E5FA120F8F" blockId="3.[151,1437,1721,2028]" pageId="3" pageNumber="53">
<emphasis id="DC974836FFB9C70B378555E5FED20EB0" bold="true" box="[197,327,1721,1747]" pageId="3" pageNumber="53">Diagnosis:</emphasis>
A small elongate gymnophthalmid (maximum SVL
<quantity id="291B39C1FFB9C70B34F555E6FB9D0EB7" box="[949,1032,1722,1748]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="53" unit="mm" value="53.0">53 mm</quantity>
) with a prominent snout and body slightly longer than tail. Forelimbs absent, hindlimbs rudimentary, styliform, ending in a small apical scale. Ear opening and eyelid absent. Frontonasal single, contacting parietal; prefrontals, frontal, frontoparietals, supraoculars, and loreal absent. Parietals longer than wide. Nostril in the center of nasal. Two supraciliaries, the anterior one larger, diagonally disposed, contacting subocular. Collar fold absent. One pair of chin shields. Dorsal scales cycloid, smooth anteriorly, posteriorly hexagonal, keeled, in 39–41 regular transverse series; lateral scales enlarged, smooth. Ventral scales smooth, in 32–34 transverse rows and four regular longitudinal series, the external ones wider; 15–19 scales around body. Males with four preanal pores, absent in females.
</paragraph>
</subSubSection>
<caption id="BA9CC4ACFFBEC70C37D751B4FD580B41" httpUri="https://zenodo.org/record/184044/files/figure.png" pageId="4" pageNumber="54" targetBox="[188,1411,205,710]" targetPageId="4">
<paragraph id="EE5C9424FFBEC70C37D751B4FD580B41" blockId="4.[151,1436,744,802]" pageId="4" pageNumber="54">
<emphasis id="DC974836FFBEC70C37D751B4FE8E0B63" bold="true" box="[151,283,744,768]" pageId="4" pageNumber="54">FIGURE 2.</emphasis>
Ventral view of the hindlimbs and preanal region, and a schematic view of the midbody scalation of the holotype of
<taxonomicName id="29E3EFA7FFBEC70C36585057FD9E0B41" box="[280,523,779,802]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="54" phylum="Chordata" rank="species" species="catimbau">
<emphasis id="DC974836FFBEC70C36585057FD9E0B41" box="[280,523,779,802]" italics="true" pageId="4" pageNumber="54">Scriptosaura catimbau</emphasis>
</taxonomicName>
(MZUSP 98059).
</paragraph>
</caption>
<caption id="BA9CC4ACFFBEC70C37D7559BFBE70EBC" box="[151,1138,1735,1759]" httpUri="https://zenodo.org/record/184045/files/figure.png" pageId="4" pageNumber="54" targetBox="[151,1435,856,1710]" targetPageId="4">
<paragraph id="EE5C9424FFBEC70C37D7559BFBE70EBC" blockId="4.[151,1138,1735,1759]" box="[151,1138,1735,1759]" pageId="4" pageNumber="54">
<emphasis id="DC974836FFBEC70C37D7559BFE8C0EBC" bold="true" box="[151,281,1735,1759]" pageId="4" pageNumber="54">FIGURE 3.</emphasis>
Living specimen of
<taxonomicName id="29E3EFA7FFBEC70C36B65594FD7C0EBC" box="[502,745,1736,1759]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="54" phylum="Chordata" rank="species" species="catimbau">
<emphasis id="DC974836FFBEC70C36B65594FD7C0EBC" box="[502,745,1736,1759]" italics="true" pageId="4" pageNumber="54">Scriptosaura catimbau</emphasis>
</taxonomicName>
from Parque Nacional do
<taxonomicName id="29E3EFA7FFBEC70C33435594FBFB0EBC" box="[1027,1134,1736,1759]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="58" phylum="Chordata" rank="species" species="catimbau">Catimbau</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="A6F9C7AFFFBEC70E3785544EFEC60BCF" lastPageId="6" lastPageNumber="56" pageId="4" pageNumber="54" type="description">
<paragraph id="EE5C9424FFBEC70D3785544EFDB10BC9" blockId="4.[151,1437,1810,2036]" lastBlockId="5.[151,1437,152,2018]" lastPageId="5" lastPageNumber="55" pageId="4" pageNumber="54">
<emphasis id="DC974836FFBEC70C3785544EFD8F0F4F" bold="true" box="[197,538,1810,1836]" pageId="4" pageNumber="54">
Description of the
<typeStatus id="31582A86FFBEC70C36EF544EFD8F0F4F" box="[431,538,1810,1836]" pageId="4" pageNumber="54" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="76D888A1FFBEC70C356B544EFDE80F4F" box="[555,637,1810,1836]" captionStart="FIGURE 1" captionStartId="3.[151,255,1652,1676]" captionTargetBox="[265,1318,211,1617]" captionTargetId="figure@3.[265,1329,198,1621]" captionTargetPageId="3" captionText="FIGURE 1. Lateral, ventral, and dorsal views of the head of the holotype of Scriptosaura catimbau (MZUSP 98059)." httpUri="https://zenodo.org/record/184043/files/figure.png" pageId="4" pageNumber="54">Figs. 1</figureCitation>
,
<figureCitation id="76D888A1FFBEC70C35CD544EFD0A0F4F" box="[653,671,1810,1836]" captionStart="FIGURE 2" captionStartId="4.[151,255,744,768]" captionTargetBox="[188,1411,205,710]" captionTargetId="figure@4.[168,1419,194,720]" captionTargetPageId="4" captionText="FIGURE 2. Ventral view of the hindlimbs and preanal region, and a schematic view of the midbody scalation of the holotype of Scriptosaura catimbau (MZUSP 98059)." httpUri="https://zenodo.org/record/184044/files/figure.png" pageId="4" pageNumber="54">2</figureCitation>
): Body long, snout prominent, without pronounced constrictions in neck or base of tail; no evidence of collar fold. Rostral broad, broadly projecting over symphysial; slightly wider than high, contacting first supralabial, nasal and frontonasal. In lateral view, the rostral forms an acute angle, with about 3/4 of its total length visible from above, the other quarter extends straight ventrally, in the same plane with the symphysial. Frontonasal hexagonal, longer than wide, reaching the level of posterior margin of the eye, wider posteriorly, in contact with rostral, nasal, first and second supraciliaries, parietal, and interparietal. Sutures with rostral and interparietal, broad and straight, that of first supraciliary concave, and those of second superciliary and parietal smaller. Prefrontals, frontals, supraoculars, and frontoparietals absent. Interparietal about twice as long as wide, as long as frontonasal and slightly longer than parietals; wider and straight anteriorly, posteriorly rounded; its lateral margins straight, converging posteriorly; separated from second supraciliary by parietal-frontonasal contact. Parietals irregularly hexagonal, longer than wide, contacting frontonasal, second supraciliary, a small postocular granule, an enlarged postocular, an enlarged elongate temporal, interparietal, and first dorsal row. A diagonally directed, postero-anterior oriented and polygonal supraciliary is longer than wide; anteriorly wider and contacting nasal, second supralabial, subocular, frontonasal and second supraciliary; longest suture with frontonasal. Second supraciliary small, contacting first supraciliary, frontonasal, parietal, and a small postocular. Subocular mostly above third supralabial, longer than wide, smaller than first supraciliary, contacting first supraciliary anteriorly and an enlarged postocular posteriorly which separates parietal and fourth supralabial. Eye small, as large as second supralabial, inserted between subocular, supraciliaries and three small postocular granules. Eyelids absent; eyeball round. Nasal pentagonal, above first and second supralabials, longer than wide, wider anteriorly, contacting additionally rostral, frontonasal and first supraciliary; nostril in center of nasal. Loreal and frenocular absent, probably fused with first supraciliary and corresponding to its wider portion. Five supralabials in the following order of increasing size of their base: first, fourth, second, third, and fifth; third below eye, fourth the highest. A longer than wide diagonally disposed temporal above fourth and fifth supralabials followed posteriorly by a series of dorso-ventrally oriented and imbricate higher than wide scales which cover tympanum; external ear opening absent.
</paragraph>
<paragraph id="EE5C9424FFBFC70D378650E4FBB70D01" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="55">Symphysial broad, with convex anterior border and straight posterior margin, slightly wider than long, contacting first infralabial and postmental, and situated clearly posterior to ventral edge of rostral. Postmental single, roughly pentagonal, longer than wide, contacting first and second infralabials. A single pair of genials in broad contact at midline and contacting second and third infralabials. Four infralabials, first the smallest, second and third the largest and with the same approximate size. All head scales smooth and juxtaposed, with irregularly distributed small sensorial organs. Two longitudinal series of transversally enlarged gular scales, smooth, imbricate, disposed in seven rows and extending from genials to the interbrachial region. Gular scales separated from lateral neck scales by a series of much smaller and longitudinally elongated and diagonally disposed scales. Interbrachial region with four longer than wide transversally disposed scales, the central ones larger. Ventrals in four longitudinal rows, the external one larger, the others longer than wide, imbricate, smooth, quadrangular; 33 transverse rows between gulars and preanal region.</paragraph>
<paragraph id="EE5C9424FFBFC70D378D562CFD1A0EC1" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="55">Anterior dorsal scales cycloid, smooth and imbricate, wider than long or irregular in size, becoming longer than wide and rounded posteriorly at the level of fifth row and then progressively changing to hexagonal, slightly keeled and mucronate until the level of leg insertion; 40 regularly transverse rows between interparietal and posterior level of leg insertion; 19 scales around midbody. Lateral neck scales smooth, cycloid, imbricate; the uppermost rows being larger and higher than long; other lateral neck scales smaller, irregular in size and shape and disposed in two irregularly longitudinal rows. Flanks with a ventrolateral row of smooth, imbricate and elongate scales with approximately the same size as dorsals. Above it, two irregularly longitudinal rows of subrectangular enlarged scales.</paragraph>
<paragraph id="EE5C9424FFBFC70D378555ECFF710F09" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="55">
Preanal region with four scales (
<figureCitation id="76D888A1FFBFC70D350755ECFD050EA9" box="[583,656,1712,1738]" captionStart="FIGURE 2" captionStartId="4.[151,255,744,768]" captionTargetBox="[188,1411,205,710]" captionTargetId="figure@4.[168,1419,194,720]" captionTargetPageId="4" captionText="FIGURE 2. Ventral view of the hindlimbs and preanal region, and a schematic view of the midbody scalation of the holotype of Scriptosaura catimbau (MZUSP 98059)." httpUri="https://zenodo.org/record/184044/files/figure.png" pageId="5" pageNumber="55">Fig. 2</figureCitation>
), the two lateral ones larger than and separated on the midline by centrals, which are longitudinally in contact. Four preanal pores. Dorsal part of tail with slightly smaller scales than posterior dorsals but keeled and strongly imbricate. Other parts of tail identical to their corresponding parts of the body. Differences between dorsal, lateral, and ventral scales of tail disappear gradually towards tip of tail.
</paragraph>
<paragraph id="EE5C9424FFBFC70D37865424FE450FD9" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="55">Forelimbs absent. Hindlimbs reduced to a needle-shape vestigial leg bud covered by five scales from the base to the tip; nail absent.</paragraph>
<paragraph id="EE5C9424FFBFC70E37865494FBE40919" blockId="5.[151,1437,152,2018]" lastBlockId="6.[151,1437,152,940]" lastPageId="6" lastPageNumber="56" pageId="5" pageNumber="55">Dorsal surface of body and tail light brown with a regular series of transverse darker dots corresponding to the posterior part of dorsal scales. A wide lateral dark brown stripe, formed by a highly irregular reticulate pattern and decreasing ventrally in conspicuity, extends along head, body, and tail from nasal to tip of tail. Ventral parts of head and gular region cream with an irregular dark brown reticulum. Ventral parts of body cream with a transverse series of dark brown dots, larger than dorsal ones. Tail color identical to body except ventrally where occasionally there are some dark spots, all the rest immaculate. Limb buds light brown dorsally with a reticulate pattern identical to that of lateral stripe; ventrally immaculate.</paragraph>
<paragraph id="EE5C9424FFBCC70E378552D4FE4709A9" blockId="6.[151,1437,152,940]" pageId="6" pageNumber="56">
Measurements of the
<typeStatus id="31582A86FFBCC70E368752D4FDA709C1" box="[455,562,392,418]" pageId="6" pageNumber="56" type="holotype">holotype</typeStatus>
: snout-vent length
<quantity id="291B39C1FFBCC70E345452D4FCFE09C1" box="[788,875,392,418]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="56" unit="mm" value="45.0">45 mm</quantity>
; tail
<quantity id="291B39C1FFBCC70E34E752D4FC6E09C1" box="[935,1019,392,418]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="56" unit="mm" value="19.0">19 mm</quantity>
(broken); diameter at midbody
<quantity id="291B39C1FFBCC70E323752D4FF5D09A9" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="56" unit="mm" value="4.2">4,2 mm</quantity>
; right leg bud
<quantity id="291B39C1FFBCC70E363152ECFE5A09A9" box="[369,463,432,458]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="56" unit="mm" value="2.2">2,2 mm</quantity>
.
</paragraph>
<paragraph id="EE5C9424FFBCC70E3785528BFEC60BCF" blockId="6.[151,1437,152,940]" pageId="6" pageNumber="56">
<emphasis id="DC974836FFBCC70E3785528BFEAD0992" bold="true" box="[197,312,471,497]" pageId="6" pageNumber="56">Variation</emphasis>
: Males are slightly smaller and have slightly longer tails than females: maximum SVL for males and females were
<quantity id="291B39C1FFBCC70E36F8515CFD9F0A79" box="[440,522,512,538]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="6" pageNumber="56" unit="mm" value="48.0">48 mm</quantity>
and
<quantity id="291B39C1FFBCC70E3503515CFD0C0A79" box="[579,665,512,538]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="56" unit="mm" value="53.0">53 mm</quantity>
, respectively. SVL varied respectively in males and females from 1.04 to 1.15 and 1.06 to 1.36 times tail length (r
<specimenCount id="F8E55FADFFBCC70E358E5174FCB70A27" box="[718,802,552,580]" pageId="6" pageNumber="56" type="male">2 males</specimenCount>
0.99, n = 4; r
<specimenCount id="F8E55FADFFBCC70E34855174FBA50A27" box="[965,1072,552,580]" pageId="6" pageNumber="56" type="female">2 females</specimenCount>
0.84, n = 4). No sexual differences were found in squamation. Variation in meristic characters (n = 19) was the following (mean and standard deviation, respectively, in parenthesis): dorsal rows, 39–41 (40.1
<emphasis id="DC974836FFBCC70E34A85127FC6D0AF7" box="[1000,1016,635,660]" italics="true" pageId="6" pageNumber="56">±</emphasis>
0.65); ventrals 32–34 (33
<emphasis id="DC974836FFBCC70E327C5127FAD90AF7" box="[1340,1356,635,660]" italics="true" pageId="6" pageNumber="56">±</emphasis>
0.66); scales around midbody 15–19 (17.7
<emphasis id="DC974836FFBCC70E350251FFFDC70ADF" box="[578,594,675,700]" italics="true" pageId="6" pageNumber="56">±</emphasis>
1.04). Like the
<typeStatus id="31582A86FFBCC70E344E51FEFCE20ADF" box="[782,887,674,700]" pageId="6" pageNumber="56" type="holotype">holotype</typeStatus>
, all specimens have two supraciliaries and one subocular, except for MZUSP 98074, with two suboculars. Likewise all the specimens have five supralabials, four infralabials, and seven gular rows, except for MZUSP 98075 which has eight. Four preanal pores are present in all males examined; they are absent in females. The styliform hindlimb is covered by five scales in all specimens. The color pattern of males and females is very similar except in adult males where ventral surfaces are characteristically reticulated with dark brown whereas in adult females and juveniles ventral surfaces are immaculate.
</paragraph>
</subSubSection>
<caption id="BA9CC4ACFFBCC70E37D754CDFE690FA8" httpUri="https://zenodo.org/record/184046/files/figure.png" pageId="6" pageNumber="56" targetBox="[249,1336,1003,1907]" targetPageId="6">
<paragraph id="EE5C9424FFBCC70E37D754CDFE690FA8" blockId="6.[151,1436,1937,1995]" pageId="6" pageNumber="56">
<emphasis id="DC974836FFBCC70E37D754CDFE8C0FCA" bold="true" box="[151,281,1937,1961]" pageId="6" pageNumber="56">FIGURE 4.</emphasis>
Type locality of
<taxonomicName id="29E3EFA7FFBCC70E368D54CDFD540FCB" box="[461,705,1937,1960]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="56" phylum="Chordata" rank="species" species="catimbau">
<emphasis id="DC974836FFBCC70E368D54CDFD540FCB" box="[461,705,1937,1960]" italics="true" pageId="6" pageNumber="56">Scriptosaura catimbau</emphasis>
</taxonomicName>
in the context of the Caatingas of northeastern Brazil (gray) and that of Parque Nacional do
<taxonomicName id="29E3EFA7FFBCC70E36CC54E8FE620FA8" box="[396,503,1972,1995]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="catimbau">Catimbau</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="A6F9C7AFFFBDC700378653CBFCDF0991" lastPageId="8" lastPageNumber="58" pageId="7" pageNumber="57" type="distribution">
<paragraph id="EE5C9424FFBDC70F378653CBFAED0B6F" blockId="7.[151,1437,151,780]" pageId="7" pageNumber="57">
<emphasis id="DC974836FFBDC70F378653CBFDF708D2" bold="true" box="[198,610,151,177]" pageId="7" pageNumber="57">Distribution and natural history</emphasis>
: The Parque Nacional do
<taxonomicName id="29E3EFA7FFBDC70F34E453C4FB8308D1" box="[932,1046,152,178]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="catimbau">Catimbau</taxonomicName>
(
<figureCitation id="76D888A1FFBDC70F336953C4FBE608D1" box="[1065,1139,152,178]" captionStart="FIGURE 4" captionStartId="6.[151,255,1937,1961]" captionTargetBox="[249,1336,1003,1907]" captionTargetId="figure@6.[244,1344,997,1913]" captionTargetPageId="6" captionText="FIGURE 4. Type locality of Scriptosaura catimbau in the context of the Caatingas of northeastern Brazil (gray) and that of Parque Nacional do Catimbau." httpUri="https://zenodo.org/record/184046/files/figure.png" pageId="7" pageNumber="57">Fig. 4</figureCitation>
) was recently created to protect the habitats included in or surrounding the highly eroded plateau of Silurian-Devonian sedimentary deposits of Tacaratu formation (
<bibRefCitation id="8A72E9D5FFBDC70F355853B4FC030961" author="Projeto" box="[536,918,232,258]" pageId="7" pageNumber="59" refString="Projeto RADAMBRASIL (1983) Levantamento dos Recursos Naturais. Vol. 30, Folhas SC 24 / 25, Aracaju, Recife. Ministerio das Minas e Energia, Rio de Janeiro, 852 pp." type="book" year="1983">Projeto RADAMBRASIL 1983</bibRefCitation>
;
<bibRefCitation id="8A72E9D5FFBDC70F34E653B4FBA80961" author="CPRM" box="[934,1085,232,258]" pageId="7" pageNumber="59" refString="CPRM (2005) Projeto cadastro de fontes de abastecimento por agua subterranea, Pernambuco: diagnostico do municipio de Buique. Ministerio de Minas e Energia, CPRM. 10 pp." type="book" year="2005">CPRM 2005</bibRefCitation>
) in the Caatingas of State of Pernambuco. The area, with about 62,000 hectares, consists of extensive arenitic and conglomerate outcrops of the plateau and areas of accumulation of sandy soils resulting from its erosion (
<bibRefCitation id="8A72E9D5FFBDC70F332F5264FA1A0931" author="RADAMBRASIL" box="[1135,1423,312,338]" pageId="7" pageNumber="59" refString="Projeto RADAMBRASIL (1983) Levantamento dos Recursos Naturais. Vol. 30, Folhas SC 24 / 25, Aracaju, Recife. Ministerio das Minas e Energia, Rio de Janeiro, 852 pp." type="book" year="1983">RADAMBRASIL 1983</bibRefCitation>
). Sandstones are irregularly capped with conglomerates which gives some protection from erosion. The plateau, with elevations varying between 700 and about
<quantity id="291B39C1FFBDC70F358E52D4FCBE09C1" box="[718,811,392,418]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="57" unit="m" value="1000.0">1000 m</quantity>
, is highly dissected at the borders (
<figureCitation id="76D888A1FFBDC70F338D52D4FA8609C1" box="[1229,1299,392,418]" captionStart="FIGURE 5" captionStartId="7.[151,255,1766,1790]" captionTargetBox="[154,1433,840,1742]" captionTargetId="figure@7.[154,1434,840,1743]" captionTargetPageId="7" captionText="FIGURE 5. General aspect of the eroded border of the sandstone Silurian Devonian plateau at Parque Nacional do Catimbau, type locality of Scriptosaura catimbau (A); typical sandy habitat of Scriptosaura catimbau, note the abundance of the “ caroá ” (Neoglaziovia sp.), the thin bromeliad (B); track of Scriptosaura catimbau (C)." httpUri="https://zenodo.org/record/184047/files/figure.png" pageId="7" pageNumber="57">Fig. 5</figureCitation>
A) and predominantly covered by a xeromorphic Caatinga vegetation. Plant families dominant in the area are the
<taxonomicName id="29E3EFA7FFBDC70F37D75284FEDD0991" box="[151,328,472,498]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="57" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
,
<taxonomicName id="29E3EFA7FFBDC70F36195284FD8C0991" box="[345,537,472,498]" class="Magnoliopsida" family="Caesalpiniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="7" pageNumber="57" phylum="Magnoliophyta" rank="family">Caesalpiniaceae</taxonomicName>
,
<taxonomicName id="29E3EFA7FFBDC70F35695284FD3D0991" box="[553,680,472,498]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="57" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
, Mymosaceae,
<taxonomicName id="29E3EFA7FFBDC70F34285284FC400991" box="[872,981,472,498]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="7" pageNumber="57" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
and
<taxonomicName id="29E3EFA7FFBDC70F33585284FA0D0991" authority="Gomes et al. 2006" authorityName="Gomes et al." authorityYear="2006" box="[1048,1432,472,498]" class="Magnoliopsida" family="Cactaceae" kingdom="Plantae" order="Caryophyllales" pageId="7" pageNumber="57" phylum="Tracheophyta" rank="family">
Cactaceae (
<bibRefCitation id="8A72E9D5FFBDC70F33E45284FA050991" author="Gomes" box="[1188,1424,472,498]" pageId="7" pageNumber="59" refString="Gomes, A. P. S., Rodal, M. J. N. &amp; Melo, A. L. (2006) Floristica e fitogeografia da vegetacao arbustiva subcaducifolia da Chapada de Sao Jose, Buique, PE, Brasil. Acta Botanica Brasilica, 20, 37 - 48." type="journal article" year="2006">Gomes et al. 2006</bibRefCitation>
)
</taxonomicName>
. Although less abundant, the
<taxonomicName id="29E3EFA7FFBDC70F36A4515CFD100A79" box="[484,645,512,538]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="57" phylum="Tracheophyta" rank="family">Bromeliaceae</taxonomicName>
are also characteristic either in the outcrops or in the sandy soils; in the latter habitat the caroá (
<taxonomicName id="29E3EFA7FFBDC70F36A45174FDEA0A22" box="[484,639,552,577]" class="Liliopsida" family="Bromeliaceae" genus="Neoglaziovia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="57" phylum="Tracheophyta" rank="genus">
<emphasis id="DC974836FFBDC70F36A45174FDEA0A22" box="[484,639,552,577]" italics="true" pageId="7" pageNumber="57">Neoglaziovia</emphasis>
</taxonomicName>
sp.) is particularly frequent (
<figureCitation id="76D888A1FFBDC70F34A05174FBBD0A21" box="[992,1064,552,578]" captionStart="FIGURE 5" captionStartId="7.[151,255,1766,1790]" captionTargetBox="[154,1433,840,1742]" captionTargetId="figure@7.[154,1434,840,1743]" captionTargetPageId="7" captionText="FIGURE 5. General aspect of the eroded border of the sandstone Silurian Devonian plateau at Parque Nacional do Catimbau, type locality of Scriptosaura catimbau (A); typical sandy habitat of Scriptosaura catimbau, note the abundance of the “ caroá ” (Neoglaziovia sp.), the thin bromeliad (B); track of Scriptosaura catimbau (C)." httpUri="https://zenodo.org/record/184047/files/figure.png" pageId="7" pageNumber="57">Fig. 5</figureCitation>
B). On sandy areas vegetation cover consists basically of scattered thickets of variable size with a dry, and thin leaf litter generally concentrated at the basis of trees and bushes and separated by spaces of bare sand (
<figureCitation id="76D888A1FFBDC70F334C5124FBC70AF1" box="[1036,1106,632,658]" captionStart="FIGURE 5" captionStartId="7.[151,255,1766,1790]" captionTargetBox="[154,1433,840,1742]" captionTargetId="figure@7.[154,1434,840,1743]" captionTargetPageId="7" captionText="FIGURE 5. General aspect of the eroded border of the sandstone Silurian Devonian plateau at Parque Nacional do Catimbau, type locality of Scriptosaura catimbau (A); typical sandy habitat of Scriptosaura catimbau, note the abundance of the “ caroá ” (Neoglaziovia sp.), the thin bromeliad (B); track of Scriptosaura catimbau (C)." httpUri="https://zenodo.org/record/184047/files/figure.png" pageId="7" pageNumber="57">Fig. 5</figureCitation>
B). At some places near the lower slopes of the highly eroded plateau where water is more abundant vegetation is higher and evergreen, and a denser and humid leaf litter is present. Annual mean temperatures and precipitation in the area are 26o C and
<quantity id="291B39C1FFBDC70F378851AEFEB80B6F" box="[200,301,754,780]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="57" unit="mm" value="600.0">600 mm</quantity>
, respectively; most of the rain falls between April and June (
<bibRefCitation id="8A72E9D5FFBDC70F34B351AEFB100B6F" author="Nimer" box="[1011,1157,754,780]" pageId="7" pageNumber="59" refString="Nimer, E. (1979) Climatologia do Brasil. Superintendencia de Recursos Naturais e Meio Ambiente, SUPREN, Rio de Janeiro, 421 pp." type="book" year="1979">Nimer 1979</bibRefCitation>
;
<bibRefCitation id="8A72E9D5FFBDC70F33D151AEFAF90B6F" author="Gomes" box="[1169,1388,754,780]" pageId="7" pageNumber="59" refString="Gomes, A. P. S., Rodal, M. J. N. &amp; Melo, A. L. (2006) Floristica e fitogeografia da vegetacao arbustiva subcaducifolia da Chapada de Sao Jose, Buique, PE, Brasil. Acta Botanica Brasilica, 20, 37 - 48." type="journal article" year="2006">Gomes et al. 2006</bibRefCitation>
).
</paragraph>
<caption id="BA9CC4ACFFBDC70F37D755BAFB160F20" httpUri="https://zenodo.org/record/184047/files/figure.png" pageId="7" pageNumber="57" targetBox="[154,1433,840,1742]" targetPageId="7">
<paragraph id="EE5C9424FFBDC70F37D755BAFB160F20" blockId="7.[151,1436,1766,1859]" pageId="7" pageNumber="57">
<emphasis id="DC974836FFBDC70F37D755BAFE8F0E9D" bold="true" box="[151,282,1766,1790]" pageId="7" pageNumber="57">FIGURE 5.</emphasis>
General aspect of the eroded border of the sandstone Silurian Devonian plateau at Parque Nacional do
<taxonomicName id="29E3EFA7FFBDC70F323655BBFF710F43" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="catimbau">Catimbau</taxonomicName>
, type locality of
<taxonomicName id="29E3EFA7FFBDC70F36D95456FD180F42" box="[409,653,1802,1825]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="57" phylum="Chordata" rank="species" species="catimbau">
<emphasis id="DC974836FFBDC70F36D95456FD180F42" box="[409,653,1802,1825]" italics="true" pageId="7" pageNumber="57">Scriptosaura catimbau</emphasis>
</taxonomicName>
(A); typical sandy habitat of
<taxonomicName id="29E3EFA7FFBDC70F348B5456FB2A0F42" box="[971,1215,1802,1825]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="57" phylum="Chordata" rank="species" species="catimbau">
<emphasis id="DC974836FFBDC70F348B5456FB2A0F42" box="[971,1215,1802,1825]" italics="true" pageId="7" pageNumber="57">Scriptosaura catimbau</emphasis>
</taxonomicName>
, note the abundance of the “caroá”
<emphasis id="DC974836FFBDC70F367A5470FE5A0F20" box="[314,463,1836,1859]" italics="true" pageId="7" pageNumber="57">
(
<taxonomicName id="29E3EFA7FFBDC70F36015470FE5A0F20" box="[321,463,1836,1859]" class="Liliopsida" family="Bromeliaceae" genus="Neoglaziovia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="57" phylum="Tracheophyta" rank="genus">Neoglaziovia</taxonomicName>
</emphasis>
sp.), the thin bromeliad (B); track of
<taxonomicName id="29E3EFA7FFBDC70F341F5470FBC70F20" box="[863,1106,1836,1859]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="57" phylum="Chordata" rank="species" species="catimbau">
<emphasis id="DC974836FFBDC70F341F5470FBC70F20" box="[863,1106,1836,1859]" italics="true" pageId="7" pageNumber="57">Scriptosaura catimbau</emphasis>
</taxonomicName>
(C).
</paragraph>
</caption>
<paragraph id="EE5C9424FFBDC7003786542BFCDF0991" blockId="7.[151,1436,1911,2017]" lastBlockId="8.[151,1437,152,498]" lastPageId="8" lastPageNumber="58" pageId="7" pageNumber="57">
All specimens of
<taxonomicName id="29E3EFA7FFBDC70F36D9542BFD330FF3" box="[409,678,1911,1936]" class="Reptilia" family="Gymnophthalmidae" genus="Scriptosaura" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="57" phylum="Chordata" rank="species" species="catimbau">
<emphasis id="DC974836FFBDC70F36D9542BFD330FF3" box="[409,678,1911,1936]" italics="true" pageId="7" pageNumber="57">Scriptosaura catimbau</emphasis>
</taxonomicName>
were found on the slopes of the northern border of the plateau, while searching among the leaf litter around vegetation thickets in the sandy areas. They were always found buried in the sand a few centimeters below surface. When exposed they always tried to escape with fast sand swimming movements below sand. Although we obtained only
<specimenCount id="F8E55FADFFB2C70034C953C4FBBF08D1" box="[905,1066,152,178]" pageId="8" pageNumber="58" type="generic">19 specimens</specimenCount>
, several escaped; the characteristic tracks left in the sand (
<figureCitation id="76D888A1FFB2C70036A9539CFDA508B9" box="[489,560,192,218]" captionStart="FIGURE 5" captionStartId="7.[151,255,1766,1790]" captionTargetBox="[154,1433,840,1742]" captionTargetId="figure@7.[154,1434,840,1743]" captionTargetPageId="7" captionText="FIGURE 5. General aspect of the eroded border of the sandstone Silurian Devonian plateau at Parque Nacional do Catimbau, type locality of Scriptosaura catimbau (A); typical sandy habitat of Scriptosaura catimbau, note the abundance of the “ caroá ” (Neoglaziovia sp.), the thin bromeliad (B); track of Scriptosaura catimbau (C)." httpUri="https://zenodo.org/record/184047/files/figure.png" pageId="8" pageNumber="58">Fig. 5</figureCitation>
C) suggest that the species is very abundant in the area. The presence of recent tracks during the day in areas previously searched for specimens and where tracks had been absent suggest diurnality. The presence of several small juveniles in our sample indicates their recent eclosion. Other lizards obtained in the area were the tropidurids
<taxonomicName id="29E3EFA7FFB2C70035FA5264FC480932" box="[698,989,312,337]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="cocorobensis">
<emphasis id="DC974836FFB2C70035FA5264FC480932" box="[698,989,312,337]" italics="true" pageId="8" pageNumber="58">Tropidurus cocorobensis</emphasis>
</taxonomicName>
,
<taxonomicName id="29E3EFA7FFB2C70034AC5264FB4D0932" box="[1004,1240,312,337]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="hispidus">
<emphasis id="DC974836FFB2C70034AC5264FB4D0932" box="[1004,1240,312,337]" italics="true" pageId="8" pageNumber="58">Tropidurus hispidus</emphasis>
</taxonomicName>
, and
<taxonomicName id="29E3EFA7FFB2C700325B5264FEAD091A" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="semitaeniatus">
<emphasis id="DC974836FFB2C700325B5264FEAD091A" italics="true" pageId="8" pageNumber="58">Tropidurus semitaeniatus</emphasis>
</taxonomicName>
, the iguanid
<emphasis id="DC974836FFB2C700369B523CFD1C091A" box="[475,649,352,377]" italics="true" pageId="8" pageNumber="58">Iguana iguana</emphasis>
, the gymnophthalmids
<taxonomicName id="29E3EFA7FFB2C70034E8523CFB3D091A" box="[936,1192,352,377]" class="Reptilia" family="Gymnophthalmidae" genus="Acratosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="mentalis">
<emphasis id="DC974836FFB2C70034E8523CFB3D091A" box="[936,1192,352,377]" italics="true" pageId="8" pageNumber="58">Acratosaura mentalis</emphasis>
</taxonomicName>
,
<taxonomicName id="29E3EFA7FFB2C70033FA523CFF5809C2" class="Reptilia" family="Gymnophthalmidae" genus="Anotosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="vanzolinia">
<emphasis id="DC974836FFB2C70033FA523CFF5809C2" italics="true" pageId="8" pageNumber="58">Anotosaura vanzolinia</emphasis>
</taxonomicName>
,
<taxonomicName id="29E3EFA7FFB2C700379E52D4FDA409C2" box="[222,561,392,417]" class="Reptilia" family="Gymnophthalmidae" genus="Micrablepharus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="maximiliani">
<emphasis id="DC974836FFB2C700379E52D4FDA409C2" box="[222,561,392,417]" italics="true" pageId="8" pageNumber="58">Micrablepharus maximiliani</emphasis>
</taxonomicName>
, and
<taxonomicName id="29E3EFA7FFB2C700353752D4FC1B09C2" box="[631,910,392,417]" class="Reptilia" family="Gymnophthalmidae" genus="Vanzosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="rubricauda">
<emphasis id="DC974836FFB2C700353752D4FC1B09C2" box="[631,910,392,417]" italics="true" pageId="8" pageNumber="58">Vanzosaura rubricauda</emphasis>
</taxonomicName>
, the teiids
<emphasis id="DC974836FFB2C700335852D4FB5B09C2" box="[1048,1230,392,417]" italics="true" pageId="8" pageNumber="58">Ameiva ameiva</emphasis>
,
<taxonomicName id="29E3EFA7FFB2C70033A152D4FF6C09AA" class="Insecta" family="Curculionidae" genus="Cnemidophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="58" phylum="Arthropoda" rank="species" species="ocellifer">
<emphasis id="DC974836FFB2C70033A152D4FF6C09AA" italics="true" pageId="8" pageNumber="58">Cnemidophorus ocellifer</emphasis>
</taxonomicName>
and
<taxonomicName id="29E3EFA7FFB2C700367D52ECFDD709AA" box="[317,578,432,457]" class="Reptilia" family="Teiidae" genus="Tupinambis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="merianae">
<emphasis id="DC974836FFB2C700367D52ECFDD709AA" box="[317,578,432,457]" italics="true" pageId="8" pageNumber="58">Tupinambis merianae</emphasis>
</taxonomicName>
, the gekkonids
<emphasis id="DC974836FFB2C700344C52ECFAC809AA" box="[780,1373,432,457]" italics="true" pageId="8" pageNumber="58">
<taxonomicName id="29E3EFA7FFB2C700344C52ECFBD409AA" box="[780,1089,432,457]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="brasilianus">Hemidactylus brasilianus</taxonomicName>
,
<taxonomicName id="29E3EFA7FFB2C700331252ECFAC809AA" box="[1106,1373,432,457]" class="Reptilia" family="Phyllodactylidae" genus="Phyllopezus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="pollicaris">Phyllopezus pollicaris</taxonomicName>
</emphasis>
, and
<taxonomicName id="29E3EFA7FFB2C70037D75284FE1A0992" box="[151,399,472,497]" class="Reptilia" family="Phyllodactylidae" genus="Phyllopezus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="periosus">
<emphasis id="DC974836FFB2C70037D75284FE1A0992" box="[151,399,472,497]" italics="true" pageId="8" pageNumber="58">Phyllopezus periosus</emphasis>
</taxonomicName>
, and the scincid
<taxonomicName id="29E3EFA7FFB2C70035155284FCD60992" box="[597,835,472,497]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="58" phylum="Chordata" rank="species" species="agmosticha">
<emphasis id="DC974836FFB2C70035155284FCD60992" box="[597,835,472,497]" italics="true" pageId="8" pageNumber="58">Mabuya agmosticha</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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