393 lines
50 KiB
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393 lines
50 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1063.69942" ID-GBIF-Dataset="28e9e42b-c1df-47e6-a327-6e4cbc06db46" ID-PMC="PMC8545786" ID-Pensoft-Pub="1313-2970-1063-1" ID-Pensoft-UUID="94FEB0FF5E515631AA6239A1BB296EEB" ID-PubMed="34720622" ID-ZooBank="3687D7229CD64D82ACF8D6701E399F4B" ModsDocID="1313-2970-1063-1" checkinTime="1634632146073" checkinUser="pensoft" docAuthor="Yin, Wen-Qi, Shi, Hong-Liang & Liang, Hong-Bin" docDate="2021" docId="66079427D5165C98984E20EF8023F913" docLanguage="en" docName="ZooKeys 1063: 1-21" docOrigin="ZooKeys 1063" docPubDate="2021-10-18" docSource="http://dx.doi.org/10.3897/zookeys.1063.69942" docTitle="Pterostichus (Koreonialoe) bellatrix" docType="treatment" docVersion="5" id="94FEB0FF5E515631AA6239A1BB296EEB" lastPageNumber="1" masterDocId="94FEB0FF5E515631AA6239A1BB296EEB" masterDocTitle="Revision of the Chinese species of subgenus Koreonialoe (Coleoptera: Carabidae: Pterostichus), with descriptions of two new species" masterLastPageNumber="21" masterPageNumber="1" pageNumber="1" updateTime="1732708058111" updateUser="ExternalLinkService">
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<mods:title id="E3BD3931D9C3BB93875E04248275BE76">Revision of the Chinese species of subgenus Koreonialoe (Coleoptera: Carabidae: Pterostichus), with descriptions of two new species</mods:title>
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<mods:namePart id="90678692B209C8FFD5E163B5A2170EF1">Yin, Wen-Qi</mods:namePart>
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<mods:affiliation id="794017AFAB8F27747DA53ADA94604302">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
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<mods:namePart id="792E6D5CAAF089F53472DDFB113A7F0F">Shi, Hong-Liang</mods:namePart>
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<mods:affiliation id="E69C65DB58FF2A5E77552ABE82DE24F4">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
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<mods:nameIdentifier id="C7F6455415AEE3DB16B77FB7F14DC653" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="4E596E5EE2000A4826D21D0C54DACEF1">Liang, Hong-Bin</mods:namePart>
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<mods:affiliation id="F0AA2FA5FB20E074F74CFCAE8611E2B4">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<subSection id="368B3087C48E7476F6C03D57751C21FA" pageId="0" pageNumber="1" type="taxonomy">
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<treatment id="66079427D5165C98984E20EF8023F913" ID-GBIF-Taxon="189468674" LSID="urn:lsid:plazi:treatment:66079427D5165C98984E20EF8023F913" httpUri="http://treatment.plazi.org/id/66079427D5165C98984E20EF8023F913" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Pterostichus">
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<subSubSection id="F69F816BCDCA8F1847908B22FC2E75B4" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="632F5708EA59D39FE8D8DA0594810C38" pageId="0" pageNumber="1">
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4.
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<taxonomicName id="3E550790EA15313BF251AD00BDDE91A6" ID-CoL="4Q3TY" LSID="66079427-D516-5C98-984E-20EF8023F913" authority="(Tschitscherine, 1895)" baseAuthorityName="Tschitscherine" baseAuthorityYear="1895" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Koreonialoe) bellatrix" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellatrix" subGenus="Koreonialoe">
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Pterostichus (Koreonialoe) bellatrix (
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<normalizedToken id="895F41A5F415BABB20B2DB190C9A2990" originalValue="Tschitschérine">Tschitscherine</normalizedToken>
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, 1895)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="530D43EEC5E734F86B4BE66B48C0B90B" pageId="0" pageNumber="1" type="description">
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<paragraph id="04347DB19C3ADE98B3A4FDDF88A35306" pageId="0" pageNumber="1">
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<figureCitation id="1850BC3E164700C971B34164A7ECF161" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus of Pterostichus (Pterostichus Koreonialoe) spp. from China 1 P. micropoides sp. nov., Holotype. 2 P. micropoides sp. nov., male, Paratype from Hunchun city, Jilin 3 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 4 P. syleus Kirschenhofer, a female from Zhuanghe city, Liaoning 5 P. quadrilobatus sp. nov., Holotype 6 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1063.69942.figures1-6" httpUri="https://binary.pensoft.net/fig/599856" pageId="0" pageNumber="1">Figs 6</figureCitation>
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<figureCitation id="930ECF97EFCA22E9D998229CFA2D5578" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Male genitalia of Pterostichus (Koreonialoe) spp. from China, left lateral view of median lobe of aedeagus 7 P. micropoides sp. nov., Holotype 8 P. micropoides sp. nov., Paratype from Changbai county, Jilin 9 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 10 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 11 P. quadrilobatus sp. nov., Holotype 12 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures7-12" httpUri="https://binary.pensoft.net/fig/599857" pageId="0" pageNumber="1">, 12</figureCitation>
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<figureCitation id="E179C4636B1AF8A19A525537155CB692" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus (Koreonialoe) spp. from China, dorsal view of median lobe of aedeagus 13 P. micropoides sp. nov., Holotype 14 P. micropoides sp. nov. Paratype from Changbai county 15 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 16 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 17 P. quadrilobatus sp. nov., Paratype from Laopingding, Liaoning 18 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures13-18" httpUri="https://binary.pensoft.net/fig/599858" pageId="0" pageNumber="1">, 18</figureCitation>
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<figureCitation id="4E43C136C73D12E1638A371DA1F4B0BA" captionStart="Figures 22–27" captionStartId="F5" captionText="Figures 22 - 27. Endophallus of Pterostichus (Koreonialoe) spp. from China, left lateral view and right lateral view 22 P. micropoides sp. nov., Paratype from Antu city, Jilin 23 P. micropoides sp. nov., Paratype from Changbaishan mt., Jilin 24 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 25 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 26 P. quadrilobatus sp. nov., Holotype 27 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures22-27" httpUri="https://binary.pensoft.net/fig/599860" pageId="0" pageNumber="1">, 27</figureCitation>
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<figureCitation id="61CE681BC56B5B57466C4729D4B9DC80" captionStart="Figures 28–35" captionStartId="F6" captionText="Figures 28 - 35. Female genitalia of Pterostichus (Koreonialoe) spp. from China, Figs 28 - 31 ventral view and inner lateral view of ovipositor. Figs 32 - 35 female reproductive system. 28, 32 P. tetralobatus, Paratype from Laopingding, Liaoning 29, 33 P. syleus, a female from Baishilazi, Liaoning 30, 34 P. micropoides, female, Paratype from Antu county, Jilin 31, 35 P. bellatrix, a female from Fusong county, Jilin. Scale bar: 0.5 mm. Abbreviations: g 1: gonocoxite I; g 2: gonocoxite II; es: ensiform setae; ns, nematiform setae; bc: bursa copulatrix; co: common oviduct; sc: seminal canal; rc: receptaculum; spc: spermathecal canal." figureDoi="10.3897/zookeys.1063.69942.figures28-35" httpUri="https://binary.pensoft.net/fig/599861" pageId="0" pageNumber="1">, 28</figureCitation>
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<figureCitation id="D5DC36576A155FBE6E76046E60473218" captionStart="Figures 28–35" captionStartId="F6" captionText="Figures 28 - 35. Female genitalia of Pterostichus (Koreonialoe) spp. from China, Figs 28 - 31 ventral view and inner lateral view of ovipositor. Figs 32 - 35 female reproductive system. 28, 32 P. tetralobatus, Paratype from Laopingding, Liaoning 29, 33 P. syleus, a female from Baishilazi, Liaoning 30, 34 P. micropoides, female, Paratype from Antu county, Jilin 31, 35 P. bellatrix, a female from Fusong county, Jilin. Scale bar: 0.5 mm. Abbreviations: g 1: gonocoxite I; g 2: gonocoxite II; es: ensiform setae; ns, nematiform setae; bc: bursa copulatrix; co: common oviduct; sc: seminal canal; rc: receptaculum; spc: spermathecal canal." figureDoi="10.3897/zookeys.1063.69942.figures28-35" httpUri="https://binary.pensoft.net/fig/599861" pageId="0" pageNumber="1">, 32</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="8976E2AD74074E416958642C5A9AF9E0" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="E4298214A2436CBF7317F2DA589B8D66" pageId="0" pageNumber="1">
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||
<taxonomicName id="C50659DAD78FEFD374016562ECEA14EE" baseAuthorityName="Tschitscherine" baseAuthorityYear="1895" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Koreonialoe) bellatrix" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellatrix" subGenus="Koreonialoe">Pterostichus (Koreonialoe) bellatrix</taxonomicName>
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<normalizedToken id="E486B589B6B1184B2C5C93D4774833C9" originalValue="Tschitschérine">Tschitscherine</normalizedToken>
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, 1895: 169 (Syntypes deposited in Zoological Institute, Russian Academy of sciences, St Petersburg, Russia; type locality: North Korea, sg.
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<taxonomicName id="4E1F08789378283A33718902AF39421E" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Feronia)" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Feronia">Pterostichus Feronia</taxonomicName>
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);
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<bibRefCitation id="52A60809788FA4F89DE2C23B49B0A30A" author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlich Museum fuer Tierkunde in Dresden" pageId="0" pageNumber="1" pagination="177 - 346" refId="B6" refString="Jedlicka, A, 1962. Monographie des Tribus Pterostichini aus Ostasien (Pterostichi, Trigonotomi, Myadi) (Coleoptera - Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlich Museum fuer Tierkunde in Dresden 26: 177 - 346" title="Monographie des Tribus Pterostichini aus Ostasien (Pterostichi, Trigonotomi, Myadi) (Coleoptera - Carabidae)." volume="26" year="1962">
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<normalizedToken id="0B80E4A279516B8C80773326C69CC6F1" originalValue="Jedlička">Jedlicka</normalizedToken>
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1962
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</bibRefCitation>
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: 271 (misspelled as
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<taxonomicName id="032607C7E4521C2A1D93FE5E380264E0" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellator" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellator">P. bellator</taxonomicName>
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, sg.
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<taxonomicName id="66FCF528C16FD1F5DF9A63C28C577F79" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Lianoe)" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Lianoe">Pterostichus Lianoe</taxonomicName>
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);
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<bibRefCitation id="2E27FBDE2FAF1EEE751576AA65A8B076" author="Kwon, YJ" journalOrPublisher="Insecta Koreana," pageId="0" pageNumber="1" pagination="1 - 56" refId="B9" refString="Kwon, YJ, Lee, SM, 1986. Check list of superfamily Caraboidea from Korea. Insecta Koreana, 6: 1 - 56" title="Check list of superfamily Caraboidea from Korea." volume="6" year="1986">Kwon and Lee 1986</bibRefCitation>
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: 28 (misspelled as
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<taxonomicName id="6C8D4B48488924220A4831685E63DBF1" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellator" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellator">P. bellator</taxonomicName>
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, sg.
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<taxonomicName id="7C71881B28B955767AA5BEEA26F36342" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe)" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Nialoe">Pterostichus Nialoe</taxonomicName>
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);
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<bibRefCitation id="5F63F84A3141CEA0B15C50CDB907383C" author="Park, JK" journalOrPublisher="Korean Journal of Entomology" pageId="0" pageNumber="1" pagination="93 - 103" refId="B13" refString="Park, JK, Kwon, YJ, 1996. A new subgenus of the Pterostichus from Korea (Coleoptera: Harpalidae). Korean Journal of Entomology 26 (1): 93 - 103" title="A new subgenus of the Pterostichus from Korea (Coleoptera: Harpalidae)." volume="26" year="1996">Park and Kwon 1996</bibRefCitation>
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: 96 (misspelled as
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<taxonomicName id="8CD2E4F8B81B4D904CFE65BE0B3D11E2" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellator" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellator">P. bellator</taxonomicName>
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);
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<bibRefCitation id="0DC012F007BD04EB32223166F47A08D4" DOI="https://doi.org/10.2108/zsj.22.1205" author="Sasakawa, K" journalOrPublisher="Zoological Science" pageId="0" pageNumber="1" pagination="1205 - 1216" refId="B16" refString="Sasakawa, K, 2005a. Phylogenetic studies of the Nialoe (s. lat.) (Coleoptera, Carabidae, genus Pterostichus), Part 1: homology of the component parts of male genitalia and higher phylogeny. Zoological Science 22: 1205 - 1216, DOI: https://doi.org/10.2108/zsj.22.1205" title="Phylogenetic studies of the Nialoe (s. lat.) (Coleoptera, Carabidae, genus Pterostichus), Part 1: homology of the component parts of male genitalia and higher phylogeny." url="https://doi.org/10.2108/zsj.22.1205" volume="22" year="2005 a">Sasakawa 2005a</bibRefCitation>
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: 1209 (misspelled as
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<taxonomicName id="FCEA052DA4206EB246B0E283E5A75021" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellator" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellator">P. bellator</taxonomicName>
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);
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<bibRefCitation id="D2775E78FB5D56E26D8CCB0D231ECAF7" DOI="https://doi.org/10.1016/j.aspen.2013.05.003" author="Sasakawa, K" journalOrPublisher="Zoological Science" pageId="0" pageNumber="1" refId="B18" refString="Sasakawa, K, Kim, JK, Kim, JK, Kubota, K, 2013. Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16(4): 429-431. https://doi.org/10.1016/j.aspen.2013.05.003" title="Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16 (4): 429 - 431." url="https://doi.org/10.1016/j.aspen.2013.05.003" year="2013">Sasakawa et al. 2013</bibRefCitation>
|
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: 430;
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||
<bibRefCitation id="2130A29D9DECE8571C797CF5D10B7ECA" author="Park, JK" journalOrPublisher="Korean Journal of Entomology" pageId="0" pageNumber="1" refId="B14" refString="Park, JK, Park, JY, 2013. Insect Fauna of Korea, Volume 12, Number 13: 1-98." title="Insect Fauna of Korea, Volume 12, Number 13: 1 - 98." year="2013">Park and Park 2013</bibRefCitation>
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: 66.
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<emphasis id="2910D125E1D6897A4216BA887A1F9EA2" bold="true" pageId="0" pageNumber="1">New record from China</emphasis>
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</paragraph>
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<paragraph id="90CEF4410921039E0B7A63CECC2BC897" pageId="0" pageNumber="1">
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<taxonomicName id="17DB863081C6D86BD62AD6AAA0F57F4F" authorityName="Jedlicka" authorityYear="1931" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus klickai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="klickai">Pterostichus klickai</taxonomicName>
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<normalizedToken id="CB77C464FDF7741A31C49580C9FD7391" originalValue="Jedlička">Jedlicka</normalizedToken>
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, 1931: 104. (Holotype deposited in Narodni Muzeum Prirodovedecke Muzeum, Prague, Czech Republic, Type locality: Seishin, Korea);
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<bibRefCitation id="0CF622E92DC7088F3E0115B4463E54D5" author="Cho, PS" journalOrPublisher="Human Immunology Korea University" pageId="0" pageNumber="1" pagination="173 - 338" refId="B4" refString="Cho, PS, 1957. A systematic catalogue of Korean Coleoptera. Human Immunology Korea University 2: 173 - 338" title="A systematic catalogue of Korean Coleoptera." volume="2" year="1957">Cho 1957</bibRefCitation>
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: 194;
|
||
<bibRefCitation id="8E9D3C8BFB698B6A8CF92FD9F11E0F9D" author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlich Museum fuer Tierkunde in Dresden" pageId="0" pageNumber="1" pagination="177 - 346" refId="B6" refString="Jedlicka, A, 1962. Monographie des Tribus Pterostichini aus Ostasien (Pterostichi, Trigonotomi, Myadi) (Coleoptera - Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlich Museum fuer Tierkunde in Dresden 26: 177 - 346" title="Monographie des Tribus Pterostichini aus Ostasien (Pterostichi, Trigonotomi, Myadi) (Coleoptera - Carabidae)." volume="26" year="1962">
|
||
<normalizedToken id="24F5E5DD499D13357BBBC325C961E40E" originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
1962
|
||
</bibRefCitation>
|
||
: 272 (sg.
|
||
<taxonomicName id="F7265B56CD5F4773FFB6C857B337F597" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Lianoe)" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Lianoe">Pterostichus Lianoe</taxonomicName>
|
||
);
|
||
<bibRefCitation id="CE2C90DE4FAECD1065DC8C47AB31069D" author="Kwon, YJ" journalOrPublisher="Insecta Koreana," pageId="0" pageNumber="1" pagination="1 - 56" refId="B9" refString="Kwon, YJ, Lee, SM, 1986. Check list of superfamily Caraboidea from Korea. Insecta Koreana, 6: 1 - 56" title="Check list of superfamily Caraboidea from Korea." volume="6" year="1986">Kwon and Lee 1986</bibRefCitation>
|
||
: 28;
|
||
<bibRefCitation id="1EADB7F702943BDD2D5C97147B7A2BD8" author="Nemoto, K" journalOrPublisher="Bulletin of the Biogeographical Society of Japan" pageId="0" pageNumber="1" pagination="39 - 46" refId="B12" refString="Nemoto, K, 1988. Pterostichus opacipennis Jedlicka and its allied species of South Korea and Japan (Coleoptera, Carabidae). Bulletin of the Biogeographical Society of Japan 43: 39 - 46" title="Pterostichus opacipennis Jedlicka and its allied species of South Korea and Japan (Coleoptera, Carabidae)." volume="43" year="1988">Nemoto 1988</bibRefCitation>
|
||
: 40;
|
||
<bibRefCitation id="A945D346C2A3B0A33AF61EC1B7551635" author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="Archostemata-Myxophaga-Adephaga, 1. Apollo Books, Stenstrup" pageId="0" pageNumber="1" pagination="469 - 520" refId="B2" refString="Bousquet, Y, 2003. Tribe Pterostichini Bonelli, 1810. In: Loebl, I, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Archostemata-Myxophaga-Adephaga, 1. Apollo Books, Stenstrup: 469 - 520" title="Tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Bousquet 2003</bibRefCitation>
|
||
: 497 (synonymized with
|
||
<taxonomicName id="945B4F8ABC697AC9BDBDFAF785446620" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellatrix" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellatrix">P. bellatrix</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6EAF8B38F906F31471E0B24E06293BA9" pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph id="D2DAC3910984CA5656CB242AA4EEC9F5" pageId="0" pageNumber="1">Material examined.</paragraph>
|
||
<paragraph id="6C460652AF9E3E6173B6A4425B4EC489" pageId="0" pageNumber="1">
|
||
(
|
||
<specimenCount id="6C729A9ACA993F81EFC43C92BD17031E" type="male">6 males</specimenCount>
|
||
and
|
||
<specimenCount id="D70AFFCD44420EF11EFD8596473BC4FF" type="female">1 female</specimenCount>
|
||
):
|
||
<specimenCount id="FB219CDB35AC22E30DB7640E0E2B855F" type="male">1 male</specimenCount>
|
||
,
|
||
<materialsCitation id="6B5113BEA4C9A163C2BBA591C687CC21" collectingDate="2019-08-09" collectingMethod="pitfall trap" country="China" county="Changbai County" elevation="1330" latitude="41.7398" location="Mixed forest" municipality="Baishan City" specimenCount="1" stateProvince="Jilin province">
|
||
"
|
||
<collectingCountry id="85331AA0CF298D7C6479156F49BCEB20" name="China">China</collectingCountry>
|
||
<collectingRegion id="39BD3F6EA01955020D844B3ED83061A3" country="China" name="Jilin">Jilin province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BBD5B942AC7500DD2B9CD8E38074ACA">Baishan City</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="115DFA0C4DD5D5727A255E2A0C776767">Changbai County</collectingCounty>
|
||
,
|
||
<locationDeviation id="9DC155827F3F835962AF906DF0309B57" location="Changsongling">E of Changsongling</locationDeviation>
|
||
tunnel;
|
||
<location id="763EACC6AE2890F848BC39290D4561A3" LSID="urn:lsid:plazi:treatment:66079427D5165C98984E20EF8023F913:763EACC6AE2890F848BC39290D4561A3" country="China" county="Changbai County" latitude="41.7398" municipality="Baishan City" name="Mixed forest" stateProvince="Jilin province">Mixed forest</location>
|
||
,
|
||
<geoCoordinate id="F0E6B604E787BD79A32892F12B82411C" degrees="41.7398" direction="north" orientation="latitude" precision="5" value="41.7398">41.7398N</geoCoordinate>
|
||
, 128.0221,
|
||
<quantity id="0BAEC8F2754D7EF3778842D4B06A238F" metricMagnitude="3" metricUnit="m" metricValue="1.33" unit="m" value="1330.0">
|
||
<elevation id="F725B95868D8F863AC37CDFD2BD4278A" metricMagnitude="3" metricUnit="m" metricValue="1.33" unit="m" value="1330.0">1330m</elevation>
|
||
</quantity>
|
||
"; "
|
||
<collectingMethod id="1EA403C4CAB77BB7491D1F50DF250249">pitfall trap</collectingMethod>
|
||
;
|
||
<collectingDate id="56ACA40FB90367CE24D9F7B086FD87F6" value="2019-08-09">2019.VIII.9</collectingDate>
|
||
, Shi HL & Liu YZ lgt."
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="1F565A58C705B1D0D13AA389586D85A7" type="male">4 males</specimenCount>
|
||
and
|
||
<specimenCount id="7D15845CFBCD14FEE407CC70A8CBA453" type="female">1 female</specimenCount>
|
||
,
|
||
<materialsCitation id="8034C85A53F8862BB2E2038BCA88DE24" collectingDate="2019-08-09" collectingMethod="pitfall trap" country="China" elevation="1577" latitude="41.7798" location="Fusong county" longLatPrecision="6" longitude="127.94" municipality="Baishan City" specimenCount="1" stateProvince="Jilin province">
|
||
"
|
||
<collectingCountry id="FEBA9DDCBFF46FF43850438ABE4BEA22" name="China">China</collectingCountry>
|
||
<collectingRegion id="58A2864221CC3FC37E5012ECBC20449B" country="China" name="Jilin">Jilin province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="DEF47855C137EA50E29CFD861BC4783F">Baishan City</collectingMunicipality>
|
||
,
|
||
<location id="C98432C52217837376B0336BB79AD648" LSID="urn:lsid:plazi:treatment:66079427D5165C98984E20EF8023F913:C98432C52217837376B0336BB79AD648" country="China" latitude="41.7798" longLatPrecision="6" longitude="127.94" municipality="Baishan City" name="Fusong county" stateProvince="Jilin province">Fusong county</location>
|
||
,
|
||
<locationDeviation id="A744C52EBF686E84D8ED98F4D78490EB" location="Changsongling">W of Changsongling</locationDeviation>
|
||
tunnel; mixed forest,
|
||
<geoCoordinate id="0792F8809BEB4DBF2D120FCB93FF5FB9" degrees="41.7798" direction="north" orientation="latitude" precision="5" value="41.7798">41.7798N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6E4A9109CD9B24B35D2E113936D9D657" degrees="127.9400" direction="east" orientation="longitude" precision="5" value="127.94">127.9400E</geoCoordinate>
|
||
,
|
||
<quantity id="79E768B04BE962D353DEEB8D6646BC31" metricMagnitude="3" metricUnit="m" metricValue="1.577" unit="m" value="1577.0">
|
||
<elevation id="323A32C8A04FD2A2C8B168B40E3DD6A2" metricMagnitude="3" metricUnit="m" metricValue="1.577" unit="m" value="1577.0">1577m</elevation>
|
||
</quantity>
|
||
"; "
|
||
<collectingMethod id="929AA5D24209C49EF771BAB58210C9D2">pitfall trap</collectingMethod>
|
||
,
|
||
<collectingDate id="D422428187595BF0C4F1A34B29A9AE16" value="2019-08-09">2019.VIII.9</collectingDate>
|
||
, Shi HL & Liu YZ lgt."
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="072845DAE9842CFAFC8230F9CD3E9BDB" type="male">1 male</specimenCount>
|
||
,
|
||
<materialsCitation id="F53889C2F855C2216EF9DC1DE7522A08" collectingDate="1982-07-13" country="China" location="Changbai Mountain" specimenCount="1" stateProvince="Jilin province">
|
||
"
|
||
<collectingRegion id="55ED524B06F487D9515109F7A006BFAE" country="China" name="Jilin">Jilin province</collectingRegion>
|
||
,
|
||
<location id="6D71F4030284568A812BB66438FE1BC5" LSID="urn:lsid:plazi:treatment:66079427D5165C98984E20EF8023F913:6D71F4030284568A812BB66438FE1BC5" country="China" name="Changbai Mountain" stateProvince="Jilin province">Changbai Mountain</location>
|
||
Hot Spring,
|
||
<collectingDate id="468B503FE44E1B261B92298818583CEF" value="1982-07-13">1982.VII.13</collectingDate>
|
||
, Liao Suyi lgt."
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F63859A8AEBF0833709B7A73D71667FE" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="2DF69A6BAD2FE1E5A5B59F042FC000EE" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="14C434A24C2B257BEF09AEA4ECBC7654" pageId="0" pageNumber="1">Dorsal surface nearly black; fifth tarsomere without ventral seta. Median lobe of aedeagus stout, ventral margin strongly tumid forming a conspicuous cuneate tubercle anterior to middle. Endophallus directed basal-ventrally, gonopore opened to the ventral-basal direction of aedeagus; three lobes present: la and lp similar in size, la a little hooked apically, ra faintly defined.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2F53C903720282D856B213DE81547003" pageId="0" pageNumber="1" type="comparisons">
|
||
<paragraph id="AF5807357A57452FA9648A68FFB633BD" pageId="0" pageNumber="1">Comparison.</paragraph>
|
||
<paragraph id="075272B115C2B7ED7965E6CA0C560DFC" pageId="0" pageNumber="1">
|
||
<taxonomicName id="B45288C0ED84F038E848CEEB517372D4" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellatrix" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellatrix">
|
||
<emphasis id="FEB17F247C39282E5679A5A7CDCB8A2D" italics="true" pageId="0" pageNumber="1">P. bellatrix</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="7E8ED4645BBB3CB1FB6C816487E301E6" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="syleus">
|
||
<emphasis id="E9F3831BC543C112D8F8253A266A1D8D" italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9C270EDD08C9BD939B6DA2ED62EA7D1C" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tetralobatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tetralobatus">
|
||
<emphasis id="C10A5C9D680ACB6E7FF2ED37F7D96D84" italics="true" pageId="0" pageNumber="1">P. tetralobatus</emphasis>
|
||
</taxonomicName>
|
||
in their external features, but can be distinguished from the latter two species by the more conspicuous ventral tubercle on male genitalia. Another species,
|
||
<taxonomicName id="A1B9964BC32451830337D5BB2F0B0918" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. woongbii" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="woongbii">
|
||
<emphasis id="B9B9DD010BAF1E641FEC64E991F17A9C" italics="true" pageId="0" pageNumber="1">P. woongbii</emphasis>
|
||
</taxonomicName>
|
||
from South Korea, also has a conspicuous ventral tubercle on male genitalia, but is quite different from
|
||
<taxonomicName id="EF3F243AF0441C450E51D075B4D3FD99" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellatrix" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellatrix">
|
||
<emphasis id="25199E81B9FC074EE6DBF77C53DE854A" italics="true" pageId="0" pageNumber="1">P. bellatrix</emphasis>
|
||
</taxonomicName>
|
||
in the ventral tubercle located near base of median lobe, and endophallus short and straight.
|
||
</paragraph>
|
||
<paragraph id="196498ED0D7E351BC51FCDD1502BA7CE" pageId="0" pageNumber="1">
|
||
From the external features,
|
||
<taxonomicName id="1A71FDAE6ABA07271589F81A519A5915" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellatrix" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellatrix">
|
||
<emphasis id="C83170E93D64E365965B0F7861C19486" italics="true" pageId="0" pageNumber="1">P. bellatrix</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="0E7FCC364F57FABAF83851D5F15E3D31" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. micropoides" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="micropoides">
|
||
<emphasis id="60D1AB801ACBF9D7184F6EBC2508A479" italics="true" pageId="0" pageNumber="1">P. micropoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="36CACF25083538A12C2366550A384479" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="syleus">
|
||
<emphasis id="056EA60F2C3514B199349DDFB5A86602" italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
|
||
</taxonomicName>
|
||
for its larger size (BL 19.7-21.2 mm versus 13.7-19.7 mm in other two species) and dorsally nearly black (versus dark reddish brown in other two species). But, the females of
|
||
<taxonomicName id="8C535C8668F0DD42824712C1036A9B06" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellatrix" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellatrix">
|
||
<emphasis id="A91F70CA156073874A2A2DE6E6AA27CA" italics="true" pageId="0" pageNumber="1">P. bellatrix</emphasis>
|
||
</taxonomicName>
|
||
are completely identical to that of
|
||
<taxonomicName id="7E45CD0AD23A154DE92F5AF478BAA188" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tetralobatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tetralobatus">
|
||
<emphasis id="A416705462CC9B7F4DB54195E7E49091" italics="true" pageId="0" pageNumber="1">P. tetralobatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. They can be determined only by the allopatric distributions (Fig.
|
||
<figureCitation id="6A8AF941A18612A2ED667189F0FC306B" captionStart="Figures 28–35" captionStartId="F6" captionText="Figures 28 - 35. Female genitalia of Pterostichus (Koreonialoe) spp. from China, Figs 28 - 31 ventral view and inner lateral view of ovipositor. Figs 32 - 35 female reproductive system. 28, 32 P. tetralobatus, Paratype from Laopingding, Liaoning 29, 33 P. syleus, a female from Baishilazi, Liaoning 30, 34 P. micropoides, female, Paratype from Antu county, Jilin 31, 35 P. bellatrix, a female from Fusong county, Jilin. Scale bar: 0.5 mm. Abbreviations: g 1: gonocoxite I; g 2: gonocoxite II; es: ensiform setae; ns, nematiform setae; bc: bursa copulatrix; co: common oviduct; sc: seminal canal; rc: receptaculum; spc: spermathecal canal." figureDoi="10.3897/zookeys.1063.69942.figures28-35" httpUri="https://binary.pensoft.net/fig/599861" pageId="0" pageNumber="1">32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0714078130CA16CC7237F28EF6852CF2" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="1A25EE6A6519FA22DD233883976F1C60" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="80F1C9667EB9759215B8414DEA5C11B7" pageId="0" pageNumber="1">
|
||
BL 19.7-21.2 mm, BW 6.3-6.6 mm. Body form robust, dorsally dark brown, nearly black, elytra opaque, without metallic luster. Head large, widest at temporae; frons smooth; frontal grooves shallow; temporae strongly swollen, a little shorter than eyes; eyes small and hemispherical; terminal segment of labial palpus fusiform. Pronotum strongly cordate, disc evenly and densely covered with fine punctures, area anterior to sub-anterior transversal sulci well punctate; PW/HW = 1.23-1.27, PW/PL = 1.53-1.56, widest near anterior third; anterior margin a little wider than posterior margin, PAW/PBW = 1.16-1.18. Lateral margins slightly arched from anterior angles to the middle, strongly sinuate and then nearly straight before posterior angles; posterior angles rectangular; mid-lateral setae present at anterior fifth of lateral margins; lateral expansions equal width anteriorly and posteriorly. Basal foveae with inner and outer grooves faintly defined and partly fused, forming deep depression between them, outer groove slightly shorter than inner one; middle area between two basal foveae longitudinal rugose. Elytra oblong, shoulders widely rounded; basal ridge and lateral margin forming an obtuse angle; elytra 1.53-1.56 times longer than wide. Usually three discal pores present on third interval, all adjoining the second stria; the first one before middle, position of the second one variable, the last one at apical sixth to eighth. Umbilicate series on ninth interval continuous, sparse at middle. Ventral side: metepisternum nearly smooth; sternite VII of males without secondary sexual modification. Fifth tarsomere without ventral seta. Male genitalia: median lobe of aedeagus stout, gradually curved at basal third, apical portion gradually deflected ventrally; ventral surface strongly tumid near middle, forming a conspicuous cuneate tubercle a little anterior to the middle (Fig.
|
||
<figureCitation id="D81D5AA7320FBD6B9B91F9DF4C075240" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Male genitalia of Pterostichus (Koreonialoe) spp. from China, left lateral view of median lobe of aedeagus 7 P. micropoides sp. nov., Holotype 8 P. micropoides sp. nov., Paratype from Changbai county, Jilin 9 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 10 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 11 P. quadrilobatus sp. nov., Holotype 12 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures7-12" httpUri="https://binary.pensoft.net/fig/599857" pageId="0" pageNumber="1">12</figureCitation>
|
||
); apical lamella short and wide, apex rounded, not oblique to the left in dorsal view (Fig.
|
||
<figureCitation id="AC1DFCFCA526737579D2E19A46596768" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus (Koreonialoe) spp. from China, dorsal view of median lobe of aedeagus 13 P. micropoides sp. nov., Holotype 14 P. micropoides sp. nov. Paratype from Changbai county 15 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 16 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 17 P. quadrilobatus sp. nov., Paratype from Laopingding, Liaoning 18 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures13-18" httpUri="https://binary.pensoft.net/fig/599858" pageId="0" pageNumber="1">18</figureCitation>
|
||
). Endophallus (Fig.
|
||
<figureCitation id="52F6EE7722734FDF553322CE44CD3DFB" captionStart="Figures 22–27" captionStartId="F5" captionText="Figures 22 - 27. Endophallus of Pterostichus (Koreonialoe) spp. from China, left lateral view and right lateral view 22 P. micropoides sp. nov., Paratype from Antu city, Jilin 23 P. micropoides sp. nov., Paratype from Changbaishan mt., Jilin 24 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 25 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 26 P. quadrilobatus sp. nov., Holotype 27 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures22-27" httpUri="https://binary.pensoft.net/fig/599860" pageId="0" pageNumber="1">27</figureCitation>
|
||
) long, directed basal-ventrally, gonopore opened to the ventral-basal direction of aedeagus; three endophallic lobes recognized, all placed near gonopore: left apical lobe (
|
||
<emphasis id="E0B1FEFB04FD86153D4729D2338398CA" bold="true" pageId="0" pageNumber="1">la</emphasis>
|
||
) oblate, apex slightly hooked, forming an oblique upper surface; left preapical lobe (
|
||
<emphasis id="C153CAF74FFC50B7CE1F166E8084ACAC" bold="true" pageId="0" pageNumber="1">lp</emphasis>
|
||
) similar size as la, apex rounded, well separate from la; right apical lobe (
|
||
<emphasis id="FED3470E04A0EC29476B1665E153D47B" bold="true" pageId="0" pageNumber="1">ra</emphasis>
|
||
) faintly defined, nearly spherical. Female genitalia typical in this subgenus.
|
||
</paragraph>
|
||
<caption id="C38CCDCA1A9A1CDB4C2157DA4B22ADA3" doi="10.3897/zookeys.1063.69942.figures22-27" httpUri="https://binary.pensoft.net/fig/599860" pageId="0" pageNumber="1" start="Figures 22–27" startId="F5">
|
||
<paragraph id="C0E377AC41BFE7A7F987F2438C65B01C" pageId="0" pageNumber="1">
|
||
<emphasis id="58F6C869DAFFCE5CAD5FB806E67BF020" bold="true" pageId="0" pageNumber="1">Figures 22-27.</emphasis>
|
||
Endophallus of
|
||
<taxonomicName id="146BC79D0563561CFC537203009572CF" authority="" authorityName="Park & Kwon" authorityYear="1996" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Koreonialoe)" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Koreonialoe">Pterostichus (Koreonialoe)</taxonomicName>
|
||
spp. from China, left lateral view and right lateral view
|
||
<emphasis id="09D2287C2076658EEBA6DFA8F53BC3B6" bold="true" pageId="0" pageNumber="1">22</emphasis>
|
||
<taxonomicName id="889FED247860CE6B0E996AA0C7CF2FD2" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. micropoides" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="micropoides">
|
||
<emphasis id="0F473B4FC94108D0D782CDDF00A352FA" italics="true" pageId="0" pageNumber="1">P. micropoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., Paratype from Antu city, Jilin
|
||
<emphasis id="B0527D9F3B5BCAEF8E43AC0548F91DBE" bold="true" pageId="0" pageNumber="1">23</emphasis>
|
||
<taxonomicName id="33319E911CF7F39DD7D26EFBAA44D785" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. micropoides" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="micropoides">
|
||
<emphasis id="03FD1FF5EEBB8550794335F63111C5C8" italics="true" pageId="0" pageNumber="1">P. micropoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., Paratype from Changbaishan mt., Jilin
|
||
<emphasis id="B8E4C2F7778D8640E101538E7CFE2220" bold="true" pageId="0" pageNumber="1">24</emphasis>
|
||
<taxonomicName id="C4177E34CB224D2D722D746234B2712A" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="syleus">
|
||
<emphasis id="49DB18B9861E1D57EE741E7B1D732DE6" italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
|
||
</taxonomicName>
|
||
Kirschenhofer, a male from Fengcheng city, Liaoning
|
||
<emphasis id="76BBF64F73B6B7F2184611D3367E53EA" bold="true" pageId="0" pageNumber="1">25</emphasis>
|
||
<taxonomicName id="EED3AD8E08026E2223B793544D7F4010" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="syleus">
|
||
<emphasis id="EFA7C727FCA2B01C44E1AAFB6DA6CC67" italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
|
||
</taxonomicName>
|
||
Kirschenhofer, a male from Shedao Island, Liaoning
|
||
<emphasis id="E04341DCB90928785D6D55D1509C3EA9" bold="true" pageId="0" pageNumber="1">26</emphasis>
|
||
<taxonomicName id="E18196996BE0546F8851A4B2F8AB664E" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. quadrilobatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="quadrilobatus">
|
||
<emphasis id="6FC3B498BF7A4FF3A9401830D2FB9C80" italics="true" pageId="0" pageNumber="1">P. quadrilobatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., Holotype
|
||
<emphasis id="59B63D3F9D3430680C4D134176DA39B7" bold="true" pageId="0" pageNumber="1">27</emphasis>
|
||
<taxonomicName id="B5EC5BF9D144D2E155FD8E69D01CEF4B" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellatrix" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellatrix">
|
||
<emphasis id="7AFEC7E0798FBB6A2E969B23B1157605" italics="true" pageId="0" pageNumber="1">P. bellatrix</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="EBE3D2CE65A8C88A3D9402E18E15F0C9" originalValue="Tschitschérine">Tschitscherine</normalizedToken>
|
||
, a male from Changbai county, Jilin. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5396D9317E929B28750B9CB9F556FCE4" doi="10.3897/zookeys.1063.69942.figures28-35" httpUri="https://binary.pensoft.net/fig/599861" pageId="0" pageNumber="1" start="Figures 28–35" startId="F6">
|
||
<paragraph id="8D63BC35CB2820DCA5B857DC7FA2EC1D" pageId="0" pageNumber="1">
|
||
<emphasis id="26AEDBE0CFBCC5129BECE570B3DEB9EA" bold="true" pageId="0" pageNumber="1">Figures 28-35.</emphasis>
|
||
Female genitalia of
|
||
<taxonomicName id="6765D6C5DF30D64C658B77EE9C21185D" authority="" authorityName="Park & Kwon" authorityYear="1996" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Koreonialoe)" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Koreonialoe">Pterostichus (Koreonialoe)</taxonomicName>
|
||
spp. from China, Figs
|
||
<emphasis id="2D69351860633D98F53632ACE9E7A6A4" bold="true" pageId="0" pageNumber="1">28-31</emphasis>
|
||
ventral view and inner lateral view of ovipositor. Figs
|
||
<emphasis id="84BA3FBE4718D6575DF7D1EC70A78417" bold="true" pageId="0" pageNumber="1">32-35</emphasis>
|
||
female reproductive system.
|
||
<emphasis id="48691FA2EBCFC953F4DFF6D4114C5EE6" bold="true" pageId="0" pageNumber="1">28, 32</emphasis>
|
||
<taxonomicName id="73FC51B7A0FDBE7980DFC571FAB64CD5" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tetralobatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tetralobatus">
|
||
<emphasis id="B9D6AF3887FB7B3547116E42D00363ED" italics="true" pageId="0" pageNumber="1">P. tetralobatus</emphasis>
|
||
</taxonomicName>
|
||
, Paratype from Laopingding, Liaoning
|
||
<emphasis id="BA30A8A3099C1F4C6A319DEA579293F0" bold="true" pageId="0" pageNumber="1">29, 33</emphasis>
|
||
<taxonomicName id="0DB65DD5A734D09A389E52F5474867C1" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="syleus">
|
||
<emphasis id="6C8E84E4E9D62AE356EDF53445C14AEF" italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
|
||
</taxonomicName>
|
||
, a female from Baishilazi, Liaoning
|
||
<emphasis id="1BD9622440EBE5C5525FEEA1260EB437" bold="true" pageId="0" pageNumber="1">30, 34</emphasis>
|
||
<taxonomicName id="25BECFAC2F752C348DB865B23F1BE44C" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. micropoides" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="micropoides">
|
||
<emphasis id="6F78A05E02B48DF8D7F3C3015D5EB044" italics="true" pageId="0" pageNumber="1">P. micropoides</emphasis>
|
||
</taxonomicName>
|
||
, female, Paratype from Antu county, Jilin
|
||
<emphasis id="9107C9F0D7F0750440C1FA2826B165F1" bold="true" pageId="0" pageNumber="1">31, 35</emphasis>
|
||
<taxonomicName id="82B00690C34104F0AC951C9CD6FE06AF" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellatrix" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellatrix">
|
||
<emphasis id="24B9CBCB7A033563587B16FC0604B4AE" italics="true" pageId="0" pageNumber="1">P. bellatrix</emphasis>
|
||
</taxonomicName>
|
||
, a female from Fusong county, Jilin. Scale bar: 0.5 mm. Abbreviations: g1: gonocoxite I; g2:gonocoxite II; es: ensiform setae; ns, nematiform setae; bc: bursa copulatrix; co: common oviduct; sc: seminal canal; rc: receptaculum; spc: spermathecal canal.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="EB86A48469F64B85991B33429AF5C795" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="DD17774CD0009ECF2B65B70CC3EF01A4" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="0235C5FF4D1F40AA9BF59FFE199E65FC" pageId="0" pageNumber="1">
|
||
This species is widespread from the southeastern part of Jilin province to South Korea, along the eastern mountains in the Korean Peninsula. (Fig.
|
||
<figureCitation id="7B9A5DCB39E989A9BA105F86B23D18C0" captionStart="Figure 36" captionStartId="F7" captionText="Figure 36. Confirmed distribution of Pterostichus (Koreonialoe) spp. in China: Green: P. micropoides sp. nov.; Red: P. tetralobatus sp. nov.; Yellow: P. bellatrix Tschitscherine; Blue: P. syleus Kirschenhofer." figureDoi="10.3897/zookeys.1063.69942.figure36" httpUri="https://binary.pensoft.net/fig/599862" pageId="0" pageNumber="1">36</figureCitation>
|
||
, yellow, distributions in South Korea not shown)
|
||
</paragraph>
|
||
<caption id="F0EDB64F9FAA18D59CB73426101F113D" doi="10.3897/zookeys.1063.69942.figure36" httpUri="https://binary.pensoft.net/fig/599862" pageId="0" pageNumber="1" start="Figure 36" startId="F7">
|
||
<paragraph id="CB79624AA2C24C6C690554C50BFA0B03" pageId="0" pageNumber="1">
|
||
<emphasis id="56576EE08A40422059975932EF06A11E" bold="true" pageId="0" pageNumber="1">Figure 36.</emphasis>
|
||
Confirmed distribution of
|
||
<taxonomicName id="533B426C39B50685FDDBCB037312A5CF" authority="" authorityName="Park & Kwon" authorityYear="1996" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Koreonialoe)" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Koreonialoe">Pterostichus (Koreonialoe)</taxonomicName>
|
||
spp. in China: Green:
|
||
<taxonomicName id="C03419FB7AD677EE70652C3BF132ADBB" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. micropoides" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="micropoides">
|
||
<emphasis id="C8175F2280799A1EAE6783E2E9CC1F35" italics="true" pageId="0" pageNumber="1">P. micropoides</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; Red:
|
||
<taxonomicName id="A1A7556B3D6C6A72B73FAE0803C3C5D5" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tetralobatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tetralobatus">
|
||
<emphasis id="EF3752F24392E22111DFF16B391BF42F" italics="true" pageId="0" pageNumber="1">P. tetralobatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; Yellow:
|
||
<taxonomicName id="5A67EBEE5B9035998BB04B24D3F86433" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. bellatrix" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bellatrix">
|
||
<emphasis id="F2CCC4421C1DB261E9A91A8E2D65258E" italics="true" pageId="0" pageNumber="1">P. bellatrix</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="3DAA37AAABFF846D22A9F88B94ED0D9F" originalValue="Tschitschérine">Tschitscherine</normalizedToken>
|
||
; Blue:
|
||
<taxonomicName id="9FBF5249CE0DCF32D40224838948671B" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="syleus">
|
||
<emphasis id="B219955FED4420FC835AC3E3FA63FA22" italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
|
||
</taxonomicName>
|
||
Kirschenhofer.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |