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<document id="01CB873AA2855DA60C24605B4799AC3F" ID-DOI="10.15298/rusentj.28.3.05" ID-ISSN="0132-8069" ID-Zenodo-Dep="13163961" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722579353071" checkinUser="felipe" docAuthor="Lyubarsky, G. Yu. &amp; Perkovsky, E. E." docDate="2019" docId="654DF55DFFD5832993DBB0D3FE3C0DD3" docLanguage="en" docName="RussEntomolJ.28.3.263-268.pdf" docOrigin="Russian Entomological Journal 28 (3)" docSource="http://dx.doi.org/10.15298/rusentj.28.3.05" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.2:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="2" docTitle="Spaniophagus Lyubarsky et Perkovsky 2019, gen.n." docType="treatment" docVersion="2" lastPageNumber="265" masterDocId="99748D25FFD7832B931AB10AFFF30B08" masterDocTitle="Spaniophagus, first new Eocene genus of silken fungus beetle from Baltic amber (Coleoptera: Clavicornia: Cryptophagidae)" masterLastPageNumber="268" masterPageNumber="263" pageNumber="265" updateTime="1722580610524" updateUser="ExternalLinkService">
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<mods:title id="B416A66B83BB656237AD062B0BD82A95">Spaniophagus, first new Eocene genus of silken fungus beetle from Baltic amber (Coleoptera: Clavicornia: Cryptophagidae)</mods:title>
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<mods:namePart id="4144596DDA4A7687E1F183BC8C732000">Perkovsky, E. E.</mods:namePart>
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<treatment id="654DF55DFFD5832993DBB0D3FE3C0DD3" LSID="urn:lsid:plazi:treatment:654DF55DFFD5832993DBB0D3FE3C0DD3" httpUri="http://treatment.plazi.org/id/654DF55DFFD5832993DBB0D3FE3C0DD3" lastPageNumber="265" pageId="2" pageNumber="265">
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<heading id="B613F327FFD5832993DBB0D3FCF10AF9" box="[193,770,473,497]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="265" reason="2">
Genus
<taxonomicName id="2AE43FC8FFD583299216B0D3FD400AF9" authority="Lyubarsky et Perkovsky" authorityName="Lyubarsky et Perkovsky" authorityYear="2019" box="[268,691,473,497]" class="Insecta" family="Cryptophagidae" genus="Spaniophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="265" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="DF909859FFD583299216B0D3FE500AF9" box="[268,419,473,497]" italics="true" pageId="2" pageNumber="265">Spaniophagus</emphasis>
Lyubarsky et Perkovsky
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,
<emphasis id="DF909859FFD5832991A4B0D3FCF10AF9" bold="true" box="[702,770,473,497]" pageId="2" pageNumber="265">
<taxonomicNameLabel id="C4A32522FFD5832991A4B0D3FCF10AF9" box="[702,770,473,497]" pageId="2" pageNumber="265" rank="genus">gen.n.</taxonomicNameLabel>
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<paragraph id="ED5B444BFFD5832992B5B0FCFDE70906" blockId="2.[189,776,399,1756]" box="[431,532,502,526]" pageId="2" pageNumber="265">
<figureCitation id="75DF58CEFFD5832992B5B0FCFDE30906" box="[431,528,502,526]" captionStart-0="Figs 14" captionStart-1="Figs 56" captionStart-2="Fig" captionStartId-0="1.[226,263,1969,1988]" captionStartId-1="3.[226,263,1969,1988]" captionStartId-2="4.[226,257,1682,1701]" captionTargetBox-0="[189,1399,378,1932]" captionTargetBox-1="[274,1399,435,1947]" captionTargetBox-2="[220,748,314,1649]" captionTargetId-0="figure-80@1.[189,1399,378,1932]" captionTargetId-1="figure-71@3.[189,1399,435,1947]" captionTargetId-2="figure-554@4.[220,748,314,1649]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figs 14. Spaniophagus hoffeinsae gen. et sp.n. Holotype (inv. No CCHH 824-4 from the collection of the Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 1 — dorsal; 2 — ventral; 3 — frontal; 4 — lateral. Рис. 14. Spaniophagus hoffeinsae gen. et sp.n. Голотип (inv. No CCHH 824-4 в коллекции Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 1 — сверху; 2 — сниЗу; 3 — спереди; 4 — сбоку." captionText-1="Figs 56. Spaniophagus hoffeinsae gen. et sp.n. Holotype (inv. No CCHH 824-4 from the collection of the Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 5 — lateral margin of pronotum; 6 — legs. Рис. 56. Spaniophagus hoffeinsae gen. et sp.n. Голотип (inv. No CCHH 824-4 в коллекции Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 5 — боковой край переднеспинки; 6 — ноги." captionText-2="Fig. 7. Drawing of ventral view, Spaniophagus hoffeinsae gen. et sp.n. Рис. 7. Рисунок вентральной стороны Spaniophagus hoffeinsae gen. et sp.n." figureDoi-0="http://doi.org/10.5281/zenodo.13163963" figureDoi-1="http://doi.org/10.5281/zenodo.13163965" figureDoi-2="http://doi.org/10.5281/zenodo.13163967" httpUri-0="https://zenodo.org/record/13163963/files/figure.png" httpUri-1="https://zenodo.org/record/13163965/files/figure.png" httpUri-2="https://zenodo.org/record/13163967/files/figure.png" pageId="2" pageNumber="265">Figs 17</figureCitation>
.
</paragraph>
<paragraph id="ED5B444BFFD5832993F8B316FF1B094E" blockId="2.[189,776,399,1756]" pageId="2" pageNumber="265">
<typeStatus id="325FFAE9FFD5832993F8B316FEFD0927" box="[226,270,540,559]" pageId="2" pageNumber="265">Type</typeStatus>
species
<taxonomicName id="2AE43FC8FFD58329924DB311FCF50926" authority="Lyubarsky et Perkovsky" authorityName="Lyubarsky et Perkovsky" authorityYear="2019" box="[343,774,539,559]" class="Insecta" family="Cryptophagidae" genus="Spaniophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="265" phylum="Arthropoda" rank="species" species="hoffeinsae" status="sp. nov.">
<emphasis id="DF909859FFD58329924DB311FDC30926" box="[343,560,539,558]" italics="true" pageId="2" pageNumber="265">Spaniophagus hoffeinsae</emphasis>
Lyubarsky et Perkovsky
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<emphasis id="DF909859FFD5832993A7B339FF1B094E" bold="true" box="[189,232,563,582]" pageId="2" pageNumber="265">
<taxonomicNameLabel id="C4A32522FFD5832993A7B339FF1B094E" box="[189,232,563,582]" pageId="2" pageNumber="265" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A5FE17C0FFD5832993F8B341FEBB0814" pageId="2" pageNumber="265" type="diagnosis">
<paragraph id="ED5B444BFFD5832993F8B341FEBB0814" blockId="2.[189,776,399,1756]" pageId="2" pageNumber="265">
DIAGNOSIS.Body shape parallel-sided, moderately convex. Clypeus of male not notched. Antenna clubbed. Pronotum with anterior margin not emarginated; lateral margin not modified, not serrate, sinuate, or explanate; angularity absent; prominent middle tooth absent; sublateral line absent. Tibia slender, not club-shaped. Prosternal process not vault- ed. Ventrite 1 longer than remaining ventrites. Tarsal formula 5-5-
<quantity id="2A1CE9AEFFD5832993F9B20CFEF80814" box="[227,267,774,796]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="265" unit="in" value="4.0">4 in</quantity>
male.
</paragraph>
</subSubSection>
<subSubSection id="A5FE17C0FFD5832993F8B22BFE950D84" pageId="2" pageNumber="265" type="description">
<paragraph id="ED5B444BFFD5832993F8B22BFEF10FCE" blockId="2.[189,776,399,1756]" pageId="2" pageNumber="265">
DESCRIPTION. Color of body dark brown. Body shape parallel-sided, moderately convex (
<figureCitation id="75DF58CEFFD58329913FB231FD900858" box="[549,611,827,849]" captionStart="Figs 14" captionStartId="1.[226,263,1969,1988]" captionTargetBox="[189,1399,378,1932]" captionTargetId="figure-80@1.[189,1399,378,1932]" captionTargetPageId="1" captionText="Figs 14. Spaniophagus hoffeinsae gen. et sp.n. Holotype (inv. No CCHH 824-4 from the collection of the Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 1 — dorsal; 2 — ventral; 3 — frontal; 4 — lateral. Рис. 14. Spaniophagus hoffeinsae gen. et sp.n. Голотип (inv. No CCHH 824-4 в коллекции Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 1 — сверху; 2 — сниЗу; 3 — спереди; 4 — сбоку." figureDoi="http://doi.org/10.5281/zenodo.13163963" httpUri="https://zenodo.org/record/13163963/files/figure.png" pageId="2" pageNumber="265">Fig. 1</figureCitation>
). Punctation of body and elytron confused; punctation dense, distance between puncture on average less than diameter of puncture. Fronto-clypeal suture absent; subgenal spine well-developed; gular sutures present. Antenna clubbed; antennomeres not compact; last antennomere symmetrical and equal in length to penultimate (
<figureCitation id="75DF58CEFFD58329924FB2D1FE6308F8" box="[341,400,987,1009]" captionStart="Figs 14" captionStartId="1.[226,263,1969,1988]" captionTargetBox="[189,1399,378,1932]" captionTargetId="figure-80@1.[189,1399,378,1932]" captionTargetPageId="1" captionText="Figs 14. Spaniophagus hoffeinsae gen. et sp.n. Holotype (inv. No CCHH 824-4 from the collection of the Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 1 — dorsal; 2 — ventral; 3 — frontal; 4 — lateral. Рис. 14. Spaniophagus hoffeinsae gen. et sp.n. Голотип (inv. No CCHH 824-4 в коллекции Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 1 — сверху; 2 — сниЗу; 3 — спереди; 4 — сбоку." figureDoi="http://doi.org/10.5281/zenodo.13163963" httpUri="https://zenodo.org/record/13163963/files/figure.png" pageId="2" pageNumber="265">Fig. 2</figureCitation>
). Width of labial palpomere 1 equal to palpomere 2. Mentum with transverse ridge. Eye well developed, prominent. Boss on front of head absent (
<figureCitation id="75DF58CEFFD5832991A3B51AFD0A0F2E" box="[697,761,1040,1062]" captionStart="Figs 14" captionStartId="1.[226,263,1969,1988]" captionTargetBox="[189,1399,378,1932]" captionTargetId="figure-80@1.[189,1399,378,1932]" captionTargetPageId="1" captionText="Figs 14. Spaniophagus hoffeinsae gen. et sp.n. Holotype (inv. No CCHH 824-4 from the collection of the Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 1 — dorsal; 2 — ventral; 3 — frontal; 4 — lateral. Рис. 14. Spaniophagus hoffeinsae gen. et sp.n. Голотип (inv. No CCHH 824-4 в коллекции Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 1 — сверху; 2 — сниЗу; 3 — спереди; 4 — сбоку." figureDoi="http://doi.org/10.5281/zenodo.13163963" httpUri="https://zenodo.org/record/13163963/files/figure.png" pageId="2" pageNumber="265">Fig. 3</figureCitation>
). Pronotum parallel-sided, its width does not decrease from the middle to the base; anterior margin of pronotum not emarginate; sides not explanate; margin unmodified (
<figureCitation id="75DF58CEFFD583299170B56AFD570F7E" box="[618,676,1120,1142]" captionStart="Figs 56" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[274,1399,435,1947]" captionTargetId="figure-71@3.[189,1399,435,1947]" captionTargetPageId="3" captionText="Figs 56. Spaniophagus hoffeinsae gen. et sp.n. Holotype (inv. No CCHH 824-4 from the collection of the Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 5 — lateral margin of pronotum; 6 — legs. Рис. 56. Spaniophagus hoffeinsae gen. et sp.n. Голотип (inv. No CCHH 824-4 в коллекции Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 5 — боковой край переднеспинки; 6 — ноги." figureDoi="http://doi.org/10.5281/zenodo.13163965" httpUri="https://zenodo.org/record/13163965/files/figure.png" pageId="2" pageNumber="265">Fig. 5</figureCitation>
); angularity absent; prominent middle tooth absent; sublateral line absent; basal pits absent; median fold absent; basal groove absent.
</paragraph>
<paragraph id="ED5B444BFFD5832993F8B5C6FEF10ED9" blockId="2.[189,776,399,1756]" pageId="2" pageNumber="265">
Procoxae wide open posteriorly, separated by prosternal process. Prosternal process not vaulted; longer than anterior portion of prosternum. Mesosternum without parallel lines. Mesocoxae set very closely, width of mesosternal process equal to or 2/3 that of mesocoxa. Epimeron incomplete, extending to metasternum. Metasternal process subdepressed, not beyond mesocoxae. Distance between metacoxae approximately equal to diameter of metacoxa. Metasternum with median longitudinal line (
<figureCitation id="75DF58CEFFD5832992E3B4AAFDCA0EBE" box="[505,569,1440,1462]" captionStart="Fig" captionStartId="4.[226,257,1682,1701]" captionTargetBox="[220,748,314,1649]" captionTargetId="figure-554@4.[220,748,314,1649]" captionTargetPageId="4" captionText="Fig. 7. Drawing of ventral view, Spaniophagus hoffeinsae gen. et sp.n. Рис. 7. Рисунок вентральной стороны Spaniophagus hoffeinsae gen. et sp.n." figureDoi="http://doi.org/10.5281/zenodo.13163967" httpUri="https://zenodo.org/record/13163967/files/figure.png" pageId="2" pageNumber="265">Fig. 7</figureCitation>
). Metasubcoxal line absent.
</paragraph>
<paragraph id="ED5B444BFFD5832993F8B4DCFE300D29" blockId="2.[189,776,399,1756]" pageId="2" pageNumber="265">Ventrite 1 longer than remaining ventrites, ventrite 5 without thickened setae. Tibia with apical fringle of spines. Tarsi slightly lobed below.</paragraph>
<paragraph id="ED5B444BFFD5832993F8B72CFE950D84" blockId="2.[189,776,399,1756]" pageId="2" pageNumber="265">
Male. Clypeus not notched. Antennal segments not compact. Ventrite 5 unmodified. Metatibia unmodified. Tarsal formula 554 (
<figureCitation id="75DF58CEFFD583299251B751FE750D79" box="[331,390,1627,1649]" captionStart="Figs 56" captionStartId="3.[226,263,1969,1988]" captionTargetBox="[274,1399,435,1947]" captionTargetId="figure-71@3.[189,1399,435,1947]" captionTargetPageId="3" captionText="Figs 56. Spaniophagus hoffeinsae gen. et sp.n. Holotype (inv. No CCHH 824-4 from the collection of the Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 5 — lateral margin of pronotum; 6 — legs. Рис. 56. Spaniophagus hoffeinsae gen. et sp.n. Голотип (inv. No CCHH 824-4 в коллекции Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany (SDEI)): 5 — боковой край переднеспинки; 6 — ноги." figureDoi="http://doi.org/10.5281/zenodo.13163965" httpUri="https://zenodo.org/record/13163965/files/figure.png" pageId="2" pageNumber="265">Fig 6</figureCitation>
); tarsomeres 13 of pro-, meso- and metatarsi dilated.
</paragraph>
</subSubSection>
<subSubSection id="A5FE17C0FFD5832993F8B79BFE3C0DD3" pageId="2" pageNumber="265" type="etymology">
<paragraph id="ED5B444BFFD5832993F8B79BFE3C0DD3" blockId="2.[189,776,399,1756]" pageId="2" pageNumber="265">
ETYMOLOGY. As the new genus is similar to the genera
<taxonomicName id="2AE43FC8FFD5832993A7B7A1FEA30DC8" authorityName="Reitter" authorityYear="1875" box="[189,336,1707,1728]" class="Insecta" family="Cryptophagidae" genus="Spaniophaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="DF909859FFD5832993A7B7A1FEA30DC8" box="[189,336,1707,1728]" italics="true" pageId="2" pageNumber="265">Spaniophaenus</emphasis>
</taxonomicName>
and
<taxonomicName id="2AE43FC8FFD58329929FB7A1FDFD0DC8" authorityName="Herbst" authorityYear="1792" box="[389,526,1707,1728]" class="Insecta" family="Cryptophagidae" genus="Cryptophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="DF909859FFD58329929FB7A1FDFD0DC8" box="[389,526,1707,1728]" italics="true" pageId="2" pageNumber="265">Cryptophagus</emphasis>
</taxonomicName>
, the name is a combination of these generic names.
</paragraph>
</subSubSection>
</treatment>
</document>