146 lines
18 KiB
XML
146 lines
18 KiB
XML
<document id="4D393E2A87F85B82F2EF39C1A427EC35" ID-DOI="10.11646/zootaxa.2462.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312710" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629676798216" checkinUser="felipe" docAuthor="Hyman, Isabel T. & Ponder, Winston F." docDate="2010" docId="6413F378FFF66A68F28B709EFB56FC21" docLanguage="en" docName="zt02462p148.pdf" docOrigin="Zootaxa 2462 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Sesara infrendens" docType="treatment" docVersion="4" lastPageNumber="99" masterDocId="982A8B00FF976A0AF21C766BFFEBFF8B" masterDocTitle="A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462" masterLastPageNumber="148" masterPageNumber="1" pageNumber="98" updateTime="1699094437759" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="C760F620E592FE1F0C80E569169164D3">
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<mods:title id="CA89B5D6E97DBFB5396075FB76365C02">A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462</mods:title>
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<mods:namePart id="619C579C32D4C604384EC51DE832D993">Hyman, Isabel T.</mods:namePart>
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<mods:namePart id="5988C2CF56F4A14A29B3794CFBD7C1DA">Ponder, Winston F.</mods:namePart>
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<mods:typeOfResource id="26BBCF6A5F526E573A8A94ABFFDA75E5">text</mods:typeOfResource>
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<mods:title id="E71931D4A4A87266B307B66EE85B42A6">Zootaxa</mods:title>
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<mods:part id="9B3CD8FC94339BF3381F04FDC3642E89">
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<mods:date id="1AE07D33C4CDD80F26E64FD44686FA0E">2010</mods:date>
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<mods:number id="A65EA10D01F675783AF207118E28D55E">2010-05-14</mods:number>
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<mods:number id="5B07D5247245A25BDCB6DE8094BF4962">2462</mods:number>
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<mods:url id="E2D9A18C982AC23D074723F1B3C96058">https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1</mods:url>
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<mods:classification id="A98E2C91B7385400A92BDD88BFA20931">journal article</mods:classification>
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<mods:identifier id="FB5FE9CE07EAAA70ECBD4D2833F3A2F5" type="DOI">10.11646/zootaxa.2462.1.1</mods:identifier>
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<mods:identifier id="25FACC50B3315E5A1BAAFABBB8EB443C" type="ISSN">11755334</mods:identifier>
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<treatment id="6413F378FFF66A68F28B709EFB56FC21" ID-DOI="http://doi.org/10.5281/zenodo.5321049" ID-Zenodo-Dep="5321049" LSID="urn:lsid:plazi:treatment:6413F378FFF66A68F28B709EFB56FC21" httpUri="http://treatment.plazi.org/id/6413F378FFF66A68F28B709EFB56FC21" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98">
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<subSubSection id="A4A011E5FFF66A6BF28B709EFDCFF884" box="[151,548,1781,1807]" pageId="97" pageNumber="98" type="nomenclature">
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<paragraph id="EC05426EFFF66A6BF28B709EFDCFF884" blockId="97.[151,548,1781,1842]" box="[151,548,1781,1807]" pageId="97" pageNumber="98">
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<heading id="B74DF502FFF66A6BF28B709EFDCFF884" bold="true" box="[151,548,1781,1807]" fontSize="11" level="1" pageId="97" pageNumber="98" reason="1">
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<taxonomicName id="2BBA39EDFFF66A6BF28B709EFDCFF884" ID-CoL="6YKM4" authority="(Gould, 1844)" baseAuthorityName="Gould" baseAuthorityYear="1844" box="[151,548,1781,1807]" class="Gastropoda" family="Ariophantidae" genus="Sesara" kingdom="Animalia" order="Stylommatophora" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="infrendens">
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<emphasis id="DECE9E7CFFF66A6BF28B709EFDCFF884" bold="true" box="[151,548,1781,1807]" pageId="97" pageNumber="98">
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<emphasis id="DECE9E7CFFF66A6BF28B709EFE9AF884" bold="true" box="[151,369,1781,1807]" italics="true" pageId="97" pageNumber="98">Sesara infrendens</emphasis>
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(Gould, 1844)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A4A011E5FFF66A6BF28B7173FE9AF8B9" box="[151,369,1816,1842]" pageId="97" pageNumber="98" type="description">
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<paragraph id="EC05426EFFF66A6BF28B7173FE9AF8B9" blockId="97.[151,548,1781,1842]" box="[151,369,1816,1842]" pageId="97" pageNumber="98">
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<figureCitation id="74815EEBFFF66A6BF28B7173FEF7F8B9" box="[151,284,1816,1842]" captionStart="FIGURE 34" captionStartId="89.[151,255,1840,1864]" captionTargetBox="[289,1319,263,1822]" captionTargetId="figure-159@89.[245,1343,241,1829]" captionTargetPageId="89" captionText="FIGURE 34. Reproductive systems. A. Madagascan species MA1, AM C205304. B. Elisolimax flavescens, NMSA V9710. C. Sheldonia poeppigii, NMSA L5563. D. Macrochlamys petrosa, AM C205305. E. Sesara infrendens, BMNH 1995067. F. Parmarion martensi, BPBM 252738. Scale bars 1 mm (E), 2 mm (A), 5 mm (B–D, F). Abbreviations: alb, albumen gland; bc, bursa copulatrix; c, caecum; ep, epiphallus; fl, flagellum; fov, free oviduct; her, hermaphrodite duct; ls, lime-sac; m, muscle attaching penial sheath to epiphallus (only shown in E); ov, ovotestis; p, penis; pr, prostate; prm, penis retractor muscle; pt, penial sheath; st, stimulator; ut, uterus; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5312802" httpUri="https://zenodo.org/record/5312802/files/figure.png" pageId="97" pageNumber="98">Figure 34E</figureCitation>
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,
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<figureCitation id="74815EEBFFF66A6BF3347173FE9AF8B9" box="[296,369,1816,1842]" captionStart="FIGURE 36" captionStartId="92.[151,255,1898,1922]" captionTargetBox="[257,1329,453,1883]" captionTargetId="figure-122@92.[257,1330,453,1883]" captionTargetPageId="92" captionText="FIGURE 36. Radulae. SEM. A–C. Sheldonia poeppigii, NMSA V9710. D–F. Elisolimax flavescens, NMSA L5563. G– I. Macrochlamys petrosa, AM C205305. J–L. Sesara infrendens, BMNH 1995067. M–O. Parmarion martensi, BPBM 252738. P–R. Everettia sp., BMNH 20020188. Scale bars 10 mm (G–I), 20 mm (A–F, J–R)." figureDoi="http://doi.org/10.5281/zenodo.5312808" httpUri="https://zenodo.org/record/5312808/files/figure.png" pageId="97" pageNumber="98">36J–L</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="A4A011E5FFF66A6BF28B7109FDECF82F" pageId="97" pageNumber="98" type="materials_examined">
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<paragraph id="EC05426EFFF66A6BF28B7109FDECF82F" blockId="97.[151,1436,1890,2036]" pageId="97" pageNumber="98">
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<emphasis id="DECE9E7CFFF66A6BF28B7109FE66F8F7" bold="true" box="[151,397,1890,1916]" pageId="97" pageNumber="98">Material examined.</emphasis>
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<materialsCitation id="5CD24833FFF66A6BF38A7109FDE8F82F" collectionCode="BMNH" pageId="97" pageNumber="98" specimenCode="BMNH 1995067" specimenCount="1">
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<specimenCode id="BC1CEA15FFF66A6BF38A7109FDB5F8F7" box="[406,606,1890,1916]" collectionCode="BMNH" pageId="97" pageNumber="98">BMNH 1995067</specimenCode>
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(
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<specimenCount id="FABC89E7FFF66A6BF0737109FCF8F8F7" box="[623,787,1890,1916]" pageId="97" pageNumber="98" type="generic">one specimen</specimenCount>
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dissected, radula examined). Locality unknown. Godwin-Austen collection No. 161
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="A4A011E5FFF66A68F2DA71D9FC13FC81" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98" type="description">
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<paragraph id="EC05426EFFF66A68F2DA71D9FEB7FEA1" blockId="97.[151,1436,1890,2036]" lastBlockId="98.[151,1436,152,938]" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98">
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<emphasis id="DECE9E7CFFF66A6BF2DA71D9FEB5F847" bold="true" box="[198,350,1970,1996]" pageId="97" pageNumber="98">Description.</emphasis>
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<emphasis id="DECE9E7CFFF66A6BF37471D8FD8FF847" box="[360,612,1971,1996]" italics="true" pageId="97" pageNumber="98">External morphology</emphasis>
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: Shell 6.5 whorls, light orange-brown, spire and apex raised. Protoconch sculptured with very fine radial lirae; teleoconch with radial ribs; microradials present between ribs. Whorl profile flattened above and below evenly convex periphery. Internal walls of early whorls complete; umbilicus closed. Animal light yellowish brown. Mantle laps and lobes not examined due to extreme contraction of animal. Sole of foot tripartite; caudal apparatus as for family; caudal horn medium; caudal foss horizontal slit in tail.
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</paragraph>
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<paragraph id="EC05426EFFF56A68F2DF7753FAF1FED9" blockId="98.[151,1436,152,938]" box="[195,1306,312,338]" pageId="98" pageNumber="99">
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<emphasis id="DECE9E7CFFF56A68F2DF7753FE88FEDA" box="[195,355,312,337]" italics="true" pageId="98" pageNumber="99">Mantle cavity</emphasis>
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: As for family. Mantle with no visible minor blood vessels, pigmentation absent.
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</paragraph>
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<paragraph id="EC05426EFFF56A68F2DF770BFD29FEF1" blockId="98.[151,1436,152,938]" box="[195,706,352,378]" pageId="98" pageNumber="99">
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<emphasis id="DECE9E7CFFF56A68F2DF770BFE6CFEF2" box="[195,391,352,377]" italics="true" pageId="98" pageNumber="99">Digestive system</emphasis>
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: Oesophageal crop absent.
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</paragraph>
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<paragraph id="EC05426EFFF56A68F2DF77E3FC63FDC9" blockId="98.[151,1436,152,938]" pageId="98" pageNumber="99">
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<emphasis id="DECE9E7CFFF56A68F2DF77E3FE9AFE2A" box="[195,369,392,417]" italics="true" pageId="98" pageNumber="99">Genital system</emphasis>
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(
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<figureCitation id="74815EEBFFF56A68F39E77E3FDE6FE29" box="[386,525,392,418]" captionStart="FIGURE 34" captionStartId="89.[151,255,1840,1864]" captionTargetBox="[289,1319,263,1822]" captionTargetId="figure-159@89.[245,1343,241,1829]" captionTargetPageId="89" captionText="FIGURE 34. Reproductive systems. A. Madagascan species MA1, AM C205304. B. Elisolimax flavescens, NMSA V9710. C. Sheldonia poeppigii, NMSA L5563. D. Macrochlamys petrosa, AM C205305. E. Sesara infrendens, BMNH 1995067. F. Parmarion martensi, BPBM 252738. Scale bars 1 mm (E), 2 mm (A), 5 mm (B–D, F). Abbreviations: alb, albumen gland; bc, bursa copulatrix; c, caecum; ep, epiphallus; fl, flagellum; fov, free oviduct; her, hermaphrodite duct; ls, lime-sac; m, muscle attaching penial sheath to epiphallus (only shown in E); ov, ovotestis; p, penis; pr, prostate; prm, penis retractor muscle; pt, penial sheath; st, stimulator; ut, uterus; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5312802" httpUri="https://zenodo.org/record/5312802/files/figure.png" pageId="98" pageNumber="99">Figure 34E</figureCitation>
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): Specimen immature; reproductive tract poorly developed. Bursa copulatrix inserted on vagina. Vagina long. Stimulator absent. Penis long; internal sculpture not examined; penial diverticulum absent. Penial sheath not examined; penis retractor muscle attached to tip of epiphallic caecum. Epiphallus longer than penis. Epiphallic retractor caecum moderate in size, positioned in middle of epiphallus, doubled over. Spermatophore not present in dissected specimen.
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</paragraph>
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<paragraph id="EC05426EFFF56A68F2DF743BFC13FC81" blockId="98.[151,1436,152,938]" pageId="98" pageNumber="99">
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<taxonomicName id="2BBA39EDFFF56A68F2DF743BFEFCFDE2" box="[195,279,592,617]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" order="Porellales" pageId="98" pageNumber="99" phylum="Marchantiophyta" rank="genus">
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<emphasis id="DECE9E7CFFF56A68F2DF743BFEFCFDE2" box="[195,279,592,617]" italics="true" pageId="98" pageNumber="99">Radula</emphasis>
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</taxonomicName>
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(
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<figureCitation id="74815EEBFFF56A68F33B743BFE20FDE1" box="[295,459,592,618]" captionStart="FIGURE 36" captionStartId="92.[151,255,1898,1922]" captionTargetBox="[257,1329,453,1883]" captionTargetId="figure-122@92.[257,1330,453,1883]" captionTargetPageId="92" captionText="FIGURE 36. Radulae. SEM. A–C. Sheldonia poeppigii, NMSA V9710. D–F. Elisolimax flavescens, NMSA L5563. G– I. Macrochlamys petrosa, AM C205305. J–L. Sesara infrendens, BMNH 1995067. M–O. Parmarion martensi, BPBM 252738. P–R. Everettia sp., BMNH 20020188. Scale bars 10 mm (G–I), 20 mm (A–F, J–R)." figureDoi="http://doi.org/10.5281/zenodo.5312808" httpUri="https://zenodo.org/record/5312808/files/figure.png" pageId="98" pageNumber="99">Figure 36J–L</figureCitation>
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): Relatively long and narrow. Central tooth with small ectocones; mesocone lanceolate, longer than tooth base. Lateral and marginal tooth fields distinguishable. Lateral teeth with endocone smaller than central tooth ectocones; ectocone equal in size to those on central tooth; mesocone longer than tooth base. Marginal teeth with endocones absent; ectocones shorter and narrower than mesocone, not subdivided into extra teeth. Radular formula (39.10.1.10.39) × at least 45 rows.
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</paragraph>
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</subSubSection>
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<subSubSection id="A4A011E5FFF56A68F2DA757CFB56FC21" pageId="98" pageNumber="99" type="discussion">
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<paragraph id="EC05426EFFF56A68F2DA757CFB56FC21" blockId="98.[151,1436,152,938]" pageId="98" pageNumber="99">
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<emphasis id="DECE9E7CFFF56A68F2DA757CFED4FCBA" bold="true" box="[198,319,791,817]" pageId="98" pageNumber="99">Remarks.</emphasis>
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<taxonomicName id="2BBA39EDFFF56A68F3547573FE73FCBA" authorityName="Albers" authorityYear="1860" box="[328,408,792,817]" class="Gastropoda" family="Ariophantidae" genus="Sesara" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="genus">
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<emphasis id="DECE9E7CFFF56A68F3547573FE73FCBA" box="[328,408,792,817]" italics="true" pageId="98" pageNumber="99">Sesara</emphasis>
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</taxonomicName>
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is found in the Hindustan Peninsula,
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<collectingCountry id="94AD02FEFFF56A68F1447573FC27FCB9" box="[856,972,792,818]" name="Myanmar" pageId="98" pageNumber="99">Myanmar</collectingCountry>
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,
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<collectingCountry id="94AD02FEFFF56A68F1C57573FBAAFCB9" box="[985,1089,792,818]" name="Thailand" pageId="98" pageNumber="99">Thailand</collectingCountry>
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and
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<collectingCountry id="94AD02FEFFF56A68F6607573FB5EFCB9" box="[1148,1205,792,818]" name="Laos" pageId="98" pageNumber="99">Laos</collectingCountry>
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and contains about 20 species (
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<bibRefCitation id="882B3F9FFFF56A68F33F752BFE1AFCD1" author="Schileyko, A. A." box="[291,497,832,858]" pageId="98" pageNumber="99" refId="ref76904" refString="Schileyko, A. A. (2002 b) Treatise on Recent Terrestrial Pulmonate Molluscs, Part 9: Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica, Supplement 2, Moscow." type="book" year="2002">Schileyko 2002b</bibRefCitation>
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). It was included in
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<taxonomicName id="2BBA39EDFFF56A68F0F9752BFC2CFCD1" authorityName="Godwin-Austen" authorityYear="1888" box="[741,967,832,858]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="97" phylum="Mollusca" rank="subFamily" subFamily="Macrochlamydinae">Macrochlamydinae</taxonomicName>
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by
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<bibRefCitation id="882B3F9FFFF56A68F1EB752BFB77FCD1" author="Solem, A." box="[1015,1180,832,858]" pageId="98" pageNumber="99" pagination="1 - 110" refId="ref77598" refString="Solem, A. (1966) Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Spolia Zoologica Musei Hauniensis, 24, 1 - 110." type="journal article" year="1966">Solem (1966)</bibRefCitation>
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, in
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<taxonomicName id="2BBA39EDFFF56A68F6D5752BFA9EFCD1" authorityName="Bourguignat" authorityYear="1877" box="[1225,1397,832,858]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
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by
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<bibRefCitation id="882B3F9FFFF56A68F28B7503FEB6FC09" author="Hausdorf, B." box="[151,349,872,898]" pageId="98" pageNumber="99" pagination="35 - 66" refId="ref74906" refString="Hausdorf, B. (1998) Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora). Journal of Molluscan Studies, 64, 35 - 66." type="journal article" year="1998">Hausdorf (1998)</bibRefCitation>
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and in
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<taxonomicName id="2BBA39EDFFF56A68F3DE7503FD82FC09" baseAuthorityName="Zilch" baseAuthorityYear="1959" box="[450,617,872,898]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="96" phylum="Mollusca" rank="subFamily" subFamily="Ariophantinae">Ariophantinae</taxonomicName>
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by
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<bibRefCitation id="882B3F9FFFF56A68F0827503FC94FC09" author="Schileyko, A. A." box="[670,895,872,898]" pageId="98" pageNumber="99" refId="ref76904" refString="Schileyko, A. A. (2002 b) Treatise on Recent Terrestrial Pulmonate Molluscs, Part 9: Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica, Supplement 2, Moscow." type="book" year="2002">Schileyko (2002b)</bibRefCitation>
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. The species dissected was identified in the BMNH collections as
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<taxonomicName id="2BBA39EDFFF56A68F38075FBFCF5FC21" authority="(Gould, 1844)" baseAuthorityName="Gould" baseAuthorityYear="1844" box="[412,798,912,938]" class="Gastropoda" family="Ariophantidae" genus="Sesara" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="species" species="infrendens">
|
||
<emphasis id="DECE9E7CFFF56A68F38075FBFD86FC22" box="[412,621,912,937]" italics="true" pageId="98" pageNumber="99">Sesara infrendens</emphasis>
|
||
(Gould, 1844)
|
||
</taxonomicName>
|
||
, which was described from
|
||
<collectingCountry id="94AD02FEFFF56A68F67975FBFB53FC21" box="[1125,1208,912,938]" name="Myanmar" pageId="98" pageNumber="99">Burma</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |