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<document id="9075CCBD3B61180F6ED827DF90DDEDCB" ID-CLB-Dataset="264211" ID-DOI="10.5852/ejt.2023.889.2245" ID-GBIF-Dataset="db71145d-18b1-4ad1-a2a4-72d7b8928a7d" ID-ISSN="2118-9773" ID-Zenodo-Dep="8269451" ID-ZooBank="52E11625-6E26-4E4F-8B15-05195087BAB4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1692626252704" checkinUser="felipe" docAuthor="Domahovski, Alexandre C. &amp; Cavichioli, Rodney R." docDate="2023" docId="66347308574BFF98FD0C9721FEC1F833" docLanguage="en" docName="ejt-889_1-53.pdf" docOrigin="European Journal of Taxonomy 889 (1)" docSource="http://zoobank.org/52e11625-6e26-4e4f-8b15-05195087bab4" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Costanana alata Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="97D667AD-22B6-4354-B325-B4CBF2BCBEF2" docUuidSource="ZooBank" docVersion="8" lastPageNumber="7" masterDocId="9A0D0B70574FFF9EFF8A935BFF99FFB7" masterDocTitle="Eight new species of Costanana DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini): taxonomic changes, key to males, and description of Metacostana gen. nov." masterLastPageNumber="53" masterPageNumber="1" pageNumber="5" updateTime="1699595446126" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="FDBAFA770A042379C5983368312E0695" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F16E4B443C8D56F8215AD26BF13B5C53">
<mods:title id="1731D88F9C699D202ED3AD3D022619E6">Eight new species of Costanana DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini): taxonomic changes, key to males, and description of Metacostana gen. nov.</mods:title>
</mods:titleInfo>
<mods:name id="AD055757FC949E9BF9BB0427B1B379A9" type="personal">
<mods:role id="442A06EC7941A74CEA2E14D4C39B282B">
<mods:roleTerm id="3F7E908E0EB7E68F1672A1047B1CCD73">Author</mods:roleTerm>
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<mods:namePart id="3190B9FF2883CB2BAD5C4034B5059E08">Domahovski, Alexandre C.</mods:namePart>
<mods:nameIdentifier id="82EE36601A4FE6F3A817E33B86A0F450" type="LSID">0B988BD5-599B-42CE-ADCB-50FC813E104E</mods:nameIdentifier>
<mods:affiliation id="977332018BF59937641516173237BA0D">Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Caixa Postal 68044, Rio de Janeiro, 21941 - 971, RJ, Brazil.</mods:affiliation>
<mods:nameIdentifier id="FF8C6EE8C296BA7040601966E627D904" type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="8C85D37EBDECB6637DF5B8D8BE3DF3B0">Cavichioli, Rodney R.</mods:namePart>
<mods:nameIdentifier id="58E70D8DBAC1F24DBDC7763D68D13E0B" type="LSID">DFC42ABD-5602-4D6E-B691-49F716ECDFDC</mods:nameIdentifier>
<mods:affiliation id="688C705B680D777D9CA0CC8B54583A52">Departamento de Zoologia, Universidade Federal do Paraná. Caixa Postal 19020, Curitiba, 81531 - 980, PR, Brazil.</mods:affiliation>
<mods:nameIdentifier id="0AB093DE3F40558A934DA837B8AAC9AC" type="email">cavich@ufpr.br</mods:nameIdentifier>
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<mods:title id="C049B3031CDE92807215D4593ECC0671">European Journal of Taxonomy</mods:title>
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<mods:part id="AAE2B965ABD1B84FA79E744A0788A18A">
<mods:date id="7CC5D666E1EBE97C3C5A954EB9FD7693">2023</mods:date>
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<mods:number id="2D0B6E3C58A5007BE4619AD3A49B98A5">2023-08-21</mods:number>
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<mods:number id="50BD95931C42BB419902F649BB1664BF">889</mods:number>
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<treatment id="66347308574BFF98FD0C9721FEC1F833" ID-DOI="http://doi.org/10.5281/zenodo.8271519" ID-GBIF-Taxon="212449061" ID-Zenodo-Dep="8271519" LSID="urn:lsid:zoobank.org:act:97D667AD-22B6-4354-B325-B4CBF2BCBEF2" httpUri="http://treatment.plazi.org/id/66347308574BFF98FD0C9721FEC1F833" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<subSubSection id="A6879195574BFF9AFD0C9721FC37FB22" box="[646,942,1146,1173]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="EE22C21E574BFF9AFD0C9721FC37FB22" blockId="4.[360,1227,1146,1243]" box="[646,942,1146,1173]" pageId="4" pageNumber="5">
<heading id="B56A7572574BFF9AFD0C9721FC37FB22" box="[646,942,1146,1173]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="299DB99D574BFF9AFD0C9721FCD5FB22" ID-CoL="f20932d5-efc3-43cf-80cf-9ed3446f3c8b" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" box="[646,844,1146,1173]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alata" status="sp. nov.">
<emphasis id="DCE91E0C574BFF9AFD0C9721FCD5FB22" bold="true" box="[646,844,1146,1173]" italics="true" pageId="4" pageNumber="5">Costanana alata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7DAA377574BFF9AFCD99720FC37FB22" box="[851,942,1147,1173]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="A6879195574BFF9AFEE297C6FCF6FB6C" pageId="4" pageNumber="5" type="description">
<paragraph id="EE22C21E574BFF9AFEE297C6FB52FB0F" blockId="4.[360,1227,1146,1243]" box="[360,1227,1181,1208]" pageId="4" pageNumber="5">
<uri id="9A0CCE1C574BFF9AFEE297C6FB52FB0F" box="[360,1227,1181,1208]" pageId="4" pageNumber="5">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="EE22C21E574BFF9AFD4E979AFCF6FB6C" blockId="4.[360,1227,1146,1243]" box="[708,879,1217,1243]" pageId="4" pageNumber="5">
<figureCitation id="76A6DE9B574BFF9AFD4E979AFC94FB6C" box="[708,781,1217,1243]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
,
<figureCitation id="76A6DE9B574BFF9AFC91979AFCF6FB6C" box="[795,879,1217,1243]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[189,1398,742,1851]" captionTargetId="figure-195@26.[189,1398,742,1851]" captionTargetPageId="26" captionText="Fig. 11. Habitus of new species of Costanana DeLong &amp; Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. alata sp. nov., holotype, ♂ (DZUP 215484). C–D. C. bifida sp. nov., holotype, ♂ (DZUP 215493). E–F. C. cifi sp. nov., holotype, ♂ (DZUP 215484). G–H. C. luzi sp. nov., holotype ♂ (DZUP 215489). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269485" httpUri="https://zenodo.org/record/8269485/files/figure.png" pageId="4" pageNumber="5">11A–B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A6879195574BFF9AFF379659FAF3FA60" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="EE22C21E574BFF9AFF379659FEAFFAAB" blockId="4.[189,310,1282,1308]" box="[189,310,1282,1308]" pageId="4" pageNumber="5">
<heading id="B56A7572574BFF9AFF379659FEAFFAAB" bold="true" box="[189,310,1282,1308]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="DCE91E0C574BFF9AFF379659FEAFFAAB" bold="true" box="[189,310,1282,1308]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="EE22C21E574BFF9AFF37966AFAF3FA60" blockId="4.[188,1399,1329,1496]" pageId="4" pageNumber="5">
Coloration (
<figureCitation id="76A6DE9B574BFF9AFEC1966AFE2FFAFC" box="[331,438,1329,1355]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[189,1398,742,1851]" captionTargetId="figure-195@26.[189,1398,742,1851]" captionTargetPageId="26" captionText="Fig. 11. Habitus of new species of Costanana DeLong &amp; Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. alata sp. nov., holotype, ♂ (DZUP 215484). C–D. C. bifida sp. nov., holotype, ♂ (DZUP 215493). E–F. C. cifi sp. nov., holotype, ♂ (DZUP 215484). G–H. C. luzi sp. nov., holotype ♂ (DZUP 215489). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269485" httpUri="https://zenodo.org/record/8269485/files/figure.png" pageId="4" pageNumber="5">Fig. 11A</figureCitation>
) of head and pronotum brown with small black maculae, forewing costal margin with narrow light yellow macula on anterior third and small depigmented (white) macula on apical third. Style (
<figureCitation id="76A6DE9B574BFF9AFEC5962CFE33FA26" box="[335,426,1399,1425]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="4" pageNumber="5">Fig. 1K</figureCitation>
) with dentiform ventral process near base of blade; apex truncated, forming pair of acute processes directed dorsally and ventrally. Aedeagus (
<figureCitation id="76A6DE9B574BFF9AFCE696C1FC74FA03" box="[876,1005,1434,1460]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="4" pageNumber="5">Fig. 1L–M</figureCitation>
) with pair of ventral wing-shaped processes near base; shaft with slender basodorsal process and pair of minute spurs on ventral margin.
</paragraph>
</subSubSection>
<subSubSection id="A6879195574BFF9AFF3796A4FB12F9F0" box="[189,1163,1535,1608]" pageId="4" pageNumber="5" type="etymology">
<paragraph id="EE22C21E574BFF9AFF3796A4FEDDF9AE" blockId="4.[189,324,1535,1561]" box="[189,324,1535,1561]" pageId="4" pageNumber="5">
<heading id="B56A7572574BFF9AFF3796A4FEDDF9AE" bold="true" box="[189,324,1535,1561]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="DCE91E0C574BFF9AFF3796A4FEDDF9AE" bold="true" box="[189,324,1535,1561]" pageId="4" pageNumber="5">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="EE22C21E574BFF9AFF379576FB12F9F0" blockId="4.[189,1163,1581,1608]" box="[189,1163,1581,1608]" pageId="4" pageNumber="5">
The species epithet ‘
<taxonomicName id="299DB99D574BFF9AFE259576FE72F9F0" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" box="[431,491,1581,1607]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alata">
<emphasis id="DCE91E0C574BFF9AFE259576FE72F9F0" box="[431,491,1581,1607]" italics="true" pageId="4" pageNumber="5">alata</emphasis>
</taxonomicName>
’ refers to the aedeagus with processes resembling wings.
</paragraph>
</subSubSection>
<subSubSection id="A6879195574BFF9AFF379534FBC3F948" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="EE22C21E574BFF9AFF379534FE33F93E" blockId="4.[189,426,1647,1673]" box="[189,426,1647,1673]" pageId="4" pageNumber="5">
<heading id="B56A7572574BFF9AFF379534FE33F93E" bold="true" box="[189,426,1647,1673]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="DCE91E0C574BFF9AFF379534FE33F93E" bold="true" box="[189,426,1647,1673]" pageId="4" pageNumber="5">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="EE22C21E574BFF9AFF6095C5FBC3F948" blockId="4.[189,1398,1694,1791]" pageId="4" pageNumber="5">
<materialsCitation id="5EF5C843574BFF9AFF6095C5FBC3F948" ID-GBIF-Occurrence="4399003317" collectingDate="2012-12-09" collectingMethod="arm. [armadilha] Luminosa" collectionCode="DZUP" collectorName="G. Melo &amp; P. Grossi" country="Brazil" county="Brasil" elevation="1520" latitude="-22.881" location="Morro do Lopo" longLatPrecision="75" longitude="-46.308" municipality="Extrema" pageId="4" pageNumber="5" specimenCode="DZUP 215484" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<emphasis id="DCE91E0C574BFF9AFF6095C5FEC5F90F" bold="true" box="[234,348,1694,1720]" pageId="4" pageNumber="5">
<typeStatus id="31267CBC574BFF9AFF6095C5FEC5F90F" box="[234,348,1694,1720]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="968A828E574BFF9AFF37959AFEBFF96C" box="[189,294,1729,1755]" name="Brazil" pageId="4" pageNumber="5">BRAZIL</collectingCountry>
•
<specimenCount id="F89B0997574BFF9AFEC0959AFEFBF96C" box="[330,354,1729,1755]" pageId="4" pageNumber="5" type="male">♂</specimenCount>
, “Brasil,
<collectingRegion id="2C590CFC574BFF9AFE56959AFD1BF96C" box="[476,642,1729,1755]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="5">Minas Gerais</collectingRegion>
, \
<collectingMunicipality id="0E465864574BFF9AFD23959AFC96F96C" box="[681,783,1729,1755]" pageId="4" pageNumber="5">Extrema</collectingMunicipality>
,
<location id="EB4294C5574BFF9AFCAA959AFC7DF96C" LSID="urn:lsid:plazi:treatment:66347308574BFF98FD0C9721FEC1F833:EB4294C5574BFF9AFCAA959AFC7DF96C" box="[800,996,1729,1755]" country="Brazil" county="Brasil" latitude="-22.881" longLatPrecision="75" longitude="-46.308" municipality="Extrema" name="Morro do Lopo" pageId="4" pageNumber="5" stateProvince="Minas Gerais">Morro do Lopo</location>
, \
<geoCoordinate id="8BA9A4D9574BFF9AFB80959AFBEDF96C" box="[1034,1140,1729,1755]" degrees="22.881" direction="south" orientation="latitude" pageId="4" pageNumber="5" precision="55" value="-22.881">22.881ºS</geoCoordinate>
<geoCoordinate id="8BA9A4D9574BFF9AFB0B959AFB6EF96C" box="[1153,1271,1729,1755]" degrees="46.308" direction="west" orientation="longitude" pageId="4" pageNumber="5" precision="55" value="-46.308">46.308ºW</geoCoordinate>
, \
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<elevation id="65B0252D574BFF9AFA94959BFAEBF96C" box="[1310,1394,1728,1755]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="4" pageNumber="5" unit="m" value="1520.0">1520m</elevation>
</quantity>
,
<date id="9A23E4DE574BFF9AFF3795BFFEAEF949" box="[189,311,1764,1790]" pageId="4" pageNumber="5" value="2012-12-09">
<collectingDate id="8A671D36574BFF9AFF3795BFFEAEF949" box="[189,311,1764,1790]" pageId="4" pageNumber="5" value="2012-12-09">9.xii.2012</collectingDate>
</date>
,
<collectorName id="4368A7C8574BFF9AFEC995BFFE3DF949" box="[323,420,1764,1790]" pageId="4" pageNumber="5">G. Melo</collectorName>
\ &amp;
<collectorName id="4368A7C8574BFF9AFE5395BFFDDCF949" box="[473,581,1764,1790]" pageId="4" pageNumber="5">P. Grossi</collectorName>
,
<collectingMethod id="37DCBA09574BFF9AFDDA95BFFC08F949" box="[592,913,1764,1790]" pageId="4" pageNumber="5">arm.[armadilha] Luminosa</collectingMethod>
”;
<specimenCode id="BE3B6A65574BFF9AFC2D95BFFBCFF948" box="[935,1110,1764,1791]" collectionCode="DZUP" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="4" pageNumber="5" type="University or college">DZUP 215484</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A6879195574BFF98FF37947EFEA0F933" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
<paragraph id="EE22C21E574BFF9AFF37947EFED7F888" blockId="4.[189,334,1829,1855]" box="[189,334,1829,1855]" pageId="4" pageNumber="5">
<heading id="B56A7572574BFF9AFF37947EFED7F888" bold="true" box="[189,334,1829,1855]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="DCE91E0C574BFF9AFF37947EFED7F888" bold="true" box="[189,334,1829,1855]" pageId="4" pageNumber="5">Description</emphasis>
</heading>
</paragraph>
<paragraph id="EE22C21E574BFF9AFF60940FFEB3F8D9" blockId="4.[189,808,1876,1938]" box="[234,298,1876,1902]" pageId="4" pageNumber="5">
<emphasis id="DCE91E0C574BFF9AFF60940FFEB3F8D9" bold="true" box="[234,298,1876,1902]" pageId="4" pageNumber="5">Male</emphasis>
</paragraph>
<paragraph id="EE22C21E574BFF9AFF379423FCB1F825" blockId="4.[189,808,1876,1938]" box="[189,808,1911,1938]" pageId="4" pageNumber="5">
<smallCapsWord id="E8C454C2574BFF9AFF379423FEF7F827" baselines="1931,1931" box="[189,366,1912,1938]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="4" pageNumber="5">MEASUREMENTS</smallCapsWord>
.
<typeStatus id="31267CBC574BFF9AFEF7942CFE73F826" box="[381,490,1911,1937]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
male: total length
<quantity id="29656FFB574BFF9AFD4C942CFCBAF825" box="[710,803,1911,1938]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="5" unit="mm" value="6.8">6.8 mm</quantity>
.
</paragraph>
<caption id="BAE29296574AFF9BFF37941AFC1FF85E" ID-DOI="http://doi.org/10.5281/zenodo.8269453" ID-Zenodo-Dep="8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1857,1883]" targetBox="[190,1398,269,1810]" targetPageId="5">
<paragraph id="EE22C21E574AFF9BFF37941AFC1FF85E" blockId="5.[186,1398,1857,2025]" pageId="5" pageNumber="6">
<emphasis id="DCE91E0C574AFF9BFF37941AFE93F8EC" bold="true" box="[189,266,1857,1883]" pageId="5" pageNumber="6">Fig. 1.</emphasis>
<taxonomicName id="299DB99D574AFF9BFE859419FE49F8EC" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" box="[271,464,1857,1884]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="alata" status="sp. nov.">
<emphasis id="DCE91E0C574AFF9BFE859419FE49F8EC" box="[271,464,1857,1884]" italics="true" pageId="5" pageNumber="6">Costanana alata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7DAA377574AFF9BFE5F9419FDA8F8EB" box="[469,561,1858,1884]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (DZUP 215484).
<emphasis id="DCE91E0C574AFF9BFC19941AFC31F8EC" bold="true" box="[915,936,1857,1883]" pageId="5" pageNumber="6">A</emphasis>
. Head and thorax, dorsal view.
<emphasis id="DCE91E0C574AFF9BFA859419FABAF8EB" bold="true" box="[1295,1315,1858,1884]" pageId="5" pageNumber="6">B</emphasis>
. Head, ventral view.
<emphasis id="DCE91E0C574AFF9BFEEB943FFEEDF8C9" bold="true" box="[353,372,1892,1918]" pageId="5" pageNumber="6">C</emphasis>
. Head and thorax, lateral view.
<emphasis id="DCE91E0C574AFF9BFD7F943EFC91F8C8" bold="true" box="[757,776,1893,1919]" pageId="5" pageNumber="6">D</emphasis>
. Forewing.
<emphasis id="DCE91E0C574AFF9BFC16943EFC41F8C8" bold="true" box="[924,984,1893,1919]" pageId="5" pageNumber="6">E–M</emphasis>
. Male terminalia.
<emphasis id="DCE91E0C574AFF9BFB3D943EFB50F8C8" bold="true" box="[1207,1225,1893,1919]" pageId="5" pageNumber="6">E</emphasis>
. Sternite VIII, ventral view.
<emphasis id="DCE91E0C574AFF9BFED094D3FEF5F815" bold="true" box="[346,364,1928,1954]" pageId="5" pageNumber="6">F</emphasis>
. Valve, anterior view.
<emphasis id="DCE91E0C574AFF9BFDF894D3FD10F815" bold="true" box="[626,649,1928,1954]" pageId="5" pageNumber="6">G</emphasis>
. Pygofer, valve and subgenital plate, lateral view.
<emphasis id="DCE91E0C574AFF9BFB5994D3FB73F815" bold="true" box="[1235,1258,1928,1954]" pageId="5" pageNumber="6">H</emphasis>
. Subgenital plate, ventral view.
<emphasis id="DCE91E0C574AFF9BFE1594F0FE33F872" bold="true" box="[415,426,1963,1989]" pageId="5" pageNumber="6">I</emphasis>
. Connective, dorsal view.
<emphasis id="DCE91E0C574AFF9BFD5594F0FD77F872" bold="true" box="[735,750,1963,1989]" pageId="5" pageNumber="6">J</emphasis>
. Style, dorsal view.
<emphasis id="DCE91E0C574AFF9BFC5294F0FC76F872" bold="true" box="[984,1007,1963,1989]" pageId="5" pageNumber="6">K</emphasis>
. Style, lateral view.
<emphasis id="DCE91E0C574AFF9BFB5194F0FB76F872" bold="true" box="[1243,1263,1963,1989]" pageId="5" pageNumber="6">L</emphasis>
. Aedeagus, lateral view.
<emphasis id="DCE91E0C574AFF9BFED99495FEF6F85F" bold="true" box="[339,367,1998,2024]" pageId="5" pageNumber="6">M</emphasis>
. Aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
<paragraph id="EE22C21E5749FF98FF379252FDB9FDD3" blockId="6.[188,1400,265,613]" pageId="6" pageNumber="7">
<smallCapsWord id="E8C454C25749FF98FF379252FED1FE96" baselines="284,284" box="[189,328,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="6" pageNumber="7">COLORATION</smallCapsWord>
. Dorsal portion of head and thorax reddish-brown (
<figureCitation id="76A6DE9B5749FF98FC179252FB86FE94" box="[925,1055,265,291]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Figs 1A, C</figureCitation>
,
<figureCitation id="76A6DE9B5749FF98FBA09252FBC6FE94" box="[1066,1119,265,291]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[189,1398,742,1851]" captionTargetId="figure-195@26.[189,1398,742,1851]" captionTargetPageId="26" captionText="Fig. 11. Habitus of new species of Costanana DeLong &amp; Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. alata sp. nov., holotype, ♂ (DZUP 215484). C–D. C. bifida sp. nov., holotype, ♂ (DZUP 215493). E–F. C. cifi sp. nov., holotype, ♂ (DZUP 215484). G–H. C. luzi sp. nov., holotype ♂ (DZUP 215489). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269485" httpUri="https://zenodo.org/record/8269485/files/figure.png" pageId="6" pageNumber="7">11A</figureCitation>
); ventral portion yellow (
<figureCitation id="76A6DE9B5749FF98FF4E9276FEDFFEF0" box="[196,326,301,327]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Figs 1B–C</figureCitation>
,
<figureCitation id="76A6DE9B5749FF98FED99276FE1FFEF0" box="[339,390,301,327]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[189,1398,742,1851]" captionTargetId="figure-195@26.[189,1398,742,1851]" captionTargetPageId="26" captionText="Fig. 11. Habitus of new species of Costanana DeLong &amp; Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. alata sp. nov., holotype, ♂ (DZUP 215484). C–D. C. bifida sp. nov., holotype, ♂ (DZUP 215493). E–F. C. cifi sp. nov., holotype, ♂ (DZUP 215484). G–H. C. luzi sp. nov., holotype ♂ (DZUP 215489). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269485" httpUri="https://zenodo.org/record/8269485/files/figure.png" pageId="6" pageNumber="7">11B</figureCitation>
). Crown (
<figureCitation id="76A6DE9B5749FF98FE709276FDCFFEF0" box="[506,598,301,327]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
) with black maculae surrounding ocelli; pair of rounded black spots behind ocelli, near posterior margin; median line and posterior margin light brown; eyes (in life) and ocelli red. Pronotum (
<figureCitation id="76A6DE9B5749FF98FE42922FFDBFFE39" box="[456,550,372,398]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
) with black irregular maculae and light brown spots near anterior and lateral margins; lateral margin yellow. Mesonotum with pair of dark brown maculae near lateral angles and medially; scutellum yellowish, with brown areas adjacent to scutoscutellar suture. Face (
<figureCitation id="76A6DE9B5749FF98FA9892E7FAF4FE61" box="[1298,1389,444,470]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1B</figureCitation>
) with frons slightly brownish dorsally. Forewing (
<figureCitation id="76A6DE9B5749FF98FC8B9284FCC5FE4E" box="[769,860,479,505]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
) with narrow light yellow macula on anterior third of costal margin and a depigmented (white) macula on costal margin at level of outer anteapical cell, and apex of third and fourth apical cells; veins outlined by dark brown. Legs (
<figureCitation id="76A6DE9B5749FF98FB2D917CFA96FDF6" box="[1191,1295,551,577]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[189,1398,742,1851]" captionTargetId="figure-195@26.[189,1398,742,1851]" captionTargetPageId="26" captionText="Fig. 11. Habitus of new species of Costanana DeLong &amp; Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. alata sp. nov., holotype, ♂ (DZUP 215484). C–D. C. bifida sp. nov., holotype, ♂ (DZUP 215493). E–F. C. cifi sp. nov., holotype, ♂ (DZUP 215484). G–H. C. luzi sp. nov., holotype ♂ (DZUP 215489). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269485" httpUri="https://zenodo.org/record/8269485/files/figure.png" pageId="6" pageNumber="7">Fig. 11B</figureCitation>
) yellow, apex of metatibia dark brown.
</paragraph>
<paragraph id="EE22C21E5749FF98FF3791D6FD25FC73" blockId="6.[188,1401,652,965]" pageId="6" pageNumber="7">
<smallCapsWord id="E8C454C25749FF98FF3791D6FEA3FD12" baselines="672,672" box="[189,314,653,679]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Structure" pageId="6" pageNumber="7">STRUCTURE</smallCapsWord>
. Head in dorsal view (
<figureCitation id="76A6DE9B5749FF98FDC491D7FD37FD10" box="[590,686,652,679]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
): median length of crown as long as half interocular width; transocular width of head eight tenths humeral width of pronotum; in lateral view (
<figureCitation id="76A6DE9B5749FF98FB4391EBFAB1FD7D" box="[1225,1320,688,714]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1C</figureCitation>
), with anterior margin moderately thick, with 5–6 well developed carinae. Forewing (
<figureCitation id="76A6DE9B5749FF98FB0C918FFB7CFD59" box="[1158,1253,724,750]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
) with inner discal cell very short, one-third as long as the outer discal cell; apex slightly tapered. Profemur AV row with 3–4 setae restricted to basal half and PV row with 2 setae restricted to distal half. Protibia PD row with 3 long setae and apical PD
<subScript id="7219C05B5749FF98FD1E9009FD04FCD7" attach="left" box="[660,669,850,864]" fontSize="6" pageId="6" pageNumber="7">1</subScript>
seta developed; PV row developed, with 8 setae increasing in thickness and length towards apex. Metatibia rows PD, AD, and AV with 21, 12, and 12 macrosetae, respectively. Metatarsomere I apex with 3 platellae. Metatarsomere II apex with 2 apical platellae. Other characteristics as in the generic description.
</paragraph>
<paragraph id="EE22C21E5749FF98FF3790B6FAFFF9A9" blockId="6.[188,1400,1004,1567]" pageId="6" pageNumber="7">
<smallCapsWord id="E8C454C25749FF98FF3790B6FF66FBB2" baselines="1024,1024" box="[189,255,1005,1031]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="6" pageNumber="7">MALE</smallCapsWord>
<smallCapsWord id="E8C454C25749FF98FE8690A9FE14FBB2" baselines="1024" box="[268,397,1010,1029]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="terminalia" pageId="6" pageNumber="7">TERMINALIA</smallCapsWord>
. Sternite VIII (
<figureCitation id="76A6DE9B5749FF98FDC590B7FD32FBB1" box="[591,683,1004,1030]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1E</figureCitation>
) 1.2× as wide as long; lateral margins excavated at base, forming a triangular projection and then converging towards apex, narrow and rounded. Valve (
<figureCitation id="76A6DE9B5749FF98FA9D974BFAF5FB9D" box="[1303,1388,1040,1066]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1F</figureCitation>
) 3.1× as wide as long; posterior margin almost straight. Pygofer (
<figureCitation id="76A6DE9B5749FF98FC2C976FFB98FBF9" box="[934,1025,1076,1102]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1G</figureCitation>
), in lateral view, 2.3× as long as maximum height; ventral margin with a small protrusion near base; posterodorsal and ventral margins rounded; external surface near ventral margin with short filiform setae; macrosetae distributed on apical half; apex slightly tapered and rounded. Subgenital plate (
<figureCitation id="76A6DE9B5749FF98FCD697C4FC2EFB0E" box="[860,951,1183,1209]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1G</figureCitation>
), in lateral view, short, surpassing half length of pygofer; in ventral view (
<figureCitation id="76A6DE9B5749FF98FDD29798FD2EFB6A" box="[600,695,1219,1245]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1H</figureCitation>
), 5× as long as wide; inner margin straight; external margin rounded on basal third; ventral surface and external margin with long filiform setae; apex strongly tapered and weakly sclerotized. Connective (
<figureCitation id="76A6DE9B5749FF98FD5F9651FCBFFA93" box="[725,806,1290,1316]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1I</figureCitation>
) Y-shaped; arms weakly produced laterally; stalk about as long as maximum width of arms; median keel weakly developed. Style (
<figureCitation id="76A6DE9B5749FF98FB2A9675FB6AFAFF" box="[1184,1267,1326,1352]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1J</figureCitation>
), in dorsal view, with outer lobe small and rounded; inner margin with a small dentiform process near midlength of blade; in lateral view (
<figureCitation id="76A6DE9B5749FF98FE4E962EFDB9FA27" box="[452,544,1397,1424]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1K</figureCitation>
), ventral margin not serrated, with dentiform process near base of blade; apex truncated, forming a pair of acute processes directed dorsally and ventrally. Aedeagus (
<figureCitation id="76A6DE9B5749FF98FB6796C2FAF7FA04" box="[1261,1390,1433,1459]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1L–M</figureCitation>
) preatrium not developed; dorsal apodeme strongly elongated laterally; shaft short, nearly tubular, with pair of ventral wing-shaped processes near base, curved dorsally with divergent and acute apices; dorsal margin of shaft with slender basodorsal process; ventral margin with pair of minute spurs subapically.
</paragraph>
<paragraph id="EE22C21E5749FF98FF60951DFEDCF9D7" blockId="6.[189,325,1606,1668]" box="[234,325,1606,1632]" pageId="6" pageNumber="7">
<emphasis id="DCE91E0C5749FF98FF60951DFEDCF9D7" bold="true" box="[234,325,1606,1632]" pageId="6" pageNumber="7">Female</emphasis>
</paragraph>
<paragraph id="EE22C21E5749FF98FF379531FEA0F933" blockId="6.[189,325,1606,1668]" box="[189,313,1642,1668]" pageId="6" pageNumber="7">Unknown.</paragraph>
</subSubSection>
<subSubSection id="A68791955749FF98FF3795F7FEC1F833" pageId="6" pageNumber="7" type="discussion">
<paragraph id="EE22C21E5749FF98FF3795F7FEB6F971" blockId="6.[189,303,1708,1734]" box="[189,303,1708,1734]" pageId="6" pageNumber="7">
<heading id="B56A75725749FF98FF3795F7FEB6F971" bold="true" box="[189,303,1708,1734]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="DCE91E0C5749FF98FF3795F7FEB6F971" bold="true" box="[189,303,1708,1734]" pageId="6" pageNumber="7">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="EE22C21E5749FF98FF379587FEC1F833" blockId="6.[188,1401,1755,1924]" pageId="6" pageNumber="7">
<taxonomicName id="299DB99D5749FF98FF379587FE18F942" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" box="[189,385,1755,1782]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="alata" status="sp. nov.">
<emphasis id="DCE91E0C5749FF98FF379587FE18F942" box="[189,385,1755,1782]" italics="true" pageId="6" pageNumber="7">Costanana alata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7DAA3775749FF98FE019587FE71F941" box="[395,488,1756,1782]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="299DB99D5749FF98FD479587FCDBF942" authorityName="DeLong &amp; Freytag" authorityYear="1972" box="[717,834,1755,1782]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="flavina">
<emphasis id="DCE91E0C5749FF98FD479587FCDBF942" box="[717,834,1755,1782]" italics="true" pageId="6" pageNumber="7">C. flavina</emphasis>
</taxonomicName>
in having the same color pattern (
<figureCitation id="76A6DE9B5749FF98FB549580FAEAF942" box="[1246,1395,1755,1781]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[189,1398,742,1851]" captionTargetId="figure-195@26.[189,1398,742,1851]" captionTargetPageId="26" captionText="Fig. 11. Habitus of new species of Costanana DeLong &amp; Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. alata sp. nov., holotype, ♂ (DZUP 215484). C–D. C. bifida sp. nov., holotype, ♂ (DZUP 215493). E–F. C. cifi sp. nov., holotype, ♂ (DZUP 215484). G–H. C. luzi sp. nov., holotype ♂ (DZUP 215489). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269485" httpUri="https://zenodo.org/record/8269485/files/figure.png" pageId="6" pageNumber="7">Figs 11A–B</figureCitation>
.
<figureCitation id="76A6DE9B5749FF98FF3595A4FE8CF8AE" box="[191,277,1791,1817]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,1029,1884]" captionTargetId="figure-226@29.[189,1398,1029,1885]" captionTargetPageId="29" captionText="Fig. 13. Habitus of species of Costanana DeLong &amp; Freytag, 1972. Left, dorsal view; right, lateral view. A–B. C. flavina DeLong &amp; Freytag, 1972 (DZUP). C–D. C. praecellens (Stål, 1862) (DZUP). E–F. C. santana DeLong &amp; Wolda, 1983 (DZUP). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269491" httpUri="https://zenodo.org/record/8269491/files/figure.png" pageId="6" pageNumber="7">13A–B</figureCitation>
) and the aedeagal shaft short with a pair of ventral processes near the base, not contiguous to shaft and following its curvature. The new species can easily be separated from
<taxonomicName id="299DB99D5749FF98FBD59478FB48F88B" authorityName="DeLong &amp; Freytag" authorityYear="1972" box="[1119,1233,1826,1853]" class="Insecta" family="Cicadellidae" genus="Costanana" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="flavina">
<emphasis id="DCE91E0C5749FF98FBD59478FB48F88B" box="[1119,1233,1826,1853]" italics="true" pageId="6" pageNumber="7">C. flavina</emphasis>
</taxonomicName>
by having the apex of the style truncated (
<figureCitation id="76A6DE9B5749FF98FD8E941DFDF9F8D7" box="[516,608,1862,1888]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1K</figureCitation>
) and the longer and winged ventral processes of the aedeagal shaft (
<figureCitation id="76A6DE9B5749FF98FF4C9431FED2F833" box="[198,331,1898,1924]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[190,1398,269,1810]" captionTargetId="figure-11@5.[190,1398,265,1815]" captionTargetPageId="5" captionText="Fig. 1. Costanana alata sp. nov., holotype, ♂ (DZUP 215484). A. Head and thorax, dorsal view.B. Head, ventral view. C. Head and thorax, lateral view. D. Forewing. E–M. Male terminalia. E. Sternite VIII, ventral view. F. Valve, anterior view. G. Pygofer, valve and subgenital plate, lateral view. H. Subgenital plate, ventral view. I. Connective, dorsal view. J. Style, dorsal view. K. Style, lateral view. L. Aedeagus, lateral view. M. Aedeagus, posterior view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.8269453" httpUri="https://zenodo.org/record/8269453/files/figure.png" pageId="6" pageNumber="7">Fig. 1L–M</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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