182 lines
21 KiB
XML
182 lines
21 KiB
XML
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<mods:title id="675D48E31C7A6628AF38A346460C3EAE">Description Of Two Species Of Hoploscaphites (Ammonoidea: Ancyloceratina) From The Upper Cretaceous (Lower Maastrichtian) Of The U. S. Western Interior</mods:title>
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<mods:namePart id="F077A5D6CAA32857C7085DE6F3B74467">Landman, Neil H.</mods:namePart>
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<mods:affiliation id="56FCE2A6CB1C8BD6E50E8E0B83B3BA44">Division of Paleontology (Invertebrates), American Museum of Natural History, New York</mods:affiliation>
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<mods:namePart id="A30AE193E3D405C3E2606DC9D0BA1BDA">Kennedy, W. James</mods:namePart>
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<mods:affiliation id="2A8470487C07A1A4790C9E70F6634F6A">Oxford University Museum of Natural History, Oxford</mods:affiliation>
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<mods:namePart id="FB2309947CAFDE966E63CC50CB7512BA">Larson, Neal L.</mods:namePart>
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<mods:affiliation id="8E1D4DE94D87190B10BC9547AC7CBD4D">Larson Paleontology Unlimited, Keystone, South Dakota</mods:affiliation>
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<mods:namePart id="0CBADFFB2266F131003C51705DE37AC7">Grier, Joyce C.</mods:namePart>
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<mods:namePart id="5F70821C3B8A6B6DCF875CF6C524DE29">Grier, James W.</mods:namePart>
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<mods:affiliation id="C6B5DEFBB4ABD16F62C760664A48FF15">Department of Biological Sciences, North Dakota State University, Fargo</mods:affiliation>
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<treatment id="651BDA298C25A304FF1A73D188B0178D" ID-DOI="http://doi.org/10.5281/zenodo.4631215" ID-GBIF-Taxon="180670910" ID-Zenodo-Dep="4631215" LSID="urn:lsid:plazi:treatment:651BDA298C25A304FF1A73D188B0178D" httpUri="http://treatment.plazi.org/id/651BDA298C25A304FF1A73D188B0178D" lastPageNumber="22" pageId="22" pageNumber="22">
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<subSubSection id="A5A838B48C25A304FF1A73D189131719" box="[179,565,506,530]" pageId="22" pageNumber="22" type="nomenclature">
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<paragraph id="ED0D6B3F8C25A304FF1A73D189131719" blockId="22.[179,565,506,530]" box="[179,565,506,530]" pageId="22" pageNumber="22">
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Genus
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<taxonomicName id="2AB210BC8C25A304FEA873D189131719" ID-CoL="6379R" authority="Nowak, 1911" authorityName="Nowak" authorityYear="1911" box="[257,565,506,530]" class="Cephalopoda" family="Scaphitidae" genus="Hoploscaphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="genus">
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<emphasis id="DFC6B72D8C25A304FEA873D18AB91719" box="[257,415,506,530]" italics="true" pageId="22" pageNumber="22">Hoploscaphites</emphasis>
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<bibRefCitation id="892316CE8C25A304FE0F73D189131719" author="Nowak, J." box="[422,565,506,530]" pageId="22" pageNumber="22" pagination="547 - 589" refId="ref27081" refString="Nowak, J. 1911. Untersuchungen uber die Cephalopoden der oberen Kreide in Polen. II Teil: die Skaphiten. Bulletin de l'Academie des Sciences de Cracovie Series B 7: 547 - 589." type="journal article" year="1911">Nowak, 1911</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="A5A838B48C25A304FFDD7002893E1789" pageId="22" pageNumber="22" type="synonymic_list">
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<paragraph id="ED0D6B3F8C25A304FFDD7002893E1789" blockId="22.[108,637,552,1734]" pageId="22" pageNumber="22">
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<heading id="B645DC538C25A304FFDD7002893E1789" centered="true" fontSize="9" level="2" pageId="22" pageNumber="22" reason="2">
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[=
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<taxonomicName id="2AB210BC8C25A304FF3D70038930174B" authority="Atabekian, 1979: 523" authorityName="Atabekian" authorityPageNumber="523" authorityYear="1979" box="[148,534,552,576]" class="Cephalopoda" genus="Mesoscaphites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="22" phylum="Mollusca" rank="genus">
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<emphasis id="DFC6B72D8C25A304FF3D70038A0F174B" box="[148,297,552,576]" italics="true" pageId="22" pageNumber="22">Mesoscaphites</emphasis>
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Atabekian, 1979: 523
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</taxonomicName>
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(nomen nudum) fide Kennedy, 1986;
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<bibRefCitation id="892316CE8C25A304FE637062897D176A" author="Wright, C. W." box="[458,603,585,609]" pageId="22" pageNumber="22" refId="ref28011" refString="Wright, C. W. 1996. Treatise on invertebrate paleontology: part L, Mollusca 4 (revised), Cretaceous Ammonoidea: 362 pp. Boulder, CO: Geological Society of America." type="book" year="1996">Wright, 1996</bibRefCitation>
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;
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<taxonomicName id="2AB210BC8C25A304FF7B7041892A1789" authority="Riccardi, 1983: 14" authorityName="Riccardi" authorityPageNumber="14" authorityYear="1983" box="[210,524,618,642]" class="Cephalopoda" family="Scaphitidae" genus="Jeletzkytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="genus">
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<emphasis id="DFC6B72D8C25A304FF7B70418A181789" box="[210,318,618,642]" italics="true" pageId="22" pageNumber="22">Jeletzkytes</emphasis>
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<bibRefCitation id="892316CE8C25A304FEED7041892A1789" author="Riccardi, A. C." box="[324,524,618,642]" pageId="22" pageNumber="22" pagination="1 - 51" refId="ref27243" refString="Riccardi, A. C. 1983. Scaphitids from the Upper Campanian-lower Maastrichtian Bearpaw Formation of the Western Interior of Canada. Geological Survey of Canada Bulletin 354: 1 - 51." type="journal article" year="1983">Riccardi, 1983: 14</bibRefCitation>
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</taxonomicName>
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].
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="A5A838B48C25A304FF2570A78BD717EE" pageId="22" pageNumber="22" type="type_taxon">
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<paragraph id="ED0D6B3F8C25A304FF2570A78BD717EE" blockId="22.[108,637,552,1734]" pageId="22" pageNumber="22">
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<typeStatus id="3209D59D8C25A304FF2570A78BE517A8" box="[140,195,652,675]" pageId="22" pageNumber="22">TYPE</typeStatus>
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SPECIES:
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<emphasis id="DFC6B72D8C25A304FE9870A78A8C17A8" box="[305,426,652,675]" italics="true" pageId="22" pageNumber="22">Ammonites</emphasis>
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<taxonomicName id="2AB210BC8C25A304FE1A70A78A0B17CF" ID-CoL="3MN2D" authority="J. Sowerby (1817: 189" authorityName="J. Sowerby" authorityYear="1817" class="Cephalopoda" family="Scaphitidae" genus="Hoploscaphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="species" species="constrictus">
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<emphasis id="DFC6B72D8C25A304FE1A70A7890317A8" box="[435,549,652,675]" italics="true" pageId="22" pageNumber="22">constrictus</emphasis>
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J.
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<bibRefCitation id="892316CE8C25A304FDEC70A08A0B17CF" author="Sowerby, J." pageId="22" pageNumber="22" refId="ref27625" refString="Sowerby, J. 1817. The mineral conchology of Great Britain, vol. 2. London: the author. [7 vols.]" type="book" year="1817">Sowerby (1817: 189</bibRefCitation>
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</taxonomicName>
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, pl. A, fig. 1), by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="A5A838B48C25A304FF2570C48A3C11BF" pageId="22" pageNumber="22" type="diagnosis">
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<paragraph id="ED0D6B3F8C25A304FF2570C48A3C11BF" blockId="22.[108,637,552,1734]" pageId="22" pageNumber="22">
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DIAGNOSIS: “Small to large scaphites, strongly dimorphic, with broad variation in degree of whorl compression ranging from slender to robust, with involute phragmocone, short to long shaft, and weakly recurved hook; apertural angle ranging from approximately 35° to 85°; aperture constricted with dorsal projection; ribs straight to flexuous, increasing by branching and intercalation, with weak to strong adoral projection on venter; adult shell with or without umbilicolateral, flank, and ventrolateral tubercles; suture fairly indented, with symmetrically to slightly asymmetrically bifid first lateral lobe” (
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<bibRefCitation id="892316CE8C25A304FDB876508A2811BF" author="Landman, N. H. & W. J. Kennedy & W. A. Cobban & N. L. Larson" pageId="22" pageNumber="22" pagination="1 - 242" refId="ref26072" refString="Landman, N. H., W. J. Kennedy, W. A. Cobban, and N. L. Larson. 2010. Scaphites of the " nodosus group " from the Upper Cretaceous (Campanian) of the Western Interior of North America. Bulletin of the American Museum of Natural History 342: 1 - 242." type="journal article" year="2010">Landman et al., 2010: 93</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="A5A838B48C25A304FF25769688B0178D" pageId="22" pageNumber="22" type="discussion">
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<paragraph id="ED0D6B3F8C25A304FF25769688B0178D" blockId="22.[108,637,552,1734]" lastBlockId="22.[684,1214,226,647]" pageId="22" pageNumber="22">
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DISCUSSION: In their monograph on the scaphites of the “
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<taxonomicName id="2AB210BC8C25A304FE8E76F58A5B11FD" baseAuthorityName="Owen" baseAuthorityYear="1852" box="[295,381,1246,1270]" class="Cephalopoda" family="Scaphitidae" genus="Hoploscaphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="species" species="nodosus">
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<emphasis id="DFC6B72D8C25A304FE8E76F58A5B11FD" box="[295,381,1246,1270]" italics="true" pageId="22" pageNumber="22">nodosus</emphasis>
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</taxonomicName>
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group,”
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<bibRefCitation id="892316CE8C25A304FE7176F58B901013" author="Landman, N. H. & W. J. Kennedy & W. A. Cobban & N. L. Larson" pageId="22" pageNumber="22" pagination="1 - 242" refId="ref26072" refString="Landman, N. H., W. J. Kennedy, W. A. Cobban, and N. L. Larson. 2010. Scaphites of the " nodosus group " from the Upper Cretaceous (Campanian) of the Western Interior of North America. Bulletin of the American Museum of Natural History 342: 1 - 242." type="journal article" year="2010">Landman et al. (2010)</bibRefCitation>
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treated
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<taxonomicName id="2AB210BC8C25A304FEB276D4891B1013" authority="Riccardi, 1983" authorityName="Riccardi" authorityYear="1983" box="[283,573,1279,1304]" class="Cephalopoda" family="Scaphitidae" genus="Jeletzkytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="genus">
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||
<emphasis id="DFC6B72D8C25A304FEB276D48AAC101C" box="[283,394,1279,1303]" italics="true" pageId="22" pageNumber="22">Jeletzkytes</emphasis>
|
||
<bibRefCitation id="892316CE8C25A304FE3F76D4891B1013" author="Riccardi, A. C." box="[406,573,1279,1304]" pageId="22" pageNumber="22" pagination="1 - 51" refId="ref27243" refString="Riccardi, A. C. 1983. Scaphitids from the Upper Campanian-lower Maastrichtian Bearpaw Formation of the Western Interior of Canada. Geological Survey of Canada Bulletin 354: 1 - 51." type="journal article" year="1983">Riccardi, 1983</bibRefCitation>
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||
</taxonomicName>
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, as a junior subjective synonym of
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<taxonomicName id="2AB210BC8C25A304FE7A770B89501033" authorityName="Nowak" authorityYear="1911" box="[467,630,1312,1336]" class="Cephalopoda" family="Scaphitidae" genus="Hoploscaphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="genus">
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||
<emphasis id="DFC6B72D8C25A304FE7A770B89501033" box="[467,630,1312,1336]" italics="true" pageId="22" pageNumber="22">Hoploscaphites</emphasis>
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</taxonomicName>
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. They argued that the shape of the shell, the pattern of ornamentation, and the complexity of the suture are the same in both genera, the only differences being the degree of compression and, as a result, the flexuosity of the ribs and the size of the tubercles. They noted that such variation already exists within the genus
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<taxonomicName id="2AB210BC8C25A304FE157423897F132B" authorityName="Nowak" authorityYear="1911" box="[444,601,1544,1568]" class="Cephalopoda" family="Scaphitidae" genus="Hoploscaphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="genus">
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||
<emphasis id="DFC6B72D8C25A304FE157423897F132B" box="[444,601,1544,1568]" italics="true" pageId="22" pageNumber="22">Hoploscaphites</emphasis>
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</taxonomicName>
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, as previously defined, and exemplified by the
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<typeStatus id="3209D59D8C25A304FDE47401895A1349" box="[589,636,1578,1602]" pageId="22" pageNumber="22">type</typeStatus>
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species
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<taxonomicName id="2AB210BC8C25A304FF6D74608A7B1369" authorityName="J. Sowerby" authorityYear="1817" box="[196,349,1611,1634]" class="Cephalopoda" family="Scaphitidae" genus="Hoploscaphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="species" species="constrictus">
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||
<emphasis id="DFC6B72D8C25A304FF6D74608BFE1369" box="[196,216,1611,1634]" italics="true" pageId="22" pageNumber="22">H</emphasis>
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||
.
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<emphasis id="DFC6B72D8C25A304FF4074608A7B1369" box="[233,349,1611,1634]" italics="true" pageId="22" pageNumber="22">constrictus</emphasis>
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</taxonomicName>
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.
|
||
<bibRefCitation id="892316CE8C25A304FEC77461895D1369" author="Wright, C. W." box="[366,635,1610,1634]" pageId="22" pageNumber="22" refId="ref28011" refString="Wright, C. W. 1996. Treatise on invertebrate paleontology: part L, Mollusca 4 (revised), Cretaceous Ammonoidea: 362 pp. Boulder, CO: Geological Society of America." type="book" year="1996">Wright (1996: 261–262)</bibRefCitation>
|
||
cited the same reason for synonymizing the two genera: “Separation of the large and inflated species of the
|
||
<taxonomicName id="2AB210BC8C25A304FF4674868A6013CE" baseAuthorityName="Owen" baseAuthorityYear="1852" box="[239,326,1709,1733]" class="Cephalopoda" family="Scaphitidae" genus="Hoploscaphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="species" species="nodosus">
|
||
<emphasis id="DFC6B72D8C25A304FF4674868A6013CE" box="[239,326,1709,1733]" italics="true" pageId="22" pageNumber="22">nodosus</emphasis>
|
||
</taxonomicName>
|
||
group as
|
||
<taxonomicName id="2AB210BC8C25A304FE697486890813CE" authorityName="Riccardi" authorityYear="1983" box="[448,558,1709,1733]" class="Cephalopoda" family="Scaphitidae" genus="Jeletzkytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="genus">
|
||
<emphasis id="DFC6B72D8C25A304FE697486890813CE" box="[448,558,1709,1733]" italics="true" pageId="22" pageNumber="22">Jeletzkytes</emphasis>
|
||
</taxonomicName>
|
||
seems unnecessary, given the great variation within most scaphitid species.” Later, Cooper (1994) erected the new subgenus
|
||
<taxonomicName id="2AB210BC8C25A304FC5D730F8FAB1437" box="[1012,1165,292,316]" class="Cephalopoda" family="Scaphitidae" genus="Jeletzkyites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="subGenus" subGenus="Karlwaageites">
|
||
<emphasis id="DFC6B72D8C25A304FC5D730F8FAB1437" box="[1012,1165,292,316]" italics="true" pageId="22" pageNumber="22">Karlwaageites</emphasis>
|
||
</taxonomicName>
|
||
for derived members of
|
||
<taxonomicName id="2AB210BC8C25A304FC06736E8F061456" authorityName="Riccardi" authorityYear="1983" box="[943,1056,325,349]" class="Cephalopoda" family="Scaphitidae" genus="Jeletzkytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="genus">
|
||
<emphasis id="DFC6B72D8C25A304FC06736E8F061456" box="[943,1056,325,349]" italics="true" pageId="22" pageNumber="22">Jeletzkytes</emphasis>
|
||
</taxonomicName>
|
||
from North America that bear lateral tubercles. However, the number and distribution of lateral tubercles can vary even within a single species, casting doubt on the utility of this character for subgeneric separation. Until a thorough phylogenetic revision of all scaphitid genera and subgenera is undertaken, we prefer to follow a more conservative approach and treat all these species in the genus
|
||
<taxonomicName id="2AB210BC8C25A304FD5B704588B6178D" authorityName="Nowak" authorityYear="1911" box="[754,912,622,646]" class="Cephalopoda" family="Scaphitidae" genus="Hoploscaphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="22" pageNumber="22" phylum="Mollusca" rank="genus">
|
||
<emphasis id="DFC6B72D8C25A304FD5B704588B6178D" box="[754,912,622,646]" italics="true" pageId="22" pageNumber="22">Hoploscaphites</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |