103 lines
13 KiB
XML
103 lines
13 KiB
XML
<document id="405E4CA9E0E3B98D51A454A867F9CDE7" ID-CLB-Dataset="33408" ID-DOI="http://dx.doi.org/10.3897/zookeys.149.2345" ID-GBIF-Dataset="ce66510c-5f50-4191-9097-7166355bf4a8" ID-PMC="PMC3234406" ID-Pensoft-Pub="1313-2970-149-17" ID-PubMed="22207791" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-149-17" ModsDocOrigin="ZooKeys 149" ModsDocTitle="Two new species of the Hagnagora anicata complex (Geometridae, Larentiinae) from Costa Rica" checkinTime="1451249591128" checkinUser="pensoft" docAuthor="Sullivan, J. Bolling" docDate="2011" docId="6542405E9980D8CB744F1261E31660E8" docLanguage="en" docName="ZooKeys 149: 17-29" docOrigin="ZooKeys 149" docSource="http://dx.doi.org/10.3897/zookeys.149.2345" docTitle="Hagnagora anicata Felder & Rogenhofer 1875" docType="treatment" docVersion="5" lastPageNumber="19" masterDocId="EF132A7D27270911FFBF8A6E7A34FF9B" masterDocTitle="Two new species of the Hagnagora anicata complex (Geometridae, Larentiinae) from Costa Rica" masterLastPageNumber="29" masterPageNumber="17" pageNumber="18" updateTime="1732795132519" updateUser="ExternalLinkService">
|
||
<mods:mods id="A0250300B13361FF208D0271DEA11929" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="FC69C8A54C09C085D77AC17CFF6A4265">
|
||
<mods:title id="93AD8EAD1DD0A5EF42D2465603265C73">Two new species of the Hagnagora anicata complex (Geometridae, Larentiinae) from Costa Rica</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="D7D2126101C12BEB400DFE01E68C45CB" type="personal">
|
||
<mods:role id="1D86AD71A743CB37AB2F71B73B5B39A3">
|
||
<mods:roleTerm id="F2F8DDA4945A25DF78A91E17218FC4D2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AA2CBCABD3BF1A249D29B4E561902298">Sullivan, J. Bolling</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="FF9F67CAB3C6EBE9453CBBC61A7492CF">text</mods:typeOfResource>
|
||
<mods:relatedItem id="6A46C66DD93B73BC86C79D9DF9BB3044" type="host">
|
||
<mods:titleInfo id="8DF935EF4FC74BD326AB8F7F7C2C2B26">
|
||
<mods:title id="7DD6601E2F98E4A1452E1D5949A710CE">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="F83EE947E64943D1C3EDCDC35820F756">
|
||
<mods:date id="8A68B1CE2F63F1C5007A0F9A2412F8BD">2011</mods:date>
|
||
<mods:detail id="95BA1EAB4ED342FF0EE88F50F71B668F" type="volume">
|
||
<mods:number id="854F3B3E2E42BBF1F7E12528E7A08445">149</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="1E6F8E5F9429C7359E15C8B1F69363E7" unit="page">
|
||
<mods:start id="BADD2B9DCA025722D715594272C3B59A">17</mods:start>
|
||
<mods:end id="682DE105AD085530F3E8964D935CF81C">29</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="5BA260F65347F2BD6DF255C20D747D64">
|
||
<mods:url id="E633EEC9250C0809ACF9E1E442BF23E9">http://dx.doi.org/10.3897/zookeys.149.2345</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="18B5645A88E3AF90E3626B3889E37A3C">journal article</mods:classification>
|
||
<mods:identifier id="945A5DBE06B93506CDEFA684FAF13FF8" type="DOI">http://dx.doi.org/10.3897/zookeys.149.2345</mods:identifier>
|
||
<mods:identifier id="97F6F7B014D140ACBC9501F61CFEDC44" type="Pensoft-Pub">1313-2970-149-17</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="6542405E9980D8CB744F1261E31660E8" ID-GBIF-Taxon="152032293" LSID="urn:lsid:plazi:treatment:6542405E9980D8CB744F1261E31660E8" httpUri="http://treatment.plazi.org/id/6542405E9980D8CB744F1261E31660E8" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18" scope_family="Geometridae" scope_subFamily="Larentiinae">
|
||
<subSubSection id="95C6E3C1AC6CB0E160299AB66763A2CB" pageId="1" pageNumber="18" type="nomenclature">
|
||
<paragraph id="FFDAC2ABB735EA44F68954482EFD7E8D" pageId="1" pageNumber="18">
|
||
<taxonomicName id="22AB99F743AC9A7A51B2014A391BB10F" ID-CoL="934NS" LSID="http://species-id.net/wiki/Hagnagora_anicata" authority="Felder & Rogenhofer, 1875" authorityName="Felder & Rogenhofer" authorityYear="1875" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata (Felder & Rogenhofer, 1875)</taxonomicName>
|
||
Fig. 1
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2EAE0151F2C7482E56195AA8F90622DE" pageId="1" pageNumber="18" type="reference_group">
|
||
<paragraph id="E97B4D4ACCCC8D019529948C82A5618F" pageId="1" pageNumber="18">
|
||
<taxonomicName id="59D07BC2A9F1833E9E22DC048645FDDB" class="Insecta" family="Geometridae" genus="Heterusia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterusia anicata" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="anicata">Heterusia anicata</taxonomicName>
|
||
Felder & Rogenhofer, 1875, pl.130, fig. 13.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1A34560079C6345352FEBD56138DAE77" pageId="1" pageNumber="18" type="type material">
|
||
<paragraph id="FF9A2039CC64A2CF7D34B342BB55CA27" pageId="1" pageNumber="18">Type material.</paragraph>
|
||
<paragraph id="69D92ECB72EA63A89982D828C719918E" pageId="1" pageNumber="18">
|
||
Bogota, [Colombia]. A male specimen in BMNH labeled as
|
||
<normalizedToken id="C909EE5E5E10AD9F1A162499983197AE" originalValue="‘type’">'type'</normalizedToken>
|
||
is here designated as the lectotype (Fig. 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DC15869373295FE2763470C27624D7D4" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18" type="description and diagnosis">
|
||
<paragraph id="C21B754D838A7B7B28DADD7A156C672C" pageId="1" pageNumber="18">Description and diagnosis.</paragraph>
|
||
<paragraph id="87B54F12841F1EB52074BF480720BE1C" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
|
||
The maculation of
|
||
<taxonomicName id="66ECBDE4A9F0DEB1590F5D2D0EEE02A4" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
|
||
is shown in Fig 1a. A cream white band originates on the costa near the midpoint and runs diagonally to
|
||
<pageBreakToken id="92425051517CC25BF5DA973E77FB483C" pageId="2" pageNumber="19" start="start">ward</pageBreakToken>
|
||
a point 1/3 above the tornus, rounds off and ends without touching the margin. Distal to this band is dark brown scaling to the wing apex. Proximal to the band is dark brown scaling which becomes brighter toward the wing base. At the terminus of the wing veins are small white crescents which are usually worn off in flown specimens. The hindwing is the brownish color of the forewing base and with larger marginal crescents at the vein termini. The underside (Fig. 1b) is repeated with some slight variation in most
|
||
<taxonomicName id="9FE43429AD65E0B2C6B0F443E98F8598" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">Hagnagora</taxonomicName>
|
||
species. This maculation pattern is seen over almost the entire range (Jamaica and Mexico to Bolivia and Venezuela), with minor modification of the width of the white band. All have been referred to as
|
||
<taxonomicName id="5AC34A46C65D55577E0F96EBFF75D752" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5EB0006B813BC8829C882796FB13EC57" pageId="2" pageNumber="19">
|
||
The male genitalia are similar to those of
|
||
<taxonomicName id="24649E36A04D073756A4FFAE222AB935" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora marionae" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="marionae">Hagnagora marionae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="6279F224F4D868DFFA3B2397A05DCF35" author="Brehm, G" journalOrPublisher="Zeitschrift" pageId="7" pageNumber="24" pagination="256 - 260" title="Unusual flight activity of a new species of Hagnagora Druce, 1885 (Lepidoptera: Geometridae) from Costa Rica. Ent." volume="115" year="2005">Brehm and Sullivan 2005</bibRefCitation>
|
||
) and several characters are important. The uncus tapers evenly ending in a slight hook. Large hair brushes on the inner face of the valva (Fig. 1b) often obscure details of the valva, as often seen in species of another larentiine genus,
|
||
<taxonomicName id="E8648BAF35D0FF2B983ED8360058F44C" class="Insecta" family="Geometridae" genus="Hydriomena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydriomena" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">Hydriomena</taxonomicName>
|
||
<normalizedToken id="8DF5E8FE5AA6B4B969313F35C6FBEB69" originalValue="Hübner">Huebner</normalizedToken>
|
||
. However, in
|
||
<taxonomicName id="584FEF7730C2CD54F7F9E10391BAD8E4" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
|
||
the costal edge of the valva is sclerotized and terminates in a well-defined point extending dorsad at the distal margin of the valva. The anal edge of the valva is swollen, stippled with small dots but not well sclerotized. The median area of the valve is unsclerotized and serves as the anchor point for the large hair brush. A smaller hair brush originates subterminally just below the costa. The juxta appears as a broad, slightly sclerotized plate and the anellus is broadened but not medially joined. The saccus is unsclerotized and u-shaped. The aedeagus is similar to that of
|
||
<taxonomicName id="64FD728749EF83A418C3B3BD50AA086E" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora marionae" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="marionae">Hagnagora marionae</taxonomicName>
|
||
and the vesica has two dorsal, granulated wing-like projections but no cornutus.
|
||
</paragraph>
|
||
<paragraph id="7891A9A980A47C5CC7262EA1242771D3" pageId="2" pageNumber="19">
|
||
Beside the lectotype there is a female from Bogota in the collections at the BMNH but its wing length is only 18 mm, i.e. smaller than that of the male lectotype. Because all known females of
|
||
<taxonomicName id="E95199D16CBDF6E90A7E9E44880424B5" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">Hagnagora</taxonomicName>
|
||
species are larger than their respective males, this female is not considered to be a female of
|
||
<taxonomicName id="215B169AEA5ECE7270031FF8F79F2082" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
|
||
. Because there are at least two other species occurring in Colombia with maculation extremely similar to
|
||
<taxonomicName id="5E4FFBD968E628B3784EADCAAE49525E" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
|
||
(Sullivan, unpubl. data), it would be best to obtain fresh male and female specimens from near Bogota for barcode sequencing to be sure of correctly associating a female as
|
||
<taxonomicName id="EDED2F42120ADA34AC55610F92631EDF" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="0CA2971F3E357CC91ECEB889602B149F" pageId="2" pageNumber="19">
|
||
<paragraph id="CB147325EB0975765F16EF206944A4A1" pageId="2" pageNumber="19">
|
||
Figure 1.
|
||
<taxonomicName id="9F96C44E29226696227FB02C7DEEFD85" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
|
||
(F. & R.), male lectotype a dorsal view b ventral view c genital capsule and d aedeagus (BMNH slide 22252).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |