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<document id="DC8E4C03573F781FAECA8E1AD2A7054A" ID-CLB-Dataset="158406" ID-DOI="10.11646/zootaxa.5194.2.2" ID-GBIF-Dataset="83757748-ba0b-403d-9d82-26e0907f716b" ID-ISSN="1175-5326" ID-Zenodo-Dep="7147068" ID-ZooBank="A89D71A5-8C75-434E-BD7D-8054ED97220C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664955361361" checkinUser="plazi" docAuthor="Zhu, Xiaoju &amp; Wang, Shuxia" docDate="2022" docId="652AE37CFFD5AC58FF76C821FE53F93B" docLanguage="en" docName="zootaxa.5194.2.2.pdf" docOrigin="Zootaxa 5194 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ethmia anomala Zhu &amp; Wang 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="178" masterDocId="99139B04FFD7AC5AFFE1C907FFC8FFDA" masterDocTitle="Study of the genus Ethmia Hübner, [1819] (Lepidoptera: Ethmiidae) from China with descriptions of six new species" masterLastPageNumber="192" masterPageNumber="176" pageNumber="178" updateTime="1699387694017" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BFE13027413D9386082EAA0141482341">Study of the genus Ethmia Hübner, [1819] (Lepidoptera: Ethmiidae) from China with descriptions of six new species</mods:title>
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<mods:namePart id="29199318468EAA0572A3B78F08079C55">Zhu, Xiaoju</mods:namePart>
<mods:nameIdentifier id="039A691F9886168EE3039E2641542ED0" type="ORCID">0000-0002-4590-8480</mods:nameIdentifier>
<mods:affiliation id="19971D4101323C4A162FFB34465CCF95">College of Life Sciences, Nankai University, Tianjin 300071, China. &amp; https: // orcid. org / 0000 - 0002 - 4590 - 8480</mods:affiliation>
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<mods:namePart id="B721395C4C90ED721AA99B44D7A302F2">Wang, Shuxia</mods:namePart>
<mods:affiliation id="CF51F4239AB48CAE2F0B1E2B04A26DF3">College of Life Sciences, Nankai University, Tianjin 300071, China.</mods:affiliation>
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<mods:title id="78A9C6046FF667DD457E594BD97017EE">Zootaxa</mods:title>
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<mods:part id="E525FA017CFE812F4E8BED9E63D5EAD7">
<mods:date id="FD2AD12960525FB2B964C6BB9FAF7E4E">2022</mods:date>
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<mods:number id="20425B926C54D5FA3B0A6BB4F491EF61">2022-10-05</mods:number>
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<mods:number id="9481ADB32E1B9EF3707604A91F59B724">5194</mods:number>
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<mods:identifier id="6070F21DAC7F4CC2A4EE189619BEC759" type="DOI">10.11646/zootaxa.5194.2.2</mods:identifier>
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<treatment id="652AE37CFFD5AC58FF76C821FE53F93B" ID-DOI="http://doi.org/10.5281/zenodo.7147084" ID-GBIF-Taxon="202142533" ID-Zenodo-Dep="7147084" LSID="urn:lsid:plazi:treatment:652AE37CFFD5AC58FF76C821FE53F93B" httpUri="http://treatment.plazi.org/id/652AE37CFFD5AC58FF76C821FE53F93B" lastPageNumber="178" pageId="2" pageNumber="178">
<subSubSection id="A59901E1FFD5AC58FF76C821FE0FFE9B" box="[151,455,294,321]" pageId="2" pageNumber="178" type="nomenclature">
<paragraph id="ED3C526AFFD5AC58FF76C821FE0FFE9B" blockId="2.[151,455,294,357]" box="[151,455,294,321]" pageId="2" pageNumber="178">
<heading id="B674E506FFD5AC58FF76C821FE0FFE9B" bold="true" box="[151,455,294,321]" fontSize="11" level="1" pageId="2" pageNumber="178" reason="1">
<emphasis id="DFF78E78FFD5AC58FF76C821FE0FFE9B" bold="true" box="[151,455,294,321]" pageId="2" pageNumber="178">
<taxonomicName id="2A8329E9FFD5AC58FF76C821FEAAFE9B" ID-CoL="9H5P4" authority="Zhu &amp; Wang, 2022" authorityName="Zhu &amp; Wang" authorityYear="2022" box="[151,354,294,321]" class="Insecta" family="Ethmiidae" genus="Ethmia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="178" phylum="Arthropoda" rank="species" species="anomala" status="sp. nov.">
<emphasis id="DFF78E78FFD5AC58FF76C821FEAAFE9B" bold="true" box="[151,354,294,321]" italics="true" pageId="2" pageNumber="178">Ethmia anomala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C4C43303FFD5AC58FE88C820FE0FFE9B" box="[361,455,295,321]" pageId="2" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="A59901E1FFD5AC58FF76C84CFED0FEBF" box="[151,280,331,357]" pageId="2" pageNumber="178" type="description">
<paragraph id="ED3C526AFFD5AC58FF76C84CFED0FEBF" blockId="2.[151,455,294,357]" box="[151,280,331,357]" pageId="2" pageNumber="178">
(
<figureCitation id="75B84EEFFFD5AC58FF7EC84CFF20FEBF" box="[159,232,331,357]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,957,981]" captionTargetBox="[165,1424,190,928]" captionTargetId="figure-17@4.[155,1431,181,933]" captionTargetPageId="4" captionText="FIGURES 1–4. Adults of Ethmia spp. 1, E. anatiformis, ♂; 2, E. anomala sp. nov., holotype, ♂; 3, E. corrugata sp. nov., holotype, ♂; 4, E. adunatimacula sp. nov., holotype, ♂. Scales = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7147070" httpUri="https://zenodo.org/record/7147070/files/figure.png" pageId="2" pageNumber="178">Figs 2</figureCitation>
,
<figureCitation id="75B84EEFFFD5AC58FF15C84CFEC7FEBF" box="[244,271,331,357]" captionStart="FIGURES 10–15" captionStartId="11.[151,264,1898,1922]" captionTargetBox="[232,1362,194,1860]" captionTargetId="figure-21@11.[206,1382,181,1874]" captionTargetPageId="11" captionText="FIGURES 10–15. Male genitalia of Ethmia spp. 10, E. anatiformis, slide No. ZXJ20107; 11, E. anomala sp. nov., holotype, slide No. ZXJ20076; 12, E. corrugata sp. nov., holotype, slide No. ZXJ20108; 13, E. adunatimacula sp. nov., holotype, slide No. ZXJ20202; 14, E. crocosoma, slide No. ZXJ20129; 15, E. rectiuga sp. nov., holotype, slide No. ZXJ19650. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7147076" httpUri="https://zenodo.org/record/7147076/files/figure.png" pageId="2" pageNumber="178">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A59901E1FFD5AC58FF76C894FDB0FE0B" pageId="2" pageNumber="178" type="materials_examined">
<paragraph id="ED3C526AFFD5AC58FF76C894FDB0FE0B" blockId="2.[151,1437,402,1761]" pageId="2" pageNumber="178">
<emphasis id="DFF78E78FFD5AC58FF76C894FE8EFE76" bold="true" box="[151,326,402,429]" pageId="2" pageNumber="178">Type material.</emphasis>
<materialsCitation id="5DEB5837FFD5AC58FEACC895FDB0FE0B" ID-GBIF-Occurrence="3920945336" accessionNumber="ZXJ20076" collectingDate="2016-08-02" collectorName="WX Feng &amp; WT Shi" country="China" elevation="772" latitude="32.03" location="Niutoudian" longLatPrecision="728" longitude="109.33" pageId="2" pageNumber="178" specimenCount="1" specimenCount-male="1" stateProvince="Shaanxi" typeStatus="holotype">
<emphasis id="DFF78E78FFD5AC58FEACC895FDEBFE76" bold="true" box="[333,547,402,428]" pageId="2" pageNumber="178">
<collectingCountry id="959412FAFFD5AC58FEACC895FE64FE76" box="[333,428,402,428]" name="China" pageId="2" pageNumber="178">CHINA</collectingCountry>
,
<collectingRegion id="2F479C88FFD5AC58FE5BC895FDD7FE76" box="[442,543,402,428]" country="China" name="Shaanxi" pageId="2" pageNumber="178">Shaanxi</collectingRegion>
:
</emphasis>
<typeStatus id="3238ECC8FFD5AC58FDCAC894FD5BFE77" box="[555,659,403,429]" pageId="2" pageNumber="178" type="holotype">Holotype</typeStatus>
<specimenCount id="FB8599E3FFD5AC58FD7AC894FD7AFE77" box="[667,690,403,429]" pageId="2" pageNumber="178" type="male">♂</specimenCount>
,
<location id="E85C04B1FFD5AC58FD5EC894FCF5FE77" LSID="urn:lsid:plazi:treatment:652AE37CFFD5AC58FF76C821FE53F93B:E85C04B1FFD5AC58FD5EC894FCF5FE77" box="[703,829,403,429]" country="China" latitude="32.03" longLatPrecision="728" longitude="109.33" name="Niutoudian" pageId="2" pageNumber="178" stateProvince="Shaanxi">Niutoudian</location>
(
<geoCoordinate id="88B734ADFFD5AC58FCACC894FC65FE77" box="[845,941,403,429]" degrees="32.03" direction="north" orientation="latitude" pageId="2" pageNumber="178" precision="555" value="32.03">32.03ºN</geoCoordinate>
,
<geoCoordinate id="88B734ADFFD5AC58FC58C894FBECFE77" box="[953,1060,403,429]" degrees="109.33" direction="east" orientation="longitude" pageId="2" pageNumber="178" precision="555" value="109.33">109.33ºE</geoCoordinate>
),
<location id="E85C04B1FFD5AC58FBD9C894FBB6FE77" LSID="urn:lsid:plazi:treatment:652AE37CFFD5AC58FF76C821FE53F93B:E85C04B1FFD5AC58FBD9C894FBB6FE77" box="[1080,1150,403,429]" country="China" latitude="32.03" longLatPrecision="728" longitude="109.33" name="Pingli" pageId="2" pageNumber="178" stateProvince="Shaanxi">Pingli</location>
,
<quantity id="2A7BFF8FFFD5AC58FB6BC894FB1BFE77" box="[1162,1235,403,429]" metricMagnitude="2" metricUnit="m" metricValue="7.72" pageId="2" pageNumber="178" unit="m" value="772.0">
<elevation id="66AEB559FFD5AC58FB6BC894FB1BFE77" box="[1162,1235,403,429]" metricMagnitude="2" metricUnit="m" metricValue="7.72" pageId="2" pageNumber="178" unit="m" value="772.0">772 m</elevation>
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,
<date id="993D74AAFFD5AC58FB01C894FAAFFE77" box="[1248,1383,403,429]" pageId="2" pageNumber="178" value="2016-08-02">
<collectingDate id="89798D42FFD5AC58FB01C894FAAFFE77" box="[1248,1383,403,429]" pageId="2" pageNumber="178" value="2016-08-02">2.VIII.2016</collectingDate>
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, leg.
<collectorName id="407637BCFFD5AC58FF76C8B0FECDFE0B" box="[151,261,439,465]" pageId="2" pageNumber="178">WX Feng</collectorName>
&amp;
<collectorName id="407637BCFFD5AC58FEC9C8B0FE4AFE0B" box="[296,386,439,465]" pageId="2" pageNumber="178">WT Shi</collectorName>
, slide No. ZXJ20076.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A59901E1FFD5AC58FF26C8DDFC78FD5F" pageId="2" pageNumber="178" type="diagnosis">
<paragraph id="ED3C526AFFD5AC58FF26C8DDFC78FD5F" blockId="2.[151,1437,402,1761]" pageId="2" pageNumber="178">
<emphasis id="DFF78E78FFD5AC58FF26C8DDFE8AFE2E" bold="true" box="[199,322,474,500]" pageId="2" pageNumber="178">Diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="2A8329E9FFD5AC58FD74C8DCFC7AFE2F" authority="Sattler, 1967" authorityName="Sattler" authorityYear="1967" box="[661,946,475,501]" class="Insecta" family="Ethmiidae" genus="Ethmia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="178" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="DFF78E78FFD5AC58FD74C8DCFCD4FE2F" box="[661,796,475,501]" italics="true" pageId="2" pageNumber="178">E. maculata</emphasis>
<bibRefCitation id="89122F9BFFD5AC58FCC5C8DCFC7AFE2F" author="Sattler, K." box="[804,946,475,501]" pageId="2" pageNumber="178" refId="ref11742" refString="Sattler, K. (1967) Ethmiidae. Microlepidoptera Palaearctica. Vol. 2. Verlag Georg Fromme &amp; Co., Wien, 185 pp." type="book" year="1967">
Sattler,
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</bibRefCitation>
</taxonomicName>
in forewing pattern. It can be distinguished by the black distal stripe of the forewing not connected with the distal stripe in the cell; by the uncus bifurcated in the distal 3/4, and the costa longer than half the length of the valva in the male genitalia. In
<taxonomicName id="2A8329E9FFD5AC58FB41CB24FAEFFDE7" authorityName="Sattler" authorityYear="1967" box="[1184,1319,547,573]" class="Insecta" family="Ethmiidae" genus="Ethmia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="178" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="DFF78E78FFD5AC58FB41CB24FAEFFDE7" box="[1184,1319,547,573]" italics="true" pageId="2" pageNumber="178">E. maculata</emphasis>
</taxonomicName>
, the black distal stripe of the forewing is connected with the distal stripe in the cell; the uncus is bifurcated in the distal 2/3, and the costa is half the length of the valva (
<bibRefCitation id="89122F9BFFD5AC58FD67CB6CFC82FD5F" author="Sattler, K." box="[646,842,619,645]" pageId="2" pageNumber="178" refId="ref11742" refString="Sattler, K. (1967) Ethmiidae. Microlepidoptera Palaearctica. Vol. 2. Verlag Georg Fromme &amp; Co., Wien, 185 pp." type="book" year="1967">Sattler 1967: 121</bibRefCitation>
, fig. 60).
</paragraph>
</subSubSection>
<subSubSection id="A59901E1FFD5AC58FF26CB89FE59F9AF" pageId="2" pageNumber="178" type="description">
<paragraph id="ED3C526AFFD5AC58FF26CB89FD21FD73" blockId="2.[151,1437,402,1761]" box="[199,745,654,681]" pageId="2" pageNumber="178">
<emphasis id="DFF78E78FFD5AC58FF26CB89FE91FD72" bold="true" box="[199,345,654,680]" pageId="2" pageNumber="178">Description.</emphasis>
Adult (
<figureCitation id="75B84EEFFFD5AC58FE4CCB88FE3BFD73" box="[429,499,655,681]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,957,981]" captionTargetBox="[165,1424,190,928]" captionTargetId="figure-17@4.[155,1431,181,933]" captionTargetPageId="4" captionText="FIGURES 1–4. Adults of Ethmia spp. 1, E. anatiformis, ♂; 2, E. anomala sp. nov., holotype, ♂; 3, E. corrugata sp. nov., holotype, ♂; 4, E. adunatimacula sp. nov., holotype, ♂. Scales = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7147070" httpUri="https://zenodo.org/record/7147070/files/figure.png" pageId="2" pageNumber="178">Fig. 2</figureCitation>
). Wingspan
<quantity id="2A7BFF8FFFD5AC58FD9FCB89FD2DFD73" box="[638,741,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="2" pageNumber="178" unit="mm" value="27.5">27.5 mm</quantity>
.
</paragraph>
<paragraph id="ED3C526AFFD5AC58FF26CBB4FAC0FCE3" blockId="2.[151,1437,402,1761]" pageId="2" pageNumber="178">Head: Vertex white with a black spot, frons white. Labial palpus white, black dorsally in basal palpomere and in basal half of second palpomere on outer surface; third palpomere black at base, in distal 2/5 on ventral surface, near apex on dorsal surface. Antenna with scape black on dorsal surface and along posterior margin, white on ventral surface and along anterior margin; flagellum greyish brown, white in basal 3/5 along anterior margin.</paragraph>
<paragraph id="ED3C526AFFD5AC58FF26CA44FBEDFA8F" blockId="2.[151,1437,402,1761]" pageId="2" pageNumber="178">Thorax: Collar white, black basally. Mesonotum greyish white, with a black rounded spot anteromedially, with a pair of black spots medially and distally. Tegula greyish white, with black spot laterobasally. Forewing greyish white, black in basal 1/4 of costal margin; overlaid with 18 black markings: six black stripes in basal 3/5 below costal margin, the longest one extending from basal 1/3 transversely to distal 2/5 below costal margin, ovate silver grey spot laid between basal and second stripes, two short stripes in distal 2/5 below costal margin; cell with a short triangular black stripe beyond middle, with a black stripe at distal 1/3; distal stripe black, triangular, below its left placed a diffused spot; fold with black dot at base, rectangular stripe near base, with tadpole-shaped stripe near middle headed towards base, rounded black dot placed below its middle, narrow stripe above its middle; a black dot set near base above dorsum; ten marginal dots extending from distal part of costal margin along termen to tornus; fringe white basally, brown along middle of termen. Hindwing and fringe pale grey. Legs greyish white; coxa of foreleg with a black spot ventrally, femur and tibia black dorsally, tarsus black; femur of midleg mixed with black dorsally, with a black spot at base, tibia black dorsally, greyish white near base, at middle and at apex, tarsus black, greyish white at base of basal tarsomere and at apices of basal two tarsomeres; femur of hindleg mixed with yellowish brown dorsally, tibia with yellowish brown long hairs, tarsus mixed with black on inside, black at apical one tarsomere, black on outside, greyish white at apices of basal four tarsomeres.</paragraph>
<paragraph id="ED3C526AFFD5AC58FF26CC58FABFF98B" blockId="2.[151,1437,402,1761]" pageId="2" pageNumber="178">
Abdomen: Male genitalia (
<figureCitation id="75B84EEFFFD5AC58FE1DCC58FD86FAA3" box="[508,590,1375,1401]" captionStart="FIGURES 10–15" captionStartId="11.[151,264,1898,1922]" captionTargetBox="[232,1362,194,1860]" captionTargetId="figure-21@11.[206,1382,181,1874]" captionTargetPageId="11" captionText="FIGURES 10–15. Male genitalia of Ethmia spp. 10, E. anatiformis, slide No. ZXJ20107; 11, E. anomala sp. nov., holotype, slide No. ZXJ20076; 12, E. corrugata sp. nov., holotype, slide No. ZXJ20108; 13, E. adunatimacula sp. nov., holotype, slide No. ZXJ20202; 14, E. crocosoma, slide No. ZXJ20129; 15, E. rectiuga sp. nov., holotype, slide No. ZXJ19650. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7147076" httpUri="https://zenodo.org/record/7147076/files/figure.png" pageId="2" pageNumber="178">Fig. 11</figureCitation>
) with uncus quadrate in basal 1/4, asymmetrically bifurcate in distal 3/4, slender and bending sideways, pointed apically. Gnathos shorter than uncus; caudal part inconspicuous; oral part lip-shaped, with fine denticles on posterior margin. Labis short digitate, with dense setae. Valva with costa longer than half of its length, almost uniform in width, rounded apically; sacculus sub-quadrate, produced inward to a stout process basally, dorsal margin with a small digitate process near base, ventral half with dense long setae; cucullus sub-quadrate in basal half, digitate in distal half, covered with dense setae. Juxta widely banded, posterolateral lobes narrowed to slender apex. Phallus curved basally, with several unequal cornuti, apical one largest, drop-shaped.
</paragraph>
<paragraph id="ED3C526AFFD5AC58FF26CF5CFE59F9AF" blockId="2.[151,1437,402,1761]" box="[199,401,1627,1653]" pageId="2" pageNumber="178">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="A59901E1FFD5AC58FF26CF79FDEEF943" box="[199,550,1662,1689]" pageId="2" pageNumber="178" type="distribution">
<paragraph id="ED3C526AFFD5AC58FF26CF79FDEEF943" blockId="2.[151,1437,402,1761]" box="[199,550,1662,1689]" pageId="2" pageNumber="178">
<emphasis id="DFF78E78FFD5AC58FF26CF79FEA9F942" bold="true" box="[199,353,1662,1688]" pageId="2" pageNumber="178">Distribution.</emphasis>
<collectingCountry id="959412FAFFD5AC58FE86CF78FE62F943" box="[359,426,1663,1689]" name="China" pageId="2" pageNumber="178">China</collectingCountry>
(
<collectingRegion id="2F479C88FFD5AC58FE58CF78FDD1F943" box="[441,537,1663,1689]" country="China" name="Shaanxi" pageId="2" pageNumber="178">Shaanxi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="A59901E1FFD5AC58FF26CFA4FE53F93B" pageId="2" pageNumber="178" type="etymology">
<paragraph id="ED3C526AFFD5AC58FF26CFA4FE53F93B" blockId="2.[151,1437,402,1761]" pageId="2" pageNumber="178">
<emphasis id="DFF78E78FFD5AC58FF26CFA4FE86F967" bold="true" box="[199,334,1699,1725]" pageId="2" pageNumber="178">Etymology.</emphasis>
The specific epithet is derived from the Latin
<emphasis id="DFF78E78FFD5AC58FC8ECFA4FC14F967" box="[879,988,1699,1725]" italics="true" pageId="2" pageNumber="178">anomalus</emphasis>
, referring to the uncus asymmetrically bifurcated in distal 3/4.
</paragraph>
</subSubSection>
</treatment>
</document>
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