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<document id="9BD628C0FD55C148801C68280328A1C3" ID-DOI="10.11646/zootaxa.2462.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5312710" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629676798216" checkinUser="felipe" docAuthor="Hyman, Isabel T. &amp; Ponder, Winston F." docDate="2010" docId="6413F378FFF56A68F28B726CFE23FA77" docLanguage="en" docName="zt02462p148.pdf" docOrigin="Zootaxa 2462 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Ostracolethinae Simroth 1901" docType="treatment" docVersion="4" lastPageNumber="99" masterDocId="982A8B00FF976A0AF21C766BFFEBFF8B" masterDocTitle="A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462" masterLastPageNumber="148" masterPageNumber="1" pageNumber="99" updateTime="1699094437759" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="88CF6289A842D496B13817177D0F8B54">A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462</mods:title>
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<mods:namePart id="BD352BBB22071BD179AC5DA5626436D2">Hyman, Isabel T.</mods:namePart>
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<mods:namePart id="4EC1AA3A9862722230B13F332D52CDB6">Ponder, Winston F.</mods:namePart>
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<subSubSection id="A4A011E5FFF56A68F28B726CFE0DFBAA" box="[151,486,1031,1057]" pageId="98" pageNumber="99" type="nomenclature">
<paragraph id="EC05426EFFF56A68F28B726CFE0DFBAA" blockId="98.[151,486,1031,1057]" box="[151,486,1031,1057]" pageId="98" pageNumber="99">
<heading id="B74DF502FFF56A68F28B726CFE0DFBAA" bold="true" box="[151,486,1031,1057]" fontSize="11" level="1" pageId="98" pageNumber="99" reason="1">
<emphasis id="DECE9E7CFFF56A68F28B726CFE0DFBAA" bold="true" box="[151,486,1031,1057]" pageId="98" pageNumber="99">
Subfamily
<taxonomicName id="2BBA39EDFFF56A68F303726CFE0DFBAA" ID-CoL="7NXX2" authorityName="Simroth" authorityYear="1901" box="[287,486,1031,1057]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="subFamily" subFamily="Ostracolethinae">Ostracolethinae</taxonomicName>
</emphasis>
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<subSubSection id="A4A011E5FFF56A68F28B7239FE23FA77" pageId="98" pageNumber="99" type="discussion">
<paragraph id="EC05426EFFF56A68F28B7239FD8EFA87" blockId="98.[151,1436,1106,1532]" pageId="98" pageNumber="99">
<emphasis id="DECE9E7CFFF56A68F28B7239FEFBFBE7" bold="true" box="[151,272,1106,1132]" pageId="98" pageNumber="99">Remarks.</emphasis>
This group has in the past been referred to as
<taxonomicName id="2BBA39EDFFF56A68F13B7239FC39FBE7" box="[807,978,1106,1132]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="subFamily" subFamily="Parmarioninae">Parmarioninae</taxonomicName>
(
<bibRefCitation id="882B3F9FFFF56A68F1FC7239FB91FBE7" author="Baker, H. B." box="[992,1146,1106,1132]" pageId="98" pageNumber="99" pagination="205 - 346" refId="ref72823" refString="Baker, H. B. (1941 a) Zonitid snails from Pacific Islands Part 3: Genera other than Microcystinae. Bernice P. Bishop Museum Bulletin, 166, 205 - 346." type="journal article" year="1941">Baker 1941a</bibRefCitation>
;
<bibRefCitation id="882B3F9FFFF56A68F69A7239FAFCFBE7" author="Solem, A." box="[1158,1303,1106,1132]" pageId="98" pageNumber="99" pagination="1 - 110" refId="ref77598" refString="Solem, A. (1966) Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Spolia Zoologica Musei Hauniensis, 24, 1 - 110." type="journal article" year="1966">Solem 1966</bibRefCitation>
). However,
<bibRefCitation id="882B3F9FFFF56A68F28B7211FEB1FB1F" author="Hausdorf, B." box="[151,346,1146,1172]" pageId="98" pageNumber="99" pagination="35 - 66" refId="ref74906" refString="Hausdorf, B. (1998) Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora). Journal of Molluscan Studies, 64, 35 - 66." type="journal article" year="1998">Hausdorf (1998)</bibRefCitation>
argued that
<taxonomicName id="2BBA39EDFFF56A68F3F17210FCDFFB1F" authority="Simroth, 1901" authorityName="Simroth" authorityYear="1901" box="[493,820,1146,1172]" class="Gastropoda" family="Ariophantidae" genus="Ostracolethe" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="genus">
<emphasis id="DECE9E7CFFF56A68F3F17210FD6FFB1F" box="[493,644,1147,1172]" italics="true" pageId="98" pageNumber="99">Ostracolethe</emphasis>
Simroth, 1901
</taxonomicName>
should be grouped with
<taxonomicName id="2BBA39EDFFF56A68F6477211FAE2FB1F" box="[1115,1289,1146,1172]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="subFamily" subFamily="Parmarioninae">Parmarioninae</taxonomicName>
, citing similarities in the reproductive system and geographical distribution, and that
<taxonomicName id="2BBA39EDFFF56A68F60F72C9FB20FB37" authorityName="Simroth" authorityYear="1901" box="[1043,1227,1186,1212]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="subFamily" subFamily="Ostracolethinae">Ostracolethinae</taxonomicName>
has priority over
<taxonomicName id="2BBA39EDFFF56A68F28B72A1FEADFB6F" box="[151,326,1226,1252]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="subFamily" subFamily="Parmarioninae">Parmarioninae</taxonomicName>
.
<bibRefCitation id="882B3F9FFFF56A68F34F72A1FDC6FB6F" author="Schileyko, A. A." box="[339,557,1226,1252]" pageId="98" pageNumber="99" refId="ref76939" refString="Schileyko, A. A. (2003 a) Treatise on Recent Terrestrial Pulmonate Molluscs, Part 10: Ariophantidae, Ostracolethidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica, Supplement 2, Moscow." type="book" year="2003">Schileyko (2003a)</bibRefCitation>
does not include
<taxonomicName id="2BBA39EDFFF56A68F11A72A0FC76FB6F" authorityName="Simroth" authorityYear="1901" box="[774,925,1227,1252]" class="Gastropoda" family="Ariophantidae" genus="Ostracolethe" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="genus">
<emphasis id="DECE9E7CFFF56A68F11A72A0FC76FB6F" box="[774,925,1227,1252]" italics="true" pageId="98" pageNumber="99">Ostracolethe</emphasis>
</taxonomicName>
in
<taxonomicName id="2BBA39EDFFF56A68F1D472A1FB98FB6F" box="[968,1139,1226,1252]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="subFamily" subFamily="Parmarioninae">Parmarioninae</taxonomicName>
and recognises
<taxonomicName id="2BBA39EDFFF56A68F72B72A1FF1FFA87" authorityName="Simroth" authorityYear="1901" class="Gastropoda" family="Ostracolethidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="family">Ostracolethidae</taxonomicName>
as a family of
<taxonomicName id="2BBA39EDFFF56A68F3BE7299FD8AFA87" authorityName="Bourguignat" authorityYear="1877" box="[418,609,1266,1292]" class="Gastropoda" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="19" phylum="Mollusca" rank="superFamily" superFamily="Helicarionoidea">Helicarionoidea</taxonomicName>
.
</paragraph>
<paragraph id="EC05426EFFF56A68F2DF7371FEFDFA27" blockId="98.[151,1436,1106,1532]" pageId="98" pageNumber="99">
<taxonomicName id="2BBA39EDFFF56A68F2DF7371FE91FABF" authorityName="Simroth" authorityYear="1901" box="[195,378,1306,1332]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="subFamily" subFamily="Ostracolethinae">Ostracolethinae</taxonomicName>
was defined by
<bibRefCitation id="882B3F9FFFF56A68F0277371FD17FABF" author="Hausdorf, B." box="[571,764,1306,1332]" pageId="98" pageNumber="99" pagination="35 - 66" refId="ref74906" refString="Hausdorf, B. (1998) Phylogeny of the Limacoidea sensu lato (Gastropoda: Stylommatophora). Journal of Molluscan Studies, 64, 35 - 66." type="journal article" year="1998">Hausdorf (1998)</bibRefCitation>
based on the highly reduced shell, presence of a stimulator with a highly calcareous dart and attachment of the penis retractor directly to the epiphallus. All three characters were observed in
<taxonomicName id="2BBA39EDFFF56A68F3A57300FC8BFA0F" authority="Simroth, 1893" authorityName="Simroth" authorityYear="1893" box="[441,864,1386,1412]" class="Gastropoda" family="Ariophantidae" genus="Parmarion" kingdom="Animalia" order="Stylommatophora" pageId="98" pageNumber="99" phylum="Mollusca" rank="species" species="martensi">
<emphasis id="DECE9E7CFFF56A68F3A57300FD41FA0F" box="[441,682,1387,1412]" italics="true" pageId="98" pageNumber="99">Parmarion martensi</emphasis>
Simroth, 1893
</taxonomicName>
. We define Ostracolthinae by the following key characters:
</paragraph>
<paragraph id="EC05426EFFF56A68F2DF73D1FE23FA77" blockId="98.[151,1436,1106,1532]" pageId="98" pageNumber="99">Shell reduced; sole furrows indistinct. Flagellum and bursa copulatrix very short; epiphallic caecum not coiled; stimulator present.</paragraph>
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