331 lines
32 KiB
XML
331 lines
32 KiB
XML
<document id="70CCA8AC76CAF272FCB3AC5219477774" ID-CLB-Dataset="299340" ID-DOI="10.3897/zookeys.1205.120033" ID-GBIF-Dataset="15d96bea-75a1-4b35-a306-3aecdc1513a7" ID-ZooBank="1093570E-D1C7-4918-BD8F-0BC3D3F486E5" ID-publisher-id="120033" URI-arpha="541CAC61-3728-5591-9EFB-B1793F52FBE7" URI-zoobank="http://zoobank.org/1093570E-D1C7-4918-BD8F-0BC3D3F486E5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1719861896249" checkinUser="pensoft" docAuthor="Wood, Thomas J., Ismael, Halgurd R., Baiocchi, Daniele, Hamad, Mudhafar I., Bapir, Tahseen T. & Selis, Marco" docDate="2024" docId="64180BEE5A8355D4A3089FA3C2E06A0F" docLanguage="en" docName="ZooKeys 1205: 267-298" docOrigin="ZooKeys 1205" docSource="https://zookeys.pensoft.net/article/120033/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Andrena (Aciandrena) duhokensis Wood & Ismael & Baiocchi & Hamad & Bapir & Selis 2024, sp. nov." docType="treatment" docUuid="3C35CEE1-7063-4C23-8577-1BA21A6CCAFE" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="298" masterDocId="541CAC61372855919EFBB1793F52FBE7" masterDocTitle="A first revision of the Andrena of Iraq (Hymenoptera, Andrenidae), with the description of two new species from Iraqi Kurdistan and additional records from surrounding countries" masterLastPageNumber="298" masterPageNumber="267" pageNumber="267" updateTime="1719886393335" updateUser="ExternalLinkService">
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<mods:mods id="9008D4B05B6BD86A2E5406D4C62FF2D9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B5B24CF08D59E6EBA2075BF482BB04FD">
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<mods:title id="CA72CDFA1AE4775420E8566EDF7FC235">A first revision of the Andrena of Iraq (Hymenoptera, Andrenidae), with the description of two new species from Iraqi Kurdistan and additional records from surrounding countries</mods:title>
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</mods:titleInfo>
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<mods:name id="5324947E0F2E66FA987A6045226D2B90" type="personal">
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<mods:role id="A7C630AA704CF2441625083A79EDD53F">
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<mods:roleTerm id="7069CFB837ADB8C2989012D1A4438E50">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0C7337BA37EF94BF1D3EDA9FDD7A7500">Wood, Thomas J.</mods:namePart>
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<mods:nameIdentifier id="0EA05B0A735EDC56FB15A0E53A494237" type="ORCID">0000-0001-5653-224X</mods:nameIdentifier>
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<mods:affiliation id="069CF9F7BEC4B5254C9CD49B329B9DEF">Naturalis Biodiversity Center, 2333 CR, Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name id="6720427BAC4C5F0DE730238202B1FD94" type="personal">
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<mods:role id="75E6BF26A293BDA51E7A5DDEB617E4AB">
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<mods:roleTerm id="D8F32F92B432F3525DD0D2E123CE5F3C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="82E815EF445D9422736BD0F9DF6A34E2">Ismael, Halgurd R.</mods:namePart>
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<mods:affiliation id="767EDE94C560D1E1438362852717A26D">Department of Plant Protection, College of Agricultural Engineering Sciences, University of Duhok, Duhok, Kurdistan Region, Iraq</mods:affiliation>
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</mods:name>
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<mods:name id="174387A2E60B06097BC2349E22A85F74" type="personal">
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<mods:role id="84479813536B297AC9ADD2F25C355464">
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<mods:roleTerm id="C75906D5CC581D2308DA08F736EAACBF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="79E46BCCA4A32209D82845DF4D07E0AF">Baiocchi, Daniele</mods:namePart>
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<mods:affiliation id="DEB175CAFE0E8252074756100A38E828">Via Matteo Babini 26, 00139, Roma, Italy</mods:affiliation>
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</mods:name>
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<mods:name id="560729E3ED4438415842A2178BB80927" type="personal">
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<mods:role id="7FF2E1B47978534A22E76855CBFC99D8">
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<mods:roleTerm id="56B3576DE573F6AC822613D763CB9EAB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="098F4FCCF9221038D5DDE821B6FA63A8">Hamad, Mudhafar I.</mods:namePart>
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<mods:affiliation id="66979B4D12FF512D8D9FB6D0373D0F82">Khabat Technical Institute, Erbil Polytechnic University, Erbil, Kurdistan Region, Iraq</mods:affiliation>
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</mods:name>
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<mods:name id="300D3938A0D9865E6368396883C5B641" type="personal">
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<mods:role id="D08760653FDE27F9674DF6B29094A07B">
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<mods:roleTerm id="7EF312C72BEDDCAA7FA4506EACC05377">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0A1DAA23D67CB7004B2E7354330B65BC">Bapir, Tahseen T.</mods:namePart>
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<mods:affiliation id="315C7D8675BE2734A52E6896EC9E52D1">Department of Plant Protection, College of Agricultural Engineering Sciences, University of Duhok, Duhok, Kurdistan Region, Iraq</mods:affiliation>
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</mods:name>
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<mods:name id="A67F58571CBBF6CE65987569B1AD5046" type="personal">
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<mods:roleTerm id="C242576286CA33E0DB9240CE567FE530">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5BEC51190666191AE2270CCEF608152E">Selis, Marco</mods:namePart>
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<mods:nameIdentifier id="797D79BB9AB56168DB8FD16762AC7390" type="ORCID">0000-0002-8200-9441</mods:nameIdentifier>
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<mods:affiliation id="B445CDDC14C5BF95288181901053DF72">Via dei Tarquini 22, 01100 VT, Viterbo, Italy</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="38F1F7C41DB9EC889791CF97F0B6DE1B">text</mods:typeOfResource>
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<mods:titleInfo id="FB66D2F68462150549AAC94201A64D7F">
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<mods:title id="7F86B8FE8EA0C74ACBF832E9FDD1D2A3">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="C41C995EDE691068A3EDFC6E7E476063">
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<mods:date id="93D1DBBAFED10D27C4C5FAA38504D93C">2024</mods:date>
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<mods:detail id="E76A4673B7557C964CB24071355FAE2A" type="pubDate">
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<mods:number id="BFF590DE256F26D863A7507B47993842">2024-07-01</mods:number>
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</mods:detail>
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<mods:detail id="F27C3E39A4E2E6D8484DDAB51FF0B160" type="volume">
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<mods:number id="EF14350D1B5BC5002F06D91B94DFB2A5">1205</mods:number>
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</mods:detail>
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<mods:extent id="433C9FC07C2D6C6271214C0E747D6EDF" unit="page">
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<mods:start id="D5DCA482614547066C9CAC092537D60C">267</mods:start>
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<mods:end id="A1556BED66A0B93D5F1B63A8524C6EA7">298</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="FE7BBEE58DDB472078F94A4270BFF862">journal article</mods:classification>
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<mods:identifier id="69603DA34FA8EAB55F8A24870DB9DB6E" type="CLB-Dataset">299340</mods:identifier>
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<mods:identifier id="0746D2FC2941845257FAA5E2437FFE9D" type="DOI">10.3897/zookeys.1205.120033</mods:identifier>
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<mods:identifier id="1B36903864281A325E664A996680DDD8" type="GBIF-Dataset">15d96bea-75a1-4b35-a306-3aecdc1513a7</mods:identifier>
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<mods:identifier id="04F48F288EC68F677F17A7E0F819A1BE" type="ZooBank">1093570E-D1C7-4918-BD8F-0BC3D3F486E5</mods:identifier>
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</mods:mods>
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<treatment id="64180BEE5A8355D4A3089FA3C2E06A0F" ID-DOI="http://doi.org/10.5281/zenodo.12615775" ID-GBIF-Taxon="234396735" ID-Zenodo-Dep="12615775" ID-arpha="64180BEE-5A83-55D4-A308-9FA3C2E06A0F" ID-zoobank="https://zoobank.org/3C35CEE1-7063-4C23-8577-1BA21A6CCAFE" LSID="urn:lsid:zoobank.org:act:3C35CEE1-7063-4C23-8577-1BA21A6CCAFE" LSID-ZBK="urn:lsid:zoobank.org:act:3C35CEE1-7063-4C23-8577-1BA21A6CCAFE" httpUri="http://treatment.plazi.org/id/64180BEE5A8355D4A3089FA3C2E06A0F">
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||
<subSubSection id="BB6995671859C9DFA104E977F7845CF3" type="nomenclature">
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||
<paragraph id="0AA6C8214F5028E33964942D7D2E575F">
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||
<heading id="3F133AC0F59106E4E3F86E11EFCCB7F8" reason="label">20.</heading>
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||
<taxonomicName id="7E5DFF77950880DA2CA653D52DC95737" authority="Wood" authorityName="Wood & Ismael & Baiocchi & Hamad & Bapir & Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis" status="sp. nov." subGenus="Aciandrena">
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||
<emphasis id="D1255133B2BF3DD8CF90599C645FBBEC" italics="true">Andrena</emphasis>
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||
(
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||
<emphasis id="EF7AACFDE2A85F711CBA08CE24BE2079" italics="true">Aciandrena</emphasis>
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)
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<emphasis id="87C597DC5258A5E709D5CCAE1AF01FC1" italics="true">duhokensis</emphasis>
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Wood
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</taxonomicName>
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,
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<taxonomicNameLabel id="BAD01DDB0652A61ADE5D6DACE7E78281" rank="species">sp. nov.</taxonomicNameLabel>
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*
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EVGBG" type="material">
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||
<paragraph id="140AEC86B833FA08ABE15C0AEBDB8132">
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||
<heading id="A8A1A87C83F94B76DB98DCA2CB17B212" reason="title">Material examined.</heading>
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||
</paragraph>
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||
<paragraph id="00E8930BB27594B42AA3C946E2EE3D9D">
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||
<materialsCitation id="7ADB7E1F159A56B62683892C1F21D014" ID-GBIF-Occurrence="4900833338" collectingDate="2023-05-11" collectionCode="RMNH" collectorName="D. Baiocchi" country="Iraq" elevation="1912" latitude="37.0158" location="Mt. Gara" longLatPrecision="6" longitude="43.3506" specimenCount="1" specimenCount-male="1" stateProvince="Duhok" typeStatus="holotype">
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||
<emphasis id="54F8A1EA2A7308BD227EA7196BDDCEE2" bold="true">
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||
<emphasis id="F56CD63A55F7A72C3FBF9BA6EA46FBF2" bold="true" italics="true">
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||
<typeStatus id="65F42F68E17F57CABDB919CE8E32F67A" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
:
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<collectingCountry id="E56BDF9E67AAA2B70FCFFD776E0E9B12" name="Iraq">Iraq</collectingCountry>
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</emphasis>
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:
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<collectingRegion id="151A315C4B0C5A836537CFB5DA25EDE0" country="Iraq" name="Dihok">Duhok</collectingRegion>
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,
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||
<location id="726FDC1CF835EF2CEB05E61863D4A729" LSID="urn:lsid:plazi:treatment:64180BEE5A8355D4A3089FA3C2E06A0F:726FDC1CF835EF2CEB05E61863D4A729" country="Iraq" latitude="37.0158" longLatPrecision="6" longitude="43.3506" name="Mt. Gara" stateProvince="Duhok">Mt. Gara</location>
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[
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<locationDeviation id="ED46EDC7696A5620F87D2AC8024773D9" location="Sarsing">S of Sarsing</locationDeviation>
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||
],
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||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDHBG" specific-use="{"type":"Point","coordinates":[43.350600,37.015800]}">
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||
<geoCoordinate id="3E2D98E19051D15A86C5417144796126" degrees="37.0158" direction="north" orientation="latitude" precision="5" value="37.0158">37.0158 ° N</geoCoordinate>
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||
,
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||
<geoCoordinate id="AC72D885A1A115D344A2AEA5D0502537" degrees="43.3506" direction="east" orientation="longitude" precision="5" value="43.3506">43.3506 ° E</geoCoordinate>
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||
</named-content>
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||
,
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||
<quantity id="D738D1FD564538A383E2B08506575980" metricMagnitude="3" metricUnit="m" metricValue="1.9119999999999997" unit="m" value="1912.0">
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||
<elevation id="BE77B087F5CEA538596E7A413EF28646" metricMagnitude="3" metricUnit="m" metricValue="1.9119999999999997" unit="m" value="1912.0">1912 m</elevation>
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||
</quantity>
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||
,
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<date id="A36F671E113B0E81FDBD16523B54D4F8" value="2023-05-11">
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||
<collectingDate id="D8F10D0654934504D603E59CC0C05D64" value="2023-05-11">11. v. 2023</collectingDate>
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</date>
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,
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<specimenCount id="D24608BB8963B5332FCFDB1C05937910" count="1" type="male">1 ♂</specimenCount>
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, leg.
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<collectorName id="1E79EAEB1311BC86B77B09F16E5C910E">D. Baiocchi</collectorName>
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,
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||
<named-content id="A3CE293D764A6B9EA187921617BD40AF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-natural-history-naturalis" xlink_title="Naturalis Biodiversity Center, Leiden, the Netherlands">
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||
<collectionCode id="441111ABDDA8BCAEA9414453ED2C88B7" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
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</named-content>
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||
</materialsCitation>
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||
.
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<materialsCitation id="1E4E9B958F6B49E8DB6673F47F261991" ID-GBIF-Occurrence="4900833378" collectingDate="2023-05-11" collectionCode="MSVI, RMNH, TJWC, DUMAI" collectorName="D. Baiocchi" country="Iraq" elevation="1912" latitude="37.0158" location="Iraq" longLatPrecision="6" longitude="43.3506" specimenCount="7" specimenCount-male="7" stateProvince="Duhok" typeStatus="paratype">
|
||
<emphasis id="C6BB22E9B841E69C341536FAB2CA8BE5" bold="true">
|
||
<emphasis id="8BDCB9649CC07FA390AE48599CDC7C1A" bold="true" italics="true">
|
||
<typeStatus id="F6FB95B3AFB55B2D8E4A3A89C053F06E" type="paratype">Paratypes</typeStatus>
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</emphasis>
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||
:
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<collectingCountry id="D556241DF525407DB358A7EC8B41D8A2" name="Iraq">Iraq</collectingCountry>
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||
</emphasis>
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: Same information as holotype,
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<specimenCount id="6B2C7F5199C9A6C323C4DCA7166A37C7" count="7" type="male">7 ♂</specimenCount>
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||
,
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||
<abbrev content-type="institution" id="ABBRID0ERHBG" xlink_title="Personal collection of Marco Selis, Viterbo, Italy">
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<collectionCode id="D6289B393E6A5D671719A397BA3A3FF6">MSVI</collectionCode>
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</abbrev>
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||
/
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||
<named-content id="74B2CA9B16CC38C39D1AEB2B834E614F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-natural-history-naturalis" xlink_title="Naturalis Biodiversity Center, Leiden, the Netherlands">
|
||
<collectionCode id="3ABBFD4725242578D9FC0B4F1BDD348B" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
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||
</named-content>
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||
/
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||
<abbrev content-type="institution" id="ABBRID0E2HBG" xlink_title="Personal collection of Thomas J. Wood, Leiden, the Netherlands">
|
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<collectionCode id="1969C0916FD24826CCFBA8BFD914E736">TJWC</collectionCode>
|
||
</abbrev>
|
||
/
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||
<abbrev content-type="institution" id="ABBRID0EAIBG" xlink_title="University of Duhok, Museum of Agriculture College, Duhok, Iraq">
|
||
<collectionCode id="59BD481750119CE9AEBC6B1A2F0A89B5">DUMAI</collectionCode>
|
||
</abbrev>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EFIBG" type="diagnosis">
|
||
<paragraph id="E7FC112FC5A7EA66320C13EB0AE86C53">
|
||
<heading id="84E4AC04E6EA3A8239F09D7DAEE4F2E3" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="E7E267655A78F3B27C341F2E57691719">
|
||
<taxonomicName id="FC56A65DA3030319029E8755672BFA76" authorityName="Wood & Ismael & Baiocchi & Hamad & Bapir & Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis">
|
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<emphasis id="9E10CCEFD8AF06269B9365C2AAB39183" italics="true">Andrena duhokensis</emphasis>
|
||
</taxonomicName>
|
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can be recognised as part of the subgenus
|
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<taxonomicName id="1D1511589782D20DC361952BE4E4BB96" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="14BC667555CA14C2E4EA771F56BD7AE3" italics="true">Aciandrena</emphasis>
|
||
Warncke, 1968
|
||
</taxonomicName>
|
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due to the small body size, dark integument with the exception of the pale-marked clypeus (Fig.
|
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<figureCitation id="CF59674746FE9AD5B3D8FDF35F1BC08B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 A</figureCitation>
|
||
), finely shagreened propodeal triangle (Fig.
|
||
<figureCitation id="CA414881187B2557E0E011A6ACADCDA4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 D</figureCitation>
|
||
; without lateral or basal rugae), more or less impunctate terga (Fig.
|
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<figureCitation id="7BB791D67879B1B406FFCFC25A2E4B63" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 E</figureCitation>
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), strongly antefurcal nervulus, and typical simple genital capsule (Fig.
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<figureCitation id="C63F9C6870560A6446621D638C3E2CD8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 F; e</figureCitation>
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. g., without completely reduced gonocoxae as in most members of the subgenus
|
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<taxonomicName id="AB1EF55F9AA6A6DCB535B4973B95973C" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" rank="subGenus" subGenus="Graecandrena">
|
||
<emphasis id="2B986CA7DB0CD31CA37CCFDBBCE49D4B" italics="true">Graecandrena</emphasis>
|
||
Warncke, 1968
|
||
</taxonomicName>
|
||
). Identification of
|
||
<taxonomicName id="1ACB8C3B8BB06F333AD5573945AD55A1" authorityName="Warncke" authorityYear="1968" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="AC16AFDC9AE8D565BF26236FDEFB1401" italics="true">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
species is extremely challenging due to the large number of often cryptic species found in dry environments, their often very local distributions, and lack of good characters in the female sex. Fortunately, males are a little easier to identify through the examination of the genital capsule.
|
||
<taxonomicName id="72816D1E74B829FA7A3F29C7FCC7EB34" authorityName="Wood & Ismael & Baiocchi & Hamad & Bapir & Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis">
|
||
<emphasis id="E8AE63D28EA4D3EFD7B670A6097B86FF" italics="true">Andrena duhokensis</emphasis>
|
||
</taxonomicName>
|
||
can be initially recognised due to its almost entirely yellow-marked clypeus (with the exception of two small black marks [sometimes absent] and some narrow black areas at the edges of the clypeus), placing it closest to
|
||
<taxonomicName id="6694CDECA143DFE1EE46DE59EA3D426E" authority="Morawitz, 1877" authorityName="Morawitz" authorityYear="1877" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="1AD3DDBF540D188044957CBD4335850A" italics="true">A. tenuis</emphasis>
|
||
Morawitz, 1877
|
||
</taxonomicName>
|
||
which was described from the Caucasus. However,
|
||
<taxonomicName id="5C88312CCA2267D070AEF977A78FEC50" authorityName="Wood & Ismael & Baiocchi & Hamad & Bapir & Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis">
|
||
<emphasis id="C2B4944B2AA444BAFBF4E09B98C3593A" italics="true">A. duhokensis</emphasis>
|
||
</taxonomicName>
|
||
can be recognised due to the unique form of the genital capsule which has the gonocoxal teeth with their apexes strongly truncate and asymmetrical so that they diverge from each other (Fig.
|
||
<figureCitation id="7E0473549770949F39C2C997A75ADBB2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 F</figureCitation>
|
||
); in other words, they form somewhat pointed but flattened and diverging teeth. In comparative species of
|
||
<taxonomicName id="2FC47F344D0DA53F19B53D8125E5053E" authorityName="Warncke" authorityYear="1968" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="A6A1DF3580194954FF9B203D3782C2BA" italics="true">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
, the gonocoxal teeth are either strongly produced and truncate (e. g.,
|
||
<taxonomicName id="BD0D4C2034CA03193B521273DA49519B" authority="Wood 2022" authorityName="Wood" authorityYear="2022" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="deminuta">
|
||
<emphasis id="DE0CC1C76BBC25E5B5E311EBA5E8623D" italics="true">A. deminuta</emphasis>
|
||
<bibRefCitation id="1E7CFFC189CB164ECF3B345C3C529053" DOI="10.5852/ejt.2022.843.1947" author="Wood TJ & Monfared A" journalOrPublisher="European Journal of Taxonomy" pagination="1-136" refId="B41" refString="Wood TJ, Monfared A (2022) A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843. 1947" title="A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species." volume="843" year="2022">Wood 2022</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="2BCDD8F2EAD9E1E9BA1FABB9441057F5" name="Iran">Iran</collectingCountry>
|
||
), produced into narrowly pointed or more broadly rounded teeth (e. g.,
|
||
<taxonomicName id="24AE8DE89A8FCFA65CD3BF84CE56AECC" authorityName="Morawitz" authorityYear="1877" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="E8D91B739A48D3308E93F57B746504EB" italics="true">A. tenuis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="399405FE3D2308073185432919ABA30B" name="Turkey">Turkey</collectingCountry>
|
||
and the Caucasus,
|
||
<taxonomicName id="0DD565CDC38AF9E1A584BAC3612E3812" authority="Morawitz, 1886" authorityName="Morawitz" authorityYear="1886" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aciculata">
|
||
<emphasis id="28AFF22B15CF7261661A394CD940A938" italics="true">A. aciculata</emphasis>
|
||
Morawitz, 1886
|
||
</taxonomicName>
|
||
from Europe to the Caucasus), or reduced and truncate but never forming diverging teeth (e. g.,
|
||
<taxonomicName id="2278620F4C42CAF79DC2C604B97852C1" authority="Scheuchl & Pisanty, 2016" authorityName="Scheuchl & Pisanty" authorityYear="2016" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="judaea">
|
||
<emphasis id="20913C65A1DBCC9F264D47AC04E6105C" italics="true">A. judaea</emphasis>
|
||
<bibRefCitation id="4D9DFFCCA03E1A3D8FAEE16CCE304025" DOI="10.11646/zootaxa.4189.3.3" author="Pisanty G & Scheuchl E & Dorchin N" issue="3" journalOrPublisher="Zootaxa" pagination="485-515" refId="B21" refString="Pisanty G, Scheuchl E, Dorchin N (2016) Eight new species of Andrena Fabricius (Hymenoptera: Apoidea: Andrenidae) from Israel - a Mediterranean hotspot for wild bees. Zootaxa 4189 (3): 485 - 515. https: // doi. org / 10.11646 / zootaxa. 4189.3. 3" title="Eight new species of Andrena Fabricius (Hymenoptera: Apoidea: Andrenidae) from Israel - a Mediterranean hotspot for wild bees." volume="4189" year="2016">Scheuchl & Pisanty, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="63727013C3E561D6630E47A375C913C8" name="Israel">Israel</collectingCountry>
|
||
). In this context,
|
||
<taxonomicName id="EE9B9C9EDABBBEADDDC4638C1FB368D7" authorityName="Wood & Ismael & Baiocchi & Hamad & Bapir & Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis">
|
||
<emphasis id="E004B0C7AA796FD300A1BD6C29B0826C" italics="true">A. duhokensis</emphasis>
|
||
</taxonomicName>
|
||
can be separated from all similar species.
|
||
</paragraph>
|
||
<caption id="D0FA8CE8F33782DFBDEDC5D51EA331B7" ID-DOI="10.3897/zookeys.1205.120033.figure5" ID-arpha="96C67188-9C86-5C49-B2DE-977A05249501" httpUri="https://binary.pensoft.net/fig/1083013" startId="F5">
|
||
<paragraph id="834D8F7CF2A4BA8E5ED0A3C9F73EC404">
|
||
<label id="367B1D49D080EBAAE9E84A95E98281CE">Figure 5.</label>
|
||
</paragraph>
|
||
<paragraph id="FAFD3F374F1DDCAAFDC245609714D6EC">
|
||
<taxonomicName id="07D120DA8826CACA2C70C8E3673C5F4C" authorityName="Wood & Ismael & Baiocchi & Hamad & Bapir & Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis" status="sp. nov." subGenus="Aciandrena">
|
||
<emphasis id="DA0248160D1579AE4DB59A28249FEFE0" italics="true">Andrena</emphasis>
|
||
(
|
||
<emphasis id="8495E7EA28C7664D5CFAB7ACFB1B4B3E" italics="true">Aciandrena</emphasis>
|
||
)
|
||
<emphasis id="D3342A1B8264AAEFB3F3EE47735078B7" italics="true">duhokensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F9AD5D2E9AC460ACAD685D22D2C8F495" rank="species">sp. nov.</taxonomicNameLabel>
|
||
male
|
||
<emphasis id="3ECDE2227EEB5677647C30FDDD9FF218" bold="true">A</emphasis>
|
||
habitus, lateral view
|
||
<emphasis id="426CC6EC2D06AD607482042C674CB0D6" bold="true">B</emphasis>
|
||
face, frontal view
|
||
<emphasis id="28CDF0125951BC041FC1F6BC48ED7BA8" bold="true">C</emphasis>
|
||
scutum, dorsal view
|
||
<emphasis id="AC2C850F4358BD6CCAF816B6AE3A2A9D" bold="true">D</emphasis>
|
||
propodeum, dorsal view
|
||
<emphasis id="BC0EE7BB6C3564BE577625EDAAC75C3F" bold="true">E</emphasis>
|
||
terga, dorsal view
|
||
<emphasis id="FC6DF7B9BF08166E32CCBB1C52099F0E" bold="true">F</emphasis>
|
||
genital capsule, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EJOBG" type="description">
|
||
<paragraph id="2CD635EB2AE1930C40255F66AFB9451B">
|
||
<heading id="E562E88700F4505C61E9CF211FA1BFE2" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="9DC66673A52E78E99336AAA7BA160F50">
|
||
<emphasis id="31B57C859F94C5CB173FF6C81C1B7924" bold="true">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="F44579123C7D2F3786D93C231497063B">
|
||
<emphasis id="53D2D3CE4BE193277F0FA69491033C9A" bold="true">Male.</emphasis>
|
||
Body length
|
||
<quantity id="4B19AD8ACC6F12FF106F7CA5D17C9929" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
(Fig.
|
||
<figureCitation id="795C216E7784296D35264FEB3267156E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 A</figureCitation>
|
||
).
|
||
<emphasis id="2E2FCBCB35BD56F7844B045CF0249A97" bold="true" italics="true">Head</emphasis>
|
||
: Dark, 1.25 × wider than long (Fig.
|
||
<figureCitation id="069D1036022C28CB82A1F99577DCE948" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 B</figureCitation>
|
||
). Clypeus weakly domed, predominantly yellow or pale yellow marked with exception of two black dots (sometimes absent) and narrow black markings laterally; surface finely shagreened and weakly shining, irregularly punctate, punctures separated by 1–3 puncture diameters. Process of labrum small, rounded trapezoidal, 2 × wider than long, apical margin weakly emarginate. Gena subequal to diameter of compound eye; ocelloccipital distance equals diameter of lateral ocellus. Head covered with dirty white hairs, none equalling length of scape. Antennae basally dark, A 4–13 ventrally lightened by presence of grey scales; A 3 exceeding length of A 4, shorter than A 4 + 5; A 4 sub-square, shorter than long, A 5 quadrate, as long as broad.
|
||
</paragraph>
|
||
<paragraph id="8FD19EFBF074221DF2480ED42881798A">
|
||
<emphasis id="70F25E41C6098B2745B20048BB0AE4B0" bold="true" italics="true">Mesosoma</emphasis>
|
||
: Scutum and scutellum with fine granular shagreen, weakly shining, irregularly punctate, punctures separated by 1–4 puncture diameters (Fig.
|
||
<figureCitation id="2812B794CEF94E21DDFF5002480D5BBE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 C</figureCitation>
|
||
). Mesepisternum and dorsolateral parts of propodeum with regular but large microreticulation, dull to weakly shining; propodeal triangle large, indicated by change in surface sculpture, internal surface with fine granular microreticulation, in some parts with weakly raised network of carinae between microreticulation, surface dull (Fig.
|
||
<figureCitation id="AE26C1D8705FC8B6049744FF797D5082" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 D</figureCitation>
|
||
). Mesosoma with sparse short whitish hairs. Legs predominantly dark, apical tarsal segments slightly lightened orange, pubescence white. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma, and venation pale orange, nervulus very strongly antefurcal.
|
||
</paragraph>
|
||
<paragraph id="774B4FF9F2491536CCB9826A147BDF25">
|
||
<emphasis id="6F590F1A4BE1A0F28754936EB73B8E12" bold="true" italics="true">Metasoma</emphasis>
|
||
: Terga predominantly dark, tergal margins weakly but distinctly depressed, broadly lightened hyaline yellow-brown (Fig.
|
||
<figureCitation id="934ADD3678552A0C5C59266FC0D58122" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 E</figureCitation>
|
||
). Tergal discs finely microreticulate to shagreened, weakly shining, surface essentially impunctate, with scattered obscure punctures disappearing into underlying sculpture. Terga with sparse scattered short hairs, not forming hairbands. T 6 and T 7 with long light brown hairs overlying pseudopygidial plate of T 7. S 8 narrow, apex slightly broadened like a fish-tail, truncate, ventral surface with dense lateral fan of brown hairs. Genital capsule compact, gonocoxae produced into apical teeth, teeth strongly truncate, apexes angled and diverging from each other to form angular anteriorly projecting teeth (Fig.
|
||
<figureCitation id="652DF2F903460D81F1B4C261B03C07C8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 F</figureCitation>
|
||
). Gonostyli flattened and spatulate, internal margins raised, forming slight bump on inner margin basally. Penis valves moderate, occupying ½ space between gonostyli basally, strongly tapering apically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E5PBG" type="etymology">
|
||
<paragraph id="A3FE49049E56B1D5DBE3822E3DCB2759">
|
||
<heading id="142C13DCE607CEEE71E9261E3BB446F3" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="C515FE34D1BCC8B0BB7CCDDE568B3CE5">
|
||
The name is derived from the city of
|
||
<collectingRegion id="B64A90FBD251D047CE89342FA06F8E2B" country="Iraq" name="Dihok">Duhok</collectingRegion>
|
||
which gives its name to the province in which these specimens were collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EDQBG" type="distribution">
|
||
<paragraph id="F88FBCC73DF2B8EFCCCE3AD4CDFBA2E0">
|
||
<heading id="C7BFEA600DD6A4AB7CE62F2D01232B1C" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="599E7DBB4E346A0691CBB0BB134C3AD3">
|
||
<collectingCountry id="BB25C866A0F8CE811118215BABB7A8FA" name="Iraq">Iraq</collectingCountry>
|
||
(Kurdistan region).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |