treatments-xml/data/65/16/CC/6516CC460D16FFFFFF7A8AB665FD66B1.xml

211 lines
25 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="524D5DE8EBFD0757E922D76E9C6D7EFB" ID-CLB-Dataset="46982" ID-DOI="10.5281/zenodo.199681" ID-GBIF-Dataset="62e1623e-d85e-46ea-8edf-6cb01d7c88eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="199681" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460343126729" checkinUser="plazi" docAuthor="Blaimer, Bonnie B." docDate="2010" docId="6516CC460D16FFFFFF7A8AB665FD66B1" docLanguage="en" docName="zt02714p039.pdf" docOrigin="Zootaxa 2714" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Crematogaster malala Blaimer, 2010, NEW SPECIES" docType="treatment" docVersion="10" lastPageNumber="25" masterDocId="992FB43E0D01FFE7FFED8A216475640D" masterDocTitle="Taxonomy and Natural History of the Crematogaster (Decacrema) - group (Hymenoptera: Formicidae) in Madagascar" masterLastPageNumber="39" masterPageNumber="1" pageNumber="24" updateTime="1732492176645" updateUser="ExternalLinkService">
<mods:mods id="B0B37A46B4CAFE32B20148D15D9F7AB6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="80807AA993B58E43D9D9C29A7FE99B0F">
<mods:title id="24123527ED586C7635A442C12E3BA3FF">Taxonomy and Natural History of the Crematogaster (Decacrema) - group (Hymenoptera: Formicidae) in Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="53F35E720B316CBA22AB0D30FCA4C9FC" type="personal">
<mods:role id="6A1EA6EC1537DFBBF841E7713CDCDBC0">
<mods:roleTerm id="472E1E40D864948D21C8E035CF7C7E98">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D36050014D5F8FBFEE0991D13540EC48">Blaimer, Bonnie B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C56BAEE954937E900D03E5DC09CFAAEC">text</mods:typeOfResource>
<mods:relatedItem id="C9872FA9A76C4A41E9B11F99FFBBA86C" type="host">
<mods:titleInfo id="5B326774908BF4BB896159E0EBD6FC57">
<mods:title id="A4851DAB971F15095EADB4996C015444">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7E31035EEBBADF598A2F25133C14B814">
<mods:date id="3B6A2E780F4BA18257683BC1598BE4AE">2010</mods:date>
<mods:detail id="456109DBDF25024C609B3920ACFBF50B" type="volume">
<mods:number id="AA0CA6BFEB1F1EEBA8787AE00EAA72B0">2714</mods:number>
</mods:detail>
<mods:extent id="AA68BF06BD4DD76D24D5A18121A52FF3" unit="page">
<mods:start id="0469437984D9381AA9E8CE11AF387FA1">1</mods:start>
<mods:end id="2D42040467EFF2F4E1ADFEAAE2BA76D6">39</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F0D1DB6B9ADA49D8FF72DFE303ECE737">journal article</mods:classification>
<mods:identifier id="F32FCB203CAB9EFC2BB66065147AA8D5" type="CLB-Dataset">46982</mods:identifier>
<mods:identifier id="61D99F18E41E1EC438CDF33003549C58" type="DOI">10.5281/zenodo.199681</mods:identifier>
<mods:identifier id="B9E004DAB98E9B1A0CF9B468FEE88754" type="GBIF-Dataset">62e1623e-d85e-46ea-8edf-6cb01d7c88eb</mods:identifier>
<mods:identifier id="E90B7C26F51EBDDC52F693A6CD788DCE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="37EF2AE39A842D046E193E19F919956B" type="Zenodo-Dep">199681</mods:identifier>
</mods:mods>
<treatment id="6516CC460D16FFFFFF7A8AB665FD66B1" ID-DOI="http://doi.org/10.5281/zenodo.6199376" ID-GBIF-Taxon="119406769" ID-Zenodo-Dep="6199376" LSID="urn:lsid:plazi:treatment:6516CC460D16FFFFFF7A8AB665FD66B1" httpUri="http://treatment.plazi.org/id/6516CC460D16FFFFFF7A8AB665FD66B1" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="A5A52EDB0D16FFF0FF7A8AB6653564D9" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="ED007D500D16FFF0FF7A8AB6660464BC" blockId="23.[151,625,151,212]" box="[151,625,151,177]" pageId="23" pageNumber="24">
<heading id="B648CA3C0D16FFF0FF7A8AB6660464BC" bold="true" box="[151,625,151,177]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="DFCBA1420D16FFF0FF7A8AB6660464BC" bold="true" box="[151,625,151,177]" pageId="23" pageNumber="24">
<taxonomicName id="2ABF06D30D16FFF0FF7A8AB665D664BC" ID-CoL="CLMN9" box="[151,419,151,177]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="malala" status="sp. nov.">
<emphasis id="DFCBA1420D16FFF0FF7A8AB665D664BC" bold="true" box="[151,419,151,177]" italics="true" pageId="23" pageNumber="24">Crematogaster malala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C4F81C390D16FFF0FE478AB6660464BC" box="[426,625,151,177]" pageId="23" pageNumber="24" rank="species">NEW SPECIES</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="ED007D500D16FFF0FF7A8A9B653564D9" blockId="23.[151,625,151,212]" box="[151,320,186,212]" pageId="23" pageNumber="24">
(
<figureCitation id="758461D50D16FFF0FF728A9B648664D9" box="[159,243,186,212]" captionStart="FIGURES 32 37" captionStartId="29.[151,270,1974,1998]" captionTargetBox="[190,1397,196,1939]" captionTargetId="figure@29.[179,1407,166,1953]" captionTargetPageId="29" captionText="FIGURES 32 37. Malagasy Crematogaster (Decacrema) species; a: full-face view, b: lateral view, c: dorsal view. 32: C. nosibeensis; 33: C. sisa; 34: C. sabatra; 35: C. mahery; 36: C. malala; 37: C. grevei." httpUri="https://zenodo.org/record/199685/files/figure.png" pageId="23" pageNumber="24">Fig. 36</figureCitation>
&amp;
<figureCitation id="758461D50D16FFF0FEF48A9B654D64D9" box="[281,312,186,212]" captionStart="FIGURES 38 43" captionStartId="30.[151,270,1894,1918]" captionTargetBox="[221,1364,193,1874]" captionTargetId="figure@30.[200,1385,164,1882]" captionTargetPageId="30" captionText="FIGURES 38 43. Malagasy Crematogaster (Decacrema) species; a: petiole &amp; postpetiole in dorsal view, b: petiole &amp; postpetiole in lateral view. 38: C. nosibeensis; 39: C. sisa, 40: C. sabatra; 41: C. mahery; 42: C. malala; 43: C. grevei." httpUri="https://zenodo.org/record/199686/files/figure.png" pageId="23" pageNumber="24">42</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D16FFF0FF7A8B24674066B7" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="ED007D500D16FFF0FF7A8B2466776567" blockId="23.[151,1437,261,2004]" pageId="23" pageNumber="24">
<emphasis id="DFCBA1420D16FFF0FF7A8B2465046512" bold="true" box="[151,369,261,287]" pageId="23" pageNumber="24">
<typeStatus id="3204C3F20D16FFF0FF7A8B24657C6512" box="[151,265,261,287]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
worker.
</emphasis>
<collectingCountry id="95A83DC00D16FFF0FE948B24663B6512" box="[377,590,261,287]" name="Madagascar" pageId="23" pageNumber="24">MADAGASCAR</collectingCountry>
: P.C. Ankazomivady, -20.79751, 47.17910,
<quantity id="2A47D0B50D16FFF0FBBB8B2460DF6512" box="[1110,1194,261,287]" metricMagnitude="3" metricUnit="m" metricValue="1.782" pageId="23" pageNumber="24" unit="m" value="1782.0">1782m</quantity>
, ex dead twig above ground, disturbed montane rainforest, B.L.Fisher
<emphasis id="DFCBA1420D16FFF0FCEE8B0B6737654E" box="[771,834,298,323]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
#20466 (CASENT0140925, specimen image on Antweb) (deposited in
<collectionCode id="8BAEE5950D16FFF0FE498B7165836567" box="[420,502,336,362]" pageId="23" pageNumber="24">CASC</collectionCode>
).
</paragraph>
<paragraph id="ED007D500D16FFF0FF2B8B5460C965D7" blockId="23.[151,1437,261,2004]" pageId="23" pageNumber="24">
<emphasis id="DFCBA1420D16FFF0FF2B8B54653E6582" bold="true" box="[198,331,373,399]" pageId="23" pageNumber="24">
<typeStatus id="3204C3F20D16FFF0FF2B8B5465336582" box="[198,326,373,399]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
.
</emphasis>
4w, P.N. Befotaka-Midongy, -23.84080, 46.95750,
<quantity id="2A47D0B50D16FFF0FC398B54605D6582" box="[980,1064,373,399]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="23" pageNumber="24" unit="m" value="1250.0">1250m</quantity>
, ex dead twig above ground, montane rainforest, (B.L.Fisher
<emphasis id="DFCBA1420D16FFF0FDAE8BBB66F265BE" box="[579,647,410,435]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
#15029: CASENT0193561 and CASENT0193562; #15041: CASENT0193559 and CASENT0193560) (deposited in
<collectionCode id="8BAEE5950D16FFF0FCDC8BE167F265D7" box="[817,903,448,474]" pageId="23" pageNumber="24">MCZC</collectionCode>
,
<collectionCode id="8BAEE5950D16FFF0FC7E8BE1679C65D7" box="[915,1001,448,474]" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="23" pageNumber="24">SAMC</collectionCode>
,
<collectionCode id="8BAEE5950D16FFF0FC198BE1603865D7" box="[1012,1101,448,474]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="24">MHNG</collectionCode>
,
<collectionCode id="8BAEE5950D16FFF0FBB48BE160C565D7" box="[1113,1200,448,474]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="23" pageNumber="24">UCDC</collectionCode>
).
</paragraph>
<paragraph id="ED007D500D16FFF0FF2B8BC4674066B7" blockId="23.[151,1437,261,2004]" pageId="23" pageNumber="24">
<emphasis id="DFCBA1420D16FFF0FF2B8BC465B465F2" bold="true" box="[198,449,485,511]" pageId="23" pageNumber="24">Material examined.</emphasis>
(
<collectionCode id="8BAEE5950D16FFF0FE3B8BC4665365F2" box="[470,550,485,511]" pageId="23" pageNumber="24">CASC</collectionCode>
)
<collectingCountry id="95A83DC00D16FFF0FDD18BC4677965F2" box="[572,780,485,511]" name="Madagascar" pageId="23" pageNumber="24">MADAGASCAR</collectingCountry>
<emphasis id="DFCBA1420D16FFF0FCF78BC467CF65F3" box="[794,954,485,510]" italics="true" pageId="23" pageNumber="24">Fianarantsoa</emphasis>
<quantity id="2A47D0B50D16FFF0FC2A8BC4607F65F2" box="[967,1034,485,511]" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="23" pageNumber="24" unit="km" value="36.0">36km</quantity>
S Ambalavao, Rés. Andringitra (=P.N. Andringitra): -22.23333, 47.00000,
<quantity id="2A47D0B50D16FFF0FD40882B67776629" box="[685,770,522,548]" metricMagnitude="3" metricUnit="m" metricValue="1.925" pageId="23" pageNumber="24" unit="m" value="1925.0">1925m</quantity>
(B.L.Fisher); P.N. Andringitra: -22.18767, 46.90083,
<quantity id="2A47D0B50D16FFF0FF7A8811649C6647" box="[151,233,560,586]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="23" pageNumber="24" unit="m" value="2150.0">2150m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D16FFF0FE69881165B46644" box="[388,449,560,585]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
); P.N. Befotaka-Midongy: -23.84080, 46.95750,
<quantity id="2A47D0B50D16FFF0FBF9881160136647" box="[1044,1126,560,586]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="23" pageNumber="24" unit="m" value="1250.0">1250m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D16FFF0FAEC881161486644" box="[1281,1341,560,585]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
);
<quantity id="2A47D0B50D16FFF0FAB5881161E86647" box="[1368,1437,560,586]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="23" pageNumber="24" unit="km" value="28.0">28km</quantity>
SSW Ambositra, Ankazomivady [=P.C. Ankazomivady]: -20.77500, 47.16833,
<quantity id="2A47D0B50D16FFF0FBB0887460C56662" box="[1117,1200,597,623]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="23" pageNumber="24" unit="m" value="1680.0">1680m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D16FFF0FAA1887461FF6663" box="[1356,1418,597,622]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
); P.C. Ankazomivady: -20.77667, 47.16500,
<quantity id="2A47D0B50D16FFF0FD44885B67746699" box="[681,769,634,660]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="23" pageNumber="24" unit="m" value="1780.0">1780m</quantity>
; -20.79751, 47.17910,
<quantity id="2A47D0B50D16FFF0FBF0885B60046699" box="[1053,1137,634,660]" metricMagnitude="3" metricUnit="m" metricValue="1.782" pageId="23" pageNumber="24" unit="m" value="1782.0">1782m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D16FFF0FAE3885B6139669E" box="[1294,1356,634,659]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
); R.S. Ivohibe: -22.49667, 46.95500,
<quantity id="2A47D0B50D16FFF0FDEF8881662166B7" box="[514,596,672,698]" metricMagnitude="3" metricUnit="m" metricValue="1.575" pageId="23" pageNumber="24" unit="m" value="1575.0">1575m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D16FFF0FD078881675166B4" box="[746,804,672,697]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D16FFF0FF2B88E4653962F9" pageId="23" pageNumber="24" type="description">
<paragraph id="ED007D500D16FFF0FF2B88E465BA6742" blockId="23.[151,1437,261,2004]" pageId="23" pageNumber="24">
<emphasis id="DFCBA1420D16FFF0FF2B88E4663366D2" bold="true" box="[198,582,709,735]" pageId="23" pageNumber="24">Worker measurements (n=10).</emphasis>
HW 0.891.09; HL 0.800.99; EL 0.210.25; SL 0.730.82; WL 0.97 1.16; SPL 0.180.27; PTH 0.200.24; PTL 0.230.28; PTW 0.290.41; PPL 0.170.21; PPW 0.260.36; LHT 0.700.88; CI 1.051.17; OI 0.240.28; SI 0.810.91; SPI 0.170.25, PTHI 0.841.00, PTWI 1.181.64, PPI 1.431.72; LBI 1.231.39.
</paragraph>
<paragraph id="ED007D500D16FFF0FF2B897B600D6779" blockId="23.[151,1437,261,2004]" box="[198,1144,858,884]" pageId="23" pageNumber="24">
<emphasis id="DFCBA1420D16FFF0FF2B897B65B66779" bold="true" box="[198,451,858,884]" pageId="23" pageNumber="24">Worker description.</emphasis>
Small to medium species (HW 0.891.09, WL 0.971.16).
</paragraph>
<paragraph id="ED007D500D16FFF0FF2889A165CF67E9" blockId="23.[151,1437,261,2004]" pageId="23" pageNumber="24">Masticatory margin of mandibles with 4 teeth; posterior margin of head laterally forming round corners; antennae with scapes reaching or just surpassing posterior margin; midline of eyes situated just above midline of head in full face view.</paragraph>
<paragraph id="ED007D500D16FFF0FF2889D167046102" blockId="23.[151,1437,261,2004]" pageId="23" pageNumber="24">Promesonotum with laterally rounded shoulders; promesonotal suture weakly impressed, usually complete; outline of promesonotum convex in lateral view, sometimes broken by mesonotum which is often slightly raised over pronotum and forming a small antero-medial tubercle; mesonotum postero-laterally rounded or angular; propodeal spines medium sized (SPI 0.170.25), less than half the width between their bases, evenly tapering, from straight to weakly curved, and in dorsal view weakly diverging; petiole in dorsal view trapezoidal, anteriorly flared into rounded lobes, dorso-lateral margins carinate, terminating in a small postero-lateral tubercle; subpetiolar process present as a long, moderately acute antero-ventral tooth; postpetiole distinctly bilobed and broadly medially impressed.</paragraph>
<paragraph id="ED007D500D16FFF0FF288F3B64806237" blockId="23.[151,1437,261,2004]" pageId="23" pageNumber="24">Head sculpture reduced, usually aciculate; mesosoma with meso-and metapleuron carinulate or costulate; dorsal face of propodeum weakly costulate to reticulate; posterior face largely shiny; rest of mesosoma including legs largely aciculate; dorsal face of petiole mostly shiny; postpetiole dorsally feebly reticulate; lateral and ventral face of petiole and postpetiole coarsely rugulose-reticulate; helcium dorsally carinulate; face with 24 erect setae; pilosity on dorsum of mesosoma 24 erect humeral setae, rarely one pair of erect setae at base of mesonotal tubercules; a pair of stiff, long, erect setae present on postero-lateral tubercles of petiole; dorso-posterior erect setae also present on postpetiole; abdominal tergites 47 with scattered erect pilosity.</paragraph>
<paragraph id="ED007D500D16FFF0FF288C6460A26252" blockId="23.[151,1437,261,2004]" box="[197,1239,1605,1631]" pageId="23" pageNumber="24">Colour light to dark brown, mesosoma usually lighter coloured than head and metasoma.</paragraph>
<paragraph id="ED007D500D16FFF0FF2B8C4B653962F9" blockId="23.[151,1437,261,2004]" pageId="23" pageNumber="24">
<emphasis id="DFCBA1420D16FFF0FF2B8C4B65346289" bold="true" box="[198,321,1642,1668]" pageId="23" pageNumber="24">Variation.</emphasis>
Workers from the Midongy region are darker coloured than from P.C. Ankazomivady and P.N. Andringitra localities and have a denser appressed pubescence. The mesonotum is often more prominently raised over the pronotum in the P.C. Ankazomivady and P.N. Midongy material, than in workers from the other locations.
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D16FFFFFF2B8CDE610F64F1" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="discussion">
<paragraph id="ED007D500D16FFFFFF2B8CDE610F64F1" blockId="23.[151,1437,261,2004]" lastBlockId="24.[151,1437,152,700]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="DFCBA1420D16FFF0FF2B8CDE65206314" bold="true" box="[198,341,1791,1817]" pageId="23" pageNumber="24">Comments.</emphasis>
<taxonomicName id="2ABF06D30D16FFF0FE8F8D21661A6314" box="[354,623,1792,1817]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="malala">
<emphasis id="DFCBA1420D16FFF0FE8F8D21661A6314" box="[354,623,1792,1817]" italics="true" pageId="23" pageNumber="24">Crematogaster malala</emphasis>
</taxonomicName>
is characterized by a combination of the following features. The petiole has postero-lateral tubercles that each bear a single, stiff and long erect seta. The postpetiole is distinctly bilobed and broadly medially impressed and also bears a pair of dorso-posterior stiff, long erect setae. These characters, together with the erect pilosity on abdominal tergites 47 and the 4 mandibular teeth, are shared with
<taxonomicName id="2ABF06D30D16FFF0FEB98DB465B563A3" box="[340,448,1941,1966]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D16FFF0FEB98DB465B563A3" box="[340,448,1941,1966]" italics="true" pageId="23" pageNumber="24">C. grevei</emphasis>
</taxonomicName>
. In distinction to
<taxonomicName id="2ABF06D30D16FFF0FD798DB4668A63A3" box="[660,767,1941,1966]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D16FFF0FD798DB4668A63A3" box="[660,767,1941,1966]" italics="true" pageId="23" pageNumber="24">C. grevei</emphasis>
</taxonomicName>
however the lateral portions of the promesonotum in
<taxonomicName id="2ABF06D30D16FFF0FA6C8DB4649963D9" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="malala">
<emphasis id="DFCBA1420D16FFF0FA6C8DB4649963D9" italics="true" pageId="23" pageNumber="24">C. malala</emphasis>
</taxonomicName>
are not raised with respect to the median portion and are postero-lateral rounded or angular. The metanotal groove further lacks lateral carinae and the promesonotum forms a convex outline in lateral view. In molecular phylogenetic analyses this species is well supported as a distinct clade by both nuclear and mitochondrial data, and results suggest it is the sister taxon to the rest of the
<collectingCountry id="95A83DC00D19FFFFFBF58AC360FF64F1" box="[1048,1162,226,252]" name="Madagascar" pageId="24" pageNumber="25">Malagasy</collectingCountry>
<taxonomicName id="2ABF06D30D19FFFFFB7C8AC3616364F6" box="[1169,1302,226,251]" class="Insecta" family="Formicidae" genus="Decacrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="DFCBA1420D19FFFFFB7C8AC3616364F6" box="[1169,1302,226,251]" italics="true" pageId="24" pageNumber="25">Decacrema</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D19FFFFFF2B8B2661606641" pageId="24" pageNumber="25" type="distribution">
<paragraph id="ED007D500D19FFFFFF2B8B2661606641" blockId="24.[151,1437,152,700]" pageId="24" pageNumber="25">
<emphasis id="DFCBA1420D19FFFFFF2B8B26658B652C" bold="true" box="[198,510,263,289]" pageId="24" pageNumber="25">Distribution and biology.</emphasis>
Collections of this species are rare, with a few colonies from three locations of higher elevation rainforests (&gt;
<quantity id="2A47D0B50D19FFFFFDED8B0C6621654A" box="[512,596,301,327]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="24" pageNumber="25" unit="m" value="1250.0">1250m</quantity>
) in the Ankaratra (Ankazomivady), Andringitra and Midongy massifs (
<figureCitation id="758461D50D19FFFFFF4C8B7364836561" box="[161,246,338,364]" captionStart="FIGURES 54 57" captionStartId="33.[151,270,1948,1972]" captionTargetBox="[294,1298,205,1923]" captionTargetId="figure@33.[285,1302,194,1927]" captionTargetPageId="33" captionText="FIGURES 54 57. Distributions of Malagasy Crematogaster (Decacrema). 54: C. grevei; 55: C. mahery; 56: C. malala; 57: C. nosibeensis." httpUri="https://zenodo.org/record/199689/files/figure.png" pageId="24" pageNumber="25">
<emphasis id="DFCBA1420D19FFFFFF4C8B7364836561" bold="true" box="[161,246,338,364]" pageId="24" pageNumber="25">Fig. 56</emphasis>
</figureCitation>
). The isolated, montane distribution invites speculation that glacial cycles have played an important role in shaping this species evolutionary history and distribution, as has been suggested for other taxa in
<collectingCountry id="95A83DC00D19FFFFFF7A8BBC655D65BA" box="[151,296,413,439]" name="Madagascar" pageId="24" pageNumber="25">Madagascar</collectingCountry>
(for an overview see
<bibRefCitation id="892E00A10D19FFFFFDDF8BBC676A65BA" author="Vences" box="[562,799,413,439]" pageId="24" pageNumber="25" refString="Vences, M., Wollenberg, K. C., Vieites, D. R. &amp; Lees, D. C. (2009) Madagascar as a model region of species diversification. Trends in Ecology &amp; Evolution, 24, 456 - 465." type="journal article" year="2009">
Vences
<emphasis id="DFCBA1420D19FFFFFD7D8BBC66B865BB" box="[656,717,413,438]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
, 2009
</bibRefCitation>
). Next to nothing is known about the biology of this species. Fishers collections of
<taxonomicName id="2ABF06D30D19FFFFFDE78BE3660B65D6" box="[522,638,450,475]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="malala">
<emphasis id="DFCBA1420D19FFFFFDE78BE3660B65D6" box="[522,638,450,475]" italics="true" pageId="24" pageNumber="25">C. malala</emphasis>
</taxonomicName>
workers document these ants as nesting in dead twigs (n=4) and in carton nests (n=1); queens or males of this species have not yet been discovered. A recent search for
<taxonomicName id="2ABF06D30D19FFFFFAC48BC961E9660C" box="[1321,1436,488,513]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="malala">
<emphasis id="DFCBA1420D19FFFFFAC48BC961E9660C" box="[1321,1436,488,513]" italics="true" pageId="24" pageNumber="25">C. malala</emphasis>
</taxonomicName>
in the P.N. Andringitra at about
<quantity id="2A47D0B50D19FFFFFDE1882C6615662A" box="[524,608,525,551]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="24" pageNumber="25" unit="m" value="1800.0">1800m</quantity>
, not far from where it had been collected before, was unsuccessful and suggests that these ants might be highly restricted in habitat and possess inconspicuous life habits.
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D19FFFFFF2B887665FD66B1" pageId="24" pageNumber="25" type="etymology">
<paragraph id="ED007D500D19FFFFFF2B887665FD66B1" blockId="24.[151,1437,152,700]" pageId="24" pageNumber="25">
<emphasis id="DFCBA1420D19FFFFFF2B88766526667C" bold="true" box="[198,339,599,625]" pageId="24" pageNumber="25">Etymology.</emphasis>
This species name is derived from the
<collectingCountry id="95A83DC00D19FFFFFCDE887967D0667F" box="[819,933,600,626]" name="Madagascar" pageId="24" pageNumber="25">Malagasy</collectingCountry>
adjective “
<taxonomicName id="2ABF06D30D19FFFFFBC6887960F5667F" box="[1067,1152,600,626]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="malala">malala</taxonomicName>
”, meaning “the beloved one”, inspired by happiness about the discovery of its morphological distinctness compared to the other species in the group.
</paragraph>
</subSubSection>
</treatment>
</document>