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<document id="3F18EFC897D1CCC1632F2BC0CDA46A00" ID-DOI="10.11646/zootaxa.5570.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="14745461" ID-ZooBank="41BC11D8-E0D7-40D6-837E-DF48B4A42CA5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1737968622267" checkinUser="plazi" docAuthor="Minkina, Łukasz, Skelley, Paul, Jákl, Stanislav, Král, David &amp; Li, Xiao-Feng" docDate="2025" docId="637487AB235B045F578464ADFE9CFB18" docLanguage="en" docName="zootaxa.5570.3.7.pdf" docOrigin="Zootaxa 5570 (3)" docSource="https://doi.org/10.11646/zootaxa.5570.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyparus malaitaensis Minkina, Skelley, JAkl, KrAl &amp; Li 2025, new species" docType="treatment" docVersion="3" lastPageNumber="572" masterDocId="9F4DFFD3235D04565713643AFFA6FFDC" masterDocTitle="Review of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) in the Solomon Islands with description of two new species" masterLastPageNumber="573" masterPageNumber="563" pageNumber="569" updateTime="1738230843662" updateUser="diego" zenodo-license-document="CLOSED">
<mods:mods id="7A73F7C897FDEE40D18D7FA81591758D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="69CBBE738278F13ACE8B9073DE1CE7E8">
<mods:title id="309DAE1E1E16A8CDA09BA1FDEFA8A0AC">Review of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) in the Solomon Islands with description of two new species</mods:title>
</mods:titleInfo>
<mods:name id="F612E29FF5BA036E15B0C8D3A8933CE0" type="personal">
<mods:role id="0C5E2CBAC8D2B308CCB0891F66E20618">
<mods:roleTerm id="3F9DCF06259BAA30054DBAC6CFF60FC2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="76FCDBC1F1327CEDBC2A66EB9A86BEB1">Minkina, Łukasz</mods:namePart>
<mods:affiliation id="9B4ADE2043FC48C6E58B09FF0569A051">os. Polana Szaflarska 4 / 39, 34 - 400 Nowy Targ, Poland.</mods:affiliation>
</mods:name>
<mods:name id="463E6529AAE0C518ECF104FA33EB080D" type="personal">
<mods:role id="16021921905381EC3C690F833EA962CB">
<mods:roleTerm id="A0D4400F4A4128BB726D2C14E64B9197">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ECDDF8EC0C49C7F31997F65A7B85E5D1">Skelley, Paul</mods:namePart>
<mods:affiliation id="0079007FC7E0CF1C298C236ABE338A7F">Florida State Collection of Arthropods, 1911 SW 34 th Street, Gainesville, FL, USA.</mods:affiliation>
</mods:name>
<mods:name id="979085F6D64066EA54DCB2812B893F9E" type="personal">
<mods:role id="4ADD70BB22493DB02BC09C74719A4D7E">
<mods:roleTerm id="7E137C226AB6B5E567F4C6FF84BF7F9A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3A5AB8E41F3EA1C74A0AA08D2197C76F">Jákl, Stanislav</mods:namePart>
<mods:affiliation id="B29557894D945C4BCD0110405CA3E13C">Geologická 1218 / 2 C, CZ- 152 00 Praha 5, Czechia.</mods:affiliation>
</mods:name>
<mods:name id="8F5E1A2B0D561CCD9902553E3EC7F5BD" type="personal">
<mods:role id="48AF6F44DC5CD84A0A760795542CBD3E">
<mods:roleTerm id="C3A7026666FEE01756EDAC3B814619A2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="22DF83174A3AB34AD58734F26803762E">Král, David</mods:namePart>
<mods:affiliation id="383D7D2377DC2555B4C9541E255D7D3F">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czechia.</mods:affiliation>
</mods:name>
<mods:name id="4F42612506FE1189F41FC3DA35DCDD43" type="personal">
<mods:role id="17304728659B50854B6A7ECBDDC5E781">
<mods:roleTerm id="4C96888411F589B9C75D708487F2CC9B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6218096D3D15C8894592528A2C9D1BDF">Li, Xiao-Feng</mods:namePart>
<mods:nameIdentifier id="D07EEDD3F9A871BEA78B422EF7A3F3B9" type="ORCID">0009-0009-3982-2734</mods:nameIdentifier>
<mods:nameIdentifier id="E6618086E4E9534688131ADE13F302ED" type="email">yansi3241@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="E80A46C089810D797AA9C7AAB83A1365">text</mods:typeOfResource>
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<mods:title id="D5D4AD45BDFCFD7F8690BC2F8509FE1C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FE854D361AB804F4FEB8B1E69C6A303E">
<mods:date id="321BB34063CEB73D6FCF7D6E26C5499C">2025</mods:date>
<mods:detail id="953FD864547CD4C7B1DBFDDE8F06BA94" type="pubDate">
<mods:number id="EAE5A4BA7E0159874E4B7354E79AF574">2025-01-20</mods:number>
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<mods:number id="989EA17CCD2576B43387F34903578739">5570</mods:number>
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<mods:number id="2A41C6F224294C192A276E90C19C4E62">3</mods:number>
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<mods:url id="D9065CDA745A1012F6A04B4997C296AD">https://doi.org/10.11646/zootaxa.5570.3.7</mods:url>
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<mods:classification id="3D4D40FBB232FEE2BFAD4F54F2FB87AC">journal article</mods:classification>
<mods:identifier id="BDE2A5E659DAC29BB966399C7179310B" type="DOI">10.11646/zootaxa.5570.3.7</mods:identifier>
<mods:identifier id="459BE3847BEEAE29A447EE5B7BA053FC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D4E26147E3A2421F03E5F8A03D55AAD5" type="Zenodo-Dep">14745461</mods:identifier>
<mods:identifier id="5BF0E60A6FCB9D180211CF5BAEB98813" type="ZooBank">41BC11D8-E0D7-40D6-837E-DF48B4A42CA5</mods:identifier>
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<treatment id="637487AB235B045F578464ADFE9CFB18" ID-DOI="http://doi.org/10.5281/zenodo.14745457" ID-Zenodo-Dep="14745457" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18" httpUri="http://treatment.plazi.org/id/637487AB235B045F578464ADFE9CFB18" lastPageId="9" lastPageNumber="572" pageId="6" pageNumber="569">
<subSubSection id="A3C76536235B0450578464ADFC52FF6D" box="[151,1012,150,178]" pageId="6" pageNumber="569" type="nomenclature">
<paragraph id="EB6236BD235B0450578464ADFC52FF6D" blockId="6.[151,1012,150,213]" box="[151,1012,150,178]" pageId="6" pageNumber="569">
<heading id="B02A81D1235B0450578464ADFC52FF6D" bold="true" box="[151,1012,150,178]" fontSize="11" level="1" pageId="6" pageNumber="569" reason="1">
<emphasis id="D9A9EAAF235B0450578464ADFC52FF6D" bold="true" box="[151,1012,150,178]" pageId="6" pageNumber="569">
<taxonomicName id="2CDD4D3E235B0450578464ADFCF1FF6D" authority="Minkina, Skelley, Jakl, Kral &amp; Li" authorityName="Minkina, Skelley, JAkl, KrAl &amp; Li" authorityYear="2025" box="[151,855,150,178]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="malaitaensis" status="sp. nov.">
<emphasis id="D9A9EAAF235B0450578464ADFE0AFF6E" bold="true" box="[151,428,151,178]" italics="true" pageId="6" pageNumber="569">Rhyparus malaitaensis</emphasis>
Minkina, Skelley, Jákl, Král &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D4235B0450547764ADFC52FF6D" box="[868,1012,151,177]" pageId="6" pageNumber="569" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A3C76536235B045057846481FEDDFF09" box="[151,379,187,213]" pageId="6" pageNumber="569" type="description">
<paragraph id="EB6236BD235B045057846481FEDDFF09" blockId="6.[151,1012,150,213]" box="[151,379,187,213]" pageId="6" pageNumber="569">
(
<figureCitation id="73E62A38235B0450578C6481FE82FF09" box="[159,292,187,213]" captionStart="FIGURES 3" captionStartId="7.[151,264,1850,1874]" captionTargetBox="[156,1431,186,1825]" captionTargetId="figure-21@7.[151,1435,181,1827]" captionTargetPageId="7" captionText="FIGURES 3A–C. Rhyparus malaitaensis, new species, female, holotype. A, dorsal view; B, ventral view; C, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14745469" httpUri="https://zenodo.org/record/14745469/files/figure.png" pageId="6" pageNumber="569">Figs. 3A–C</figureCitation>
,
<figureCitation id="73E62A38235B045056236481FED5FF09" box="[304,371,187,213]" captionStart="FIGURES 4" captionStartId="8.[151,264,1820,1844]" captionTargetBox="[156,1431,601,1792]" captionTargetId="figure-195@8.[151,1435,597,1797]" captionTargetPageId="8" captionText="FIGURES 4A–G. Details of morphology. A, Rhyparus guadalcanalensis, new species, male, holotype, head; B, Rhyparus guadalcanalensis, new species, male, holotype, metatibia; C, Rhyparus helophoroides Fairmaire, 1893, male, metatibia; D, Rhyparus malaitaensis, new species, female, holotype, head; E, Rhyparus malaitaensis, new species, female, holotype, pygidium and last abdominal ventrite; F, Rhyparus argopuroensis Minkina &amp; Jákl, 2024, female, holotype, pygidium and last abdominal ventrite; G, Rhyparus bacanensis Minkina, Anichtchenko, Vasiljeva &amp; Skelley, 2022, female, holotype, pygidium and last abdominal ventrite." figureDoi="http://doi.org/10.5281/zenodo.14745471" httpUri="https://zenodo.org/record/14745471/files/figure.png" pageId="6" pageNumber="569">4D–E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A3C76536235B045057846539FB57FC81" pageId="6" pageNumber="569" type="materials_examined">
<paragraph id="EB6236BD235B045057846539FB23FEC1" blockId="6.[151,1437,259,1977]" box="[151,1157,259,286]" pageId="6" pageNumber="569">
<emphasis id="D9A9EAAF235B045057846539FE90FEC1" bold="true" box="[151,310,259,285]" pageId="6" pageNumber="569">
<typeStatus id="3466881F235B045057846539FF74FEC1" box="[151,210,259,285]" pageId="6" pageNumber="569">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="93CA762D235B0450562F6539FE5CFEC1" box="[316,506,259,285]" name="Solomon Islands" pageId="6" pageNumber="569">Solomon Islands</collectingCountry>
,
<collectingRegion id="2919F85F235B045055156539FDFAFEC1" box="[518,604,259,285]" country="Solomon Islands" name="Malaita" pageId="6" pageNumber="569">Malaita</collectingRegion>
<collectingCounty id="02034E31235B045055716539FD0FFEC1" box="[610,681,259,285]" pageId="6" pageNumber="569">Island</collectingCounty>
, south coast
<collectingMunicipality id="0B06ACC7235B0450542A6539FC5DFEC2" box="[825,1019,259,286]" pageId="6" pageNumber="569">Hahorarumu Uru</collectingMunicipality>
,
<location id="EE026066235B045053156539FB23FEC1" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B045053156539FB23FEC1" box="[1030,1157,259,285]" country="Solomon Islands" county="Island" municipality="Hahorarumu Uru" name="Tribal Area" pageId="6" pageNumber="569" stateProvince="Malaita">Tribal Area</location>
</paragraph>
<paragraph id="EB6236BD235B045057D4651DFB57FC81" blockId="6.[151,1437,259,1977]" pageId="6" pageNumber="569">
<emphasis id="D9A9EAAF235B045057D4651DFED7FE9D" bold="true" box="[199,369,295,321]" pageId="6" pageNumber="569">Type material</emphasis>
(29 exx.).
<materialsCitation id="5BB53CE0235B045056E7651DFB5DFEB9" collectingDate="2017-12-07" collectingDateMax="2017-12-13" collectingDateMin="2017-12-07" collectionCode="ISEA" collectorName="S. Jakl" country="Solomon Islands" elevation="175" location="Hahorarumu Uru" pageId="6" pageNumber="569" specimenCount="1" specimenCount-female="1" stateProvince="Malaita Island" typeStatus="holotype">
<typeStatus id="3466881F235B045056E7651DFDFAFE9D" box="[500,604,295,321]" pageId="6" pageNumber="569" type="holotype">Holotype</typeStatus>
(
<specimenCount id="FDDBFD34235B0450557D651DFD67FE9D" box="[622,705,295,321]" pageId="6" pageNumber="569" type="female">female</specimenCount>
):
<collectingCountry id="93CA762D235B045055C4651DFC33FE9D" box="[727,917,295,321]" name="Solomon Islands" pageId="6" pageNumber="569">Solomon Islands</collectingCountry>
/ South Pacific,
<collectingRegion id="2919F85F235B0450534F651DFAA6FE9D" box="[1116,1280,295,321]" pageId="6" pageNumber="569">Malaita Island</collectingRegion>
/ south coast
<location id="EE026066235B045057846571FEFEFEBA" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B045057846571FEFEFEBA" box="[151,344,331,358]" country="Solomon Islands" county="South Pacific" municipality="Island" name="Hahorarumu Uru" pageId="6" pageNumber="569" stateProvince="Malaita Island">Hahorarumu Uru</location>
/
<location id="EE026066235B0450567F6571FE4DFEB9" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B0450567F6571FE4DFEB9" box="[364,491,331,357]" country="Solomon Islands" county="South Pacific" municipality="Island" name="Tribal Area" pageId="6" pageNumber="569" stateProvince="Malaita Island">Tribal Area</location>
(conservation area) /
<quantity id="2C259B58235B045055F36571FCFBFEBA" box="[736,861,331,358]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="6" pageNumber="569" unit="m" value="175.0" valueMax="250.0" valueMin="100.0">
<elevation id="60F0D18E235B045055F36571FCFBFEBA" box="[736,861,331,358]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="6" pageNumber="569" unit="m" value="175.0" valueMax="250.0" valueMin="100.0">100–250m</elevation>
</quantity>
.,
<date id="9F63107D235B0450547E6571FBA9FEB9" box="[877,1039,331,357]" pageId="6" pageNumber="569" value="2017-12-07" valueMax="2017-12-13" valueMin="2017-12-07">
<collectingDate id="8F27E995235B0450547E6571FBA9FEB9" box="[877,1039,331,357]" pageId="6" pageNumber="569" value="2017-12-07" valueMax="2017-12-13" valueMin="2017-12-07">7.–13.xii.2017</collectingDate>
</date>
/ leg.
<collectorName id="4628536B235B045053466571FB06FEB9" box="[1109,1184,331,357]" pageId="6" pageNumber="569">S. Jakl</collectorName>
(
<collectionCode id="8DCCAE78235B045053A36571FB55FEB9" box="[1200,1267,331,357]" pageId="6" pageNumber="569">ISEA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="5BB53CE0235B045052156576FCA1FE55" collectingDate="2017-12-07" collectingDateMax="2017-12-13" collectingDateMin="2017-12-07" collectionCode="SJCP, ISEA" collectorName="S. Jakl" country="Solomon Islands" elevation="175" location="Hahorarumu Uru" pageId="6" pageNumber="569" specimenCount="11" stateProvince="Malaita Island" typeStatus="paratype">
<typeStatus id="3466881F235B045052156576FAD1FEBA" box="[1286,1399,332,358]" pageId="6" pageNumber="569" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="FDDBFD34235B045052926571FF61FE56" pageId="6" pageNumber="569">11 exx</specimenCount>
.: same data as holotype (8 exx.
<collectionCode id="8DCCAE78235B045055356555FDC0FE55" box="[550,614,367,393]" pageId="6" pageNumber="569">SJCP</collectionCode>
; 3 exx.
<collectionCode id="8DCCAE78235B045055AF6555FD59FE55" box="[700,767,367,393]" pageId="6" pageNumber="569">ISEA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5BB53CE0235B045054016555FF44FE0D" collectingDate="2017-12-07" collectingDateMax="2017-12-13" collectingDateMin="2017-12-07" collectionCode="SJCP, ISEA" collectorName="S. Jakl" country="Solomon Is." elevation="175" location="Hahorarumu Uru Tribal Area" pageId="6" pageNumber="569" specimenCount="3" stateProvince="Malaita I." typeStatus="paratype">
<specimenCount id="FDDBFD34235B045054016555FCF1FE56" box="[786,855,367,394]" pageId="6" pageNumber="569">3 exx</specimenCount>
.: South Pacific,
<collectingCountry id="93CA762D235B0450531B6555FB36FE56" box="[1032,1168,367,394]" pageId="6" pageNumber="569">Solomon Is.</collectingCountry>
/
<collectingRegion id="2919F85F235B045053B66555FAB5FE56" box="[1189,1299,367,394]" pageId="6" pageNumber="569">Malaita I.</collectingRegion>
, south coast /
<location id="EE026066235B045057BB65A9FE54FE71" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B045057BB65A9FE54FE71" box="[168,498,403,430]" country="Solomon Is." municipality="South Pacific" name="Hahorarumu Uru Tribal Area" pageId="6" pageNumber="569" stateProvince="Malaita I.">Hahorarumu Uru Tribal Area</location>
/ (conservation area)
<quantity id="2C259B58235B045055FE65A9FCD6FE72" box="[749,880,403,430]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="6" pageNumber="569" unit="m" value="175.0" valueMax="250.0" valueMin="100.0">
<elevation id="60F0D18E235B045055FE65A9FCD6FE72" box="[749,880,403,430]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="6" pageNumber="569" unit="m" value="175.0" valueMax="250.0" valueMin="100.0">100–250 m</elevation>
</quantity>
. /
<date id="9F63107D235B0450549E65A9FB89FE71" box="[909,1071,403,429]" pageId="6" pageNumber="569" value="2017-12-07" valueMax="2017-12-13" valueMin="2017-12-07">
<collectingDate id="8F27E995235B0450549E65A9FB89FE71" box="[909,1071,403,429]" pageId="6" pageNumber="569" value="2017-12-07" valueMax="2017-12-13" valueMin="2017-12-07">7.–13.xii.2017</collectingDate>
</date>
leg.
<collectorName id="4628536B235B0450537965A9FB10FE71" box="[1130,1206,403,429]" pageId="6" pageNumber="569">S. Jakl</collectorName>
(2 exx.
<collectionCode id="8DCCAE78235B0450520465A9FAF1FE71" box="[1303,1367,403,429]" pageId="6" pageNumber="569">SJCP</collectionCode>
; 1 ex.
<collectionCode id="8DCCAE78235B04505784658DFF7CFE0D" box="[151,218,439,465]" pageId="6" pageNumber="569">ISEA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5BB53CE0235B045057FF658DFD96FE29" collectingDate="2016-12-01" collectingDateMax="2016-12-18" collectingDateMin="2016-12-01" collectionCode="ISEA" collectorName="St. Jakl" country="Solomon Islands" elevation="825" location="Karukiki" municipality="Honiara" pageId="6" pageNumber="569" specimenCount="1" stateProvince="Guadalcanal I." typeStatus="paratype">
<specimenCount id="FDDBFD34235B045057FF658DFEB8FE0D" box="[236,286,439,465]" count="1" pageId="6" pageNumber="569" type="generic">1ex.</specimenCount>
:
<collectingCountry id="93CA762D235B0450563B658DFE44FE0D" box="[296,482,439,465]" name="Solomon Islands" pageId="6" pageNumber="569">Solomon Islands</collectingCountry>
/ South Pacific,
<collectingRegion id="2919F85F235B04505586658DFC90FE0E" box="[661,822,439,466]" pageId="6" pageNumber="569">Guadalcanal I.</collectingRegion>
/
<quantity id="2C259B58235B0450545B658DFC6CFE0E" box="[840,970,439,466]" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="9.0" metricValueMin="7.5" pageId="6" pageNumber="569" unit="m" value="825.0" valueMax="900.0" valueMin="750.0">
<elevation id="60F0D18E235B0450545B658DFC6CFE0E" box="[840,970,439,466]" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="9.0" metricValueMin="7.5" pageId="6" pageNumber="569" unit="m" value="825.0" valueMax="900.0" valueMin="750.0">750–900 m</elevation>
</quantity>
.,
<location id="EE026066235B045054CB658DFB9AFE0D" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B045054CB658DFB9AFE0D" box="[984,1084,439,465]" country="Solomon Islands" municipality="Honiara" name="Karukiki" pageId="6" pageNumber="569" stateProvince="Guadalcanal I.">Karukiki</location>
env. /
<locationDeviation id="FAD02F9F235B04505392658DFA9AFE0D" bearing="157.5" box="[1153,1340,439,466]" distance="22500.0" pageId="6" pageNumber="569">
<quantity id="2C259B58235B04505392658DFB4DFE0E" box="[1153,1259,439,466]" metricMagnitude="4" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="6" pageNumber="569" unit="km" value="22.5" valueMax="25.0" valueMin="20.0">20–25km</quantity>
SSE of
</locationDeviation>
<collectingMunicipality id="0B06ACC7235B04505251658DFA3AFE0D" box="[1346,1436,439,465]" pageId="6" pageNumber="569">Honiara</collectingMunicipality>
/
<date id="9F63107D235B045057B665E1FEE7FE29" box="[165,321,475,501]" pageId="6" pageNumber="569" value="2016-12-01" valueMax="2016-12-18" valueMin="2016-12-01">
<collectingDate id="8F27E995235B045057B665E1FEE7FE29" box="[165,321,475,501]" pageId="6" pageNumber="569" value="2016-12-01" valueMax="2016-12-18" valueMin="2016-12-01">1–18.xii.2016</collectingDate>
</date>
/ leg.
<collectorName id="4628536B235B0450569765E1FE70FE29" box="[388,470,475,501]" pageId="6" pageNumber="569">St. Jakl</collectorName>
(
<collectionCode id="8DCCAE78235B045056F765E1FD8EFE29" box="[484,552,475,501]" pageId="6" pageNumber="569">ISEA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5BB53CE0235B0450552865E1FB24FDC5" collectingDate="2017-12-05" collectingDateMax="2017-12-11" collectingDateMin="2017-12-05" collectionCode="JHCP, ISEA, NMPC" collectorName="J. HorAk" country="Solomon Islands" location="Waisisi vill." pageId="6" pageNumber="569" specimenCount="345" specimenCount-male="340" stateProvince="Malaita Is." typeStatus="paratype">
<specimenCount id="FDDBFD34235B0450552865E1FDD9FE2A" box="[571,639,475,502]" pageId="6" pageNumber="569">5 exx</specimenCount>
.:
<collectingCountry id="93CA762D235B0450559F65E1FCE0FE29" box="[652,838,475,501]" name="Solomon Islands" pageId="6" pageNumber="569">Solomon Islands</collectingCountry>
/
<collectingRegion id="2919F85F235B0450544865E1FC74FE2A" box="[859,978,475,502]" pageId="6" pageNumber="569">Malaita Is.</collectingRegion>
Cca
<locationDeviation id="FAD02F9F235B0450531965E1FBC9FE2A" bearing="315.0" box="[1034,1135,475,502]" distance="6000.0" pageId="6" pageNumber="569">
<quantity id="2C259B58235B0450531965E1FB9DFE29" box="[1034,1083,475,501]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="569" unit="km" value="6.0">6km</quantity>
NW
</locationDeviation>
/
<location id="EE026066235B0450539065E1FAAEFE29" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B0450539065E1FAAEFE29" box="[1155,1288,475,501]" country="Solomon Islands" name="Waisisi vill." pageId="6" pageNumber="569" stateProvince="Malaita Is.">Waisisi vill.</location>
env.
<specimenCount id="FDDBFD34235B0450525265E1FA29FE2A" box="[1345,1423,475,502]" pageId="6" pageNumber="569" type="male">
<quantity id="2C259B58235B0450525265E1FA2CFE2A" box="[1345,1418,475,502]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="6" pageNumber="569" unit="m" value="340.0">340 m</quantity>
.
</specimenCount>
/ 09˚20.1’S 16˚07.6’E /
<collectorName id="4628536B235B0450569C663AFE4AFDC0" box="[399,492,512,540]" pageId="6" pageNumber="569">J. HorÁk</collectorName>
leg.
<date id="9F63107D235B0450553365C5FD64FDC6" box="[544,706,511,538]" pageId="6" pageNumber="569" value="2017-12-05" valueMax="2017-12-11" valueMin="2017-12-05">
<collectingDate id="8F27E995235B0450553365C5FD64FDC6" box="[544,706,511,538]" pageId="6" pageNumber="569" value="2017-12-05" valueMax="2017-12-11" valueMin="2017-12-05">5.–11.xii.2017</collectingDate>
</date>
(3 exx.
<collectionCode id="8DCCAE78235B0450540B65C5FCFBFDC5" box="[792,861,511,537]" pageId="6" pageNumber="569">JHCP</collectionCode>
; 1 ex.
<collectionCode id="8DCCAE78235B045054B265C5FC44FDC5" box="[929,994,511,537]" pageId="6" pageNumber="569">ISEA</collectionCode>
, 1 ex.
<collectionCode id="8DCCAE78235B0450533565C5FBDDFDC5" box="[1062,1147,511,537]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="6" pageNumber="569">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5BB53CE0235B0450539F65C5FC60FDE1" collectingDate="1963-12" collectingMethod="light" collectionCode="BPBM" country="Solomon Is." elevation="10" location="Ulo Crater" municipality="Vella Lavella" pageId="6" pageNumber="569" specimenCount="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="FDDBFD34235B0450539F65C5FB1DFDC5" box="[1164,1211,511,537]" count="1" pageId="6" pageNumber="569" type="generic">1ex.</specimenCount>
(
<specimenCount id="FDDBFD34235B045053DA65C5FAA0FDC5" box="[1225,1286,511,537]" pageId="6" pageNumber="569" type="male">male</specimenCount>
):
<collectingCountry id="93CA762D235B0450520465C5FA3AFDC6" box="[1303,1436,511,538]" pageId="6" pageNumber="569">Solomon Is.</collectingCountry>
/
<collectingMunicipality id="0B06ACC7235B045057B66619FE9EFDE1" box="[165,312,547,573]" pageId="6" pageNumber="569">Vella Lavella</collectingMunicipality>
/
<location id="EE026066235B0450565E6619FE63FDE1" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B0450565E6619FE63FDE1" box="[333,453,547,573]" country="Solomon Is." county="Solomon Is." municipality="Vella Lavella" name="Ulo Crater" pageId="6" pageNumber="569" stateProvince="Malaita">Ulo Crater</location>
,
<quantity id="2C259B58235B045056DC6619FDA6FDE1" box="[463,512,547,573]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="569" unit="m" value="10.0">
<elevation id="60F0D18E235B045056DC6619FDA6FDE1" box="[463,512,547,573]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="569" unit="m" value="10.0">10m</elevation>
</quantity>
/
<date id="9F63107D235B04505506661EFDDCFDE1" box="[533,634,547,574]" pageId="6" pageNumber="569" value="1963-12">
<collectingDate id="8F27E995235B04505506661EFDDCFDE1" box="[533,634,547,574]" pageId="6" pageNumber="569" value="1963-12">XII.1963</collectingDate>
</date>
// at kerosene /
<collectingMethod id="329C4EAA235B0450543A6619FCFAFDE1" box="[809,860,547,573]" pageId="6" pageNumber="569">light</collectingMethod>
(
<collectionCode id="8DCCAE78235B045054796619FC18FDE1" box="[874,958,547,573]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="6" pageNumber="569" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5BB53CE0235B045054C26619FB3CFDBD" collectingDate="1964-03-14" collectingMethod="light trap" collectionCode="BPBM" collectorName="P. Shanahan &amp; Bishop" country="Solomon Is." elevation="80" location="Kitipi R." pageId="6" pageNumber="569" specimenCount="2" specimenCount-female="1" stateProvince="Choiseul I." typeStatus="paratype">
<specimenCount id="FDDBFD34235B045054C26619FBA1FDE2" box="[977,1031,547,574]" count="1" pageId="6" pageNumber="569" type="generic">1 ex.</specimenCount>
(
<specimenCount id="FDDBFD34235B045053076619FBC1FDE1" box="[1044,1127,547,573]" pageId="6" pageNumber="569" type="female">female</specimenCount>
):
<collectingCountry id="93CA762D235B0450536A6619FAA6FDE2" box="[1145,1280,547,574]" pageId="6" pageNumber="569">Solomon Is.</collectingCountry>
/
<collectingRegion id="2919F85F235B045052076619FA28FDE2" box="[1300,1422,547,574]" pageId="6" pageNumber="569">Choiseul I.</collectingRegion>
/
<location id="EE026066235B04505784667DFF51FDBE" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B04505784667DFF51FDBE" box="[151,247,583,610]" country="Solomon Is." name="Kitipi R." pageId="6" pageNumber="569" stateProvince="Choiseul I.">Kitipi R.</location>
,
<quantity id="2C259B58235B04505616667DFE91FDBD" box="[261,311,583,609]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="569" unit="m" value="80.0">
<elevation id="60F0D18E235B04505616667DFE91FDBD" box="[261,311,583,609]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="569" unit="m" value="80.0">80m</elevation>
</quantity>
/
<date id="9F63107D235B04505658667DFE6FFDBD" box="[331,457,583,610]" pageId="6" pageNumber="569" value="1964-03-14">
<collectingDate id="8F27E995235B04505658667DFE6FFDBD" box="[331,457,583,610]" pageId="6" pageNumber="569" value="1964-03-14">14.III.1964</collectingDate>
</date>
//
<collectorName id="4628536B235B045056F66672FDCDFDBD" box="[485,619,583,610]" pageId="6" pageNumber="569">P. Shanahan</collectorName>
/ collector /
<collectorName id="4628536B235B045055E6667DFCE2FDBD" box="[757,836,583,609]" pageId="6" pageNumber="569">Bishop</collectorName>
//
<collectingMethod id="329C4EAA235B04505473667DFC63FDBE" box="[864,965,583,610]" pageId="6" pageNumber="569">light trap</collectingMethod>
// Bishop (
<collectionCode id="8DCCAE78235B0450532D667DFB34FDBD" box="[1086,1170,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="6" pageNumber="569" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5BB53CE0235B045053B7667DFAC5FD59" collectingDate="1960-08-07" collectingMethod="light trap" collectionCode="BPBM" collectorName="C. W. O'Brien &amp; Collector" country="Solomon Is." location="Wugiroba" municipality="San Cristoval" pageId="6" pageNumber="569" specimenCount="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="FDDBFD34235B045053B7667DFB7CFDBE" box="[1188,1242,583,610]" count="1" pageId="6" pageNumber="569" type="generic">1 ex.</specimenCount>
(
<specimenCount id="FDDBFD34235B045053FB667DFA83FDBD" box="[1256,1317,583,609]" pageId="6" pageNumber="569" type="male">male</specimenCount>
):
<collectingCountry id="93CA762D235B04505224667DFF14FD5A" pageId="6" pageNumber="569">Solomon Is.</collectingCountry>
/
<collectingMunicipality id="0B06ACC7235B045057DC6651FECCFD59" box="[207,362,619,645]" pageId="6" pageNumber="569">San Cristoval</collectingMunicipality>
[Cristobal] /
<location id="EE026066235B0450551A6651FDDEFD59" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B0450551A6651FDDEFD59" box="[521,632,619,645]" country="Solomon Is." county="Solomon Is." municipality="San Cristoval" name="Wugiroba" pageId="6" pageNumber="569" stateProvince="Choiseul">Wugiroba</location>
/
<date id="9F63107D235B045055866656FCBEFD59" box="[661,792,619,646]" pageId="6" pageNumber="569" value="1960-08-07">
<collectingDate id="8F27E995235B045055866656FCBEFD59" box="[661,792,619,646]" pageId="6" pageNumber="569" value="1960-08-07">7.VIII.1960</collectingDate>
</date>
//
<collectingMethod id="329C4EAA235B0450542E6651FC01FD5A" box="[829,935,619,646]" pageId="6" pageNumber="569">light trap</collectingMethod>
//
<collectorName id="4628536B235B045054D86651FBD5FD59" box="[971,1139,619,646]" pageId="6" pageNumber="569">C. W. O’Brien</collectorName>
/
<collectorName id="4628536B235B045053836651FB5EFD59" box="[1168,1272,619,645]" pageId="6" pageNumber="569">Collector</collectorName>
(
<collectionCode id="8DCCAE78235B045052196651FAFDFD59" box="[1290,1371,619,645]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="6" pageNumber="569" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
<materialsCitation id="5BB53CE0235B0450527E6651FF55FD11" collectingDate="1960-12" collectionCode="FSCA" collectorName="W. W. Brandt &amp; Bishop" country="Solomon Is." elevation="150" location="Kukugai Vill." pageId="6" pageNumber="569" specimenCount="2" specimenCount-female="1" stateProvince="Bougainville" typeStatus="paratype">
<specimenCount id="FDDBFD34235B0450527E6651FA3AFD59" box="[1389,1436,619,645]" count="1" pageId="6" pageNumber="569" type="generic">1ex.</specimenCount>
(
<specimenCount id="FDDBFD34235B0450578C66B5FF54FD75" box="[159,242,655,681]" pageId="6" pageNumber="569" type="female">female</specimenCount>
): [PNG]
<collectingCountry id="93CA762D235B0450564466B5FE79FD76" box="[343,479,655,682]" pageId="6" pageNumber="569">Solomon Is.</collectingCountry>
/
<collectingRegion id="2919F85F235B045056E566B5FD21FD75" box="[502,647,655,681]" country="Papua New Guinea" name="Bougainville" pageId="6" pageNumber="569">Bougainville</collectingRegion>
/
<location id="EE026066235B0450558E66B5FC90FD75" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B0450558E66B5FC90FD75" box="[669,822,655,681]" country="Solomon Is." municipality="Solomon Is." name="Kukugai Vill." pageId="6" pageNumber="569" stateProvince="Bougainville">Kukugai Vill.</location>
/
<quantity id="2C259B58235B0450545F66B5FC30FD76" box="[844,918,655,682]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="569" unit="m" value="150.0">
<elevation id="60F0D18E235B0450545F66B5FC30FD76" box="[844,918,655,682]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="569" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="9F63107D235B045054B166AAFBA1FD75" box="[930,1031,655,682]" pageId="6" pageNumber="569" value="1960-12">
<collectingDate id="8F27E995235B045054B166AAFBA1FD75" box="[930,1031,655,682]" pageId="6" pageNumber="569" value="1960-12">XII.1960</collectingDate>
</date>
//
<collectorName id="4628536B235B0450533566AAFB11FD75" box="[1062,1207,655,682]" pageId="6" pageNumber="569">W.W. Brandt</collectorName>
/ Collector /
<collectorName id="4628536B235B0450525E66B5FA3AFD75" box="[1357,1436,655,681]" pageId="6" pageNumber="569">Bishop</collectorName>
(
<collectionCode id="8DCCAE78235B0450578C6689FF4DFD11" box="[159,235,691,717]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="6" pageNumber="569" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5BB53CE0235B045056136689FA27FD11" box="[256,1409,691,718]" collectingDate="1968-02-27" collectionCode="BPBM" collectorName="Tawi &amp; Bishop" elevation="200" location="Trapanas" municipality="Trapanas" pageId="6" pageNumber="569" specimenCount="2" specimenCount-male="1" stateProvince="Bougainville" typeStatus="paratype">
<specimenCount id="FDDBFD34235B045056136689FE9EFD12" box="[256,312,691,718]" count="1" pageId="6" pageNumber="569" type="generic">1 ex.</specimenCount>
(
<specimenCount id="FDDBFD34235B0450565A6689FE20FD11" box="[329,390,691,717]" pageId="6" pageNumber="569" type="male">male</specimenCount>
): [PNG]
<collectingRegion id="2919F85F235B045056FE6689FDD8FD11" box="[493,638,691,717]" country="Papua New Guinea" name="Bougainville" pageId="6" pageNumber="569">Bougainville</collectingRegion>
/
<collectingMunicipality id="0B06ACC7235B04505585668EFD59FD12" box="[662,767,692,718]" country="Papua New Guinea" name="Trapanas" pageId="6" pageNumber="569" stateProvince="Bougainville">Trapanas</collectingMunicipality>
,
<quantity id="2C259B58235B045054186689FCEDFD11" box="[779,843,691,717]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="569" unit="m" value="200.0">
<elevation id="60F0D18E235B045054186689FCEDFD11" box="[779,843,691,717]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="569" unit="m" value="200.0">200m</elevation>
</quantity>
/
<date id="9F63107D235B045054776689FC72FD11" box="[868,980,691,717]" pageId="6" pageNumber="569" value="1968-02-27">
<collectingDate id="8F27E995235B045054776689FC72FD11" box="[868,980,691,717]" pageId="6" pageNumber="569" value="1968-02-27">27.2.1968</collectingDate>
</date>
//
<collectorName id="4628536B235B045054E76689FB8AFD11" box="[1012,1068,691,717]" pageId="6" pageNumber="569">Tawi</collectorName>
/ Collector /
<collectorName id="4628536B235B045053D66689FAB2FD11" box="[1221,1300,691,717]" pageId="6" pageNumber="569">Bishop</collectorName>
(
<collectionCode id="8DCCAE78235B045052366689FADFFD11" box="[1317,1401,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="6" pageNumber="569" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5BB53CE0235B0450529D6689FB37FCC9" collectingDate="2023-05-09" collectionCode="NHMUK" collectorName="D. Telnov" country="Solomon Islands" elevation="30" location="Puatanarau" municipality="Is. Marau area" pageId="6" pageNumber="569" specimenCode="BMNH{E}2023-95" specimenCount="2" stateProvince="Guadalcanal" typeStatus="paratype">
<specimenCount id="FDDBFD34235B0450529D6689FF6DFD2E" pageId="6" pageNumber="569">2 exx.</specimenCount>
:
<collectingCountry id="93CA762D235B045057C566EDFE33FD2D" box="[214,405,727,753]" name="Solomon Islands" pageId="6" pageNumber="569">Solomon Islands</collectingCountry>
,
<collectingRegion id="2919F85F235B045056B266EDFD8BFD2D" box="[417,557,727,753]" country="Solomon Islands" name="Guadalcanal" pageId="6" pageNumber="569">Guadalcanal</collectingRegion>
/
<collectingMunicipality id="0B06ACC7235B0450555766E2FD4FFD2E" box="[580,745,728,754]" pageId="6" pageNumber="569">Is. Marau area</collectingMunicipality>
,
<location id="EE026066235B045055E666E2FCD7FD2E" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B045055E666E2FCD7FD2E" box="[757,881,728,754]" country="Solomon Islands" municipality="Is. Marau area" name="Puatanarau" pageId="6" pageNumber="569" stateProvince="Guadalcanal">Puatanarau</location>
vill. env., / 9˚45’44’’S 160˚46’44’’E,
<date id="9F63107D235B0450520C66EDFA2BFD2D" box="[1311,1421,727,753]" pageId="6" pageNumber="569" value="2023-05-09">
<collectingDate id="8F27E995235B0450520C66EDFA2BFD2D" box="[1311,1421,727,753]" pageId="6" pageNumber="569" value="2023-05-09">09.v.2023</collectingDate>
</date>
/
<quantity id="2C259B58235B0450578466C1FF5AFCCA" box="[151,252,763,790]" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="6" pageNumber="569" unit="m" value="30.0" valueMax="40.0" valueMin="20.0">
<elevation id="60F0D18E235B0450578466C1FF5AFCCA" box="[151,252,763,790]" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="6" pageNumber="569" unit="m" value="30.0" valueMax="40.0" valueMin="20.0">20–40 m</elevation>
</quantity>
, mangroves &amp;
<taxonomicName id="2CDD4D3E235B045056B566C1FDB0FCC9" box="[422,534,763,789]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="569" phylum="Tracheophyta" rank="genus">
<emphasis id="D9A9EAAF235B045056B566C1FDB0FCC9" box="[422,534,763,789]" italics="true" pageId="6" pageNumber="569">Pandanus</emphasis>
</taxonomicName>
/ forest, leg.
<collectorName id="4628536B235B045055BA66C6FCB1FCC9" box="[681,791,763,790]" pageId="6" pageNumber="569">D. Telnov</collectorName>
/
<specimenCode id="BB7B9EC6235B0450543E66C1FBACFCCA" box="[813,1034,763,790]" pageId="6" pageNumber="569">BMNH{E}2023-95</specimenCode>
(
<collectionCode id="8DCCAE78235B0450530966C1FB2FFCC9" box="[1050,1161,763,789]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="569" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5BB53CE0235B0450538F66C1FB4CFC81" collectingDate="2023-04-20" collectingDateMax="2023-04-23" collectingDateMin="2023-04-20" collectionCode="NHMUK" collectorName="D. Telnov" country="Solomon Islands" elevation="350" location="Barana vill." municipality="Honiara" pageId="6" pageNumber="569" specimenCode="BMNH{E}2023-95" specimenCount="1" stateProvince="Guadalcanal" typeStatus="paratype">
<specimenCount id="FDDBFD34235B0450538F66C1FB68FCC9" box="[1180,1230,763,789]" count="1" pageId="6" pageNumber="569" type="generic">1ex.</specimenCount>
:
<collectingCountry id="93CA762D235B045053C966C1FA3EFCC9" box="[1242,1432,763,789]" name="Solomon Islands" pageId="6" pageNumber="569">Solomon Islands</collectingCountry>
,
<collectingRegion id="2919F85F235B045057846725FE85FCE5" box="[151,291,799,825]" country="Solomon Islands" name="Guadalcanal" pageId="6" pageNumber="569">Guadalcanal</collectingRegion>
/ Is.,
<collectingMunicipality id="0B06ACC7235B0450564D6725FE1EFCE5" box="[350,440,799,825]" pageId="6" pageNumber="569">Honiara</collectingMunicipality>
S env.,
<location id="EE026066235B0450551D671AFD36FCE5" LSID="urn:lsid:plazi:treatment:637487AB235B045F578464ADFE9CFB18:EE026066235B0450551D671AFD36FCE5" box="[526,656,799,826]" country="Solomon Islands" municipality="Honiara" name="Barana vill." pageId="6" pageNumber="569" stateProvince="Guadalcanal">Barana vill.</location>
S / env.,
<quantity id="2C259B58235B045055E16725FC8CFCE5" box="[754,810,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="569" unit="km" value="1.0">
<locationDeviation id="FAD02F9F235B045055E16725FC8CFCE5" box="[754,810,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="569" unit="km" value="1.0">1 km</locationDeviation>
</quantity>
around 9˚29’5’’S / 159˚58’40’’E,
<date id="9F63107D235B045053B76725FAEAFCE5" box="[1188,1356,799,825]" pageId="6" pageNumber="569" value="2023-04-20" valueMax="2023-04-23" valueMin="2023-04-20">
<collectingDate id="8F27E995235B045053B76725FAEAFCE5" box="[1188,1356,799,825]" pageId="6" pageNumber="569" value="2023-04-20" valueMax="2023-04-23" valueMin="2023-04-20">20 –23.iv.2023</collectingDate>
</date>
, 220– /
<quantity id="2C259B58235B045057846779FF7CFC82" box="[151,218,835,862]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="569" unit="m" value="350.0">
<elevation id="60F0D18E235B045057846779FF7CFC82" box="[151,218,835,862]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="569" unit="m" value="350.0">350m</elevation>
</quantity>
, disturbed lowland rainforest / on limestone, leg.
<collectorName id="4628536B235B04505416677EFCD5FC81" box="[773,883,835,862]" pageId="6" pageNumber="569">D. Telnov</collectorName>
/
<specimenCode id="BB7B9EC6235B0450549A6779FBC0FC82" box="[905,1126,835,862]" pageId="6" pageNumber="569">BMNH{E}2023-95</specimenCode>
(
<collectionCode id="8DCCAE78235B045053646779FB45FC81" box="[1143,1251,835,861]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="569" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A3C76536235B045E57D4675DFAB0FE29" lastPageId="8" lastPageNumber="571" pageId="6" pageNumber="569" type="description">
<paragraph id="EB6236BD235B045057D4675DFB7FFC31" blockId="6.[151,1437,259,1977]" pageId="6" pageNumber="569">
<emphasis id="D9A9EAAF235B045057D4675DFD22FC5D" bold="true" box="[199,644,871,897]" pageId="6" pageNumber="569">
Description of the
<typeStatus id="3466881F235B045056B8675DFDB4FC5D" box="[427,530,871,897]" pageId="6" pageNumber="569" type="holotype">holotype</typeStatus>
(female).
</emphasis>
Dorsum (
<figureCitation id="73E62A38235B045055EA675DFCF0FC5D" box="[761,854,871,897]" captionStart="FIGURES 3" captionStartId="7.[151,264,1850,1874]" captionTargetBox="[156,1431,186,1825]" captionTargetId="figure-21@7.[151,1435,181,1827]" captionTargetPageId="7" captionText="FIGURES 3A–C. Rhyparus malaitaensis, new species, female, holotype. A, dorsal view; B, ventral view; C, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14745469" httpUri="https://zenodo.org/record/14745469/files/figure.png" pageId="6" pageNumber="569">Fig. 3A</figureCitation>
). Body length
<quantity id="2C259B58235B04505311675DFBFBFC5E" box="[1026,1117,871,898]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="569" unit="mm" value="7.8">7.8 mm</quantity>
; maximum width
<quantity id="2C259B58235B0450523C675DFA3FFC5E" box="[1327,1433,871,898]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="6" pageNumber="569" unit="mm" value="2.45">2.45 mm</quantity>
. Body large-sized for the genus, distinctly elongate, not distinctly convex, flattened medially; matt; apparently almost glabrous, except head and longitudinal costae on pronotum and eltyra partly clothed with very short and thin, yellowish macrosetae. Dark brownish; antennae, tarsomeres and mouth parts pale brownish.
</paragraph>
<paragraph id="EB6236BD235B045057D467CDFB61FB1A" blockId="6.[151,1437,259,1977]" pageId="6" pageNumber="569">
Head (
<figureCitation id="73E62A38235B0450560367CDFEC1FBCD" box="[272,359,1015,1041]" captionStart="FIGURES 4" captionStartId="8.[151,264,1820,1844]" captionTargetBox="[156,1431,601,1792]" captionTargetId="figure-195@8.[151,1435,597,1797]" captionTargetPageId="8" captionText="FIGURES 4A–G. Details of morphology. A, Rhyparus guadalcanalensis, new species, male, holotype, head; B, Rhyparus guadalcanalensis, new species, male, holotype, metatibia; C, Rhyparus helophoroides Fairmaire, 1893, male, metatibia; D, Rhyparus malaitaensis, new species, female, holotype, head; E, Rhyparus malaitaensis, new species, female, holotype, pygidium and last abdominal ventrite; F, Rhyparus argopuroensis Minkina &amp; Jákl, 2024, female, holotype, pygidium and last abdominal ventrite; G, Rhyparus bacanensis Minkina, Anichtchenko, Vasiljeva &amp; Skelley, 2022, female, holotype, pygidium and last abdominal ventrite." figureDoi="http://doi.org/10.5281/zenodo.14745471" httpUri="https://zenodo.org/record/14745471/files/figure.png" pageId="6" pageNumber="569">Fig. 4D</figureCitation>
) transversely sub-hexagonal; surface very weakly shiny, tops of ridges moderately shiny; clypeus trapezoidal in outline, anteriorly truncate, on each side upturned as obtuse, weak tooth, laterally sinuous; genae distinctly more excavate than eyes; clypeal disc distinctly convex, ringed by a deep groove; convexity with a pair of quite distinct, very short, ridges that converge anteriorly, nearly whole surface with distinct, fine punctures bearing small macrosetae. Frons with four distinct, longitudinal ridges with similar structure as ridges on clypeal convexity. Head covered by regularly spaced, dense, moderately large punctures bearing short macrosetae.
</paragraph>
<paragraph id="EB6236BD235B045057D460EAFE53FA39" blockId="6.[151,1437,259,1977]" pageId="6" pageNumber="569">Pronotum very weakly shiny, tops of costae distinctly shiny; with eight distinct costae and seven intercostae, lateral margin with two rounded lobes. Anterior lobe very weakly more prominent laterally and distinctly narrower than median lobe, anterior lobes are the widest part of pronotum. Costae of middle, third and fourth pair entire from posterior to anterior margins; median pair very gently convergent anteriorly, distinctly convergent in the middle of anterior third; second pair of costae distinctly interrupted in anterior half; all costae on each side with very small punctures bearing very small macrosetae. All intercostae anteriorly without additional short costae. Median intercostae with dense punctures, distinctly concentrated medially into a longitudinal line, all intercostae posteriorly with distinct, dense punctation.</paragraph>
<paragraph id="EB6236BD235B045057D461D5FD95F9D5" blockId="6.[151,1437,259,1977]" box="[199,563,1519,1545]" pageId="6" pageNumber="569">Scutellum almost imperceptible.</paragraph>
<paragraph id="EB6236BD235B045057D46229FD4FF8F5" blockId="6.[151,1437,259,1977]" pageId="6" pageNumber="569">Elytra matt, tops of costae and preapical glandular area distinctly shiny. Each elytron with six elevated, distinctly convex costae, and five flat intercostae. Costae on each side with very small punctures bearing very small setae. Intercostae 1–4 with two distinct rows of punctures, medially smooth; basally third intercostae with short costal remnant and additional median row of small sparse punctures extending to apical third of elytra; humeral region of fourth intercosta with dense, irregularly spaced additional punctures; fifth intercosta with one row of punctures. Preapical glandular area relatively small. External caudal bulb distinctly reduced, area between external caudal bulb and sides of elytra not divided; external and mediointernal caudal bulbs not divided; medio-internal caudal bulb shortened, transversely rounded, internally stretched.</paragraph>
<paragraph id="EB6236BD235B045057E4630EFE3CF892" blockId="6.[151,1437,259,1977]" box="[247,410,1844,1870]" pageId="6" pageNumber="569">Macropterous.</paragraph>
<paragraph id="EB6236BD235B045057D4636DFEA0F865" blockId="6.[151,1437,259,1977]" pageId="6" pageNumber="569">
Pygidium (
<figureCitation id="73E62A38235B04505651636DFE30F8AD" box="[322,406,1879,1905]" captionStart="FIGURES 4" captionStartId="8.[151,264,1820,1844]" captionTargetBox="[156,1431,601,1792]" captionTargetId="figure-195@8.[151,1435,597,1797]" captionTargetPageId="8" captionText="FIGURES 4A–G. Details of morphology. A, Rhyparus guadalcanalensis, new species, male, holotype, head; B, Rhyparus guadalcanalensis, new species, male, holotype, metatibia; C, Rhyparus helophoroides Fairmaire, 1893, male, metatibia; D, Rhyparus malaitaensis, new species, female, holotype, head; E, Rhyparus malaitaensis, new species, female, holotype, pygidium and last abdominal ventrite; F, Rhyparus argopuroensis Minkina &amp; Jákl, 2024, female, holotype, pygidium and last abdominal ventrite; G, Rhyparus bacanensis Minkina, Anichtchenko, Vasiljeva &amp; Skelley, 2022, female, holotype, pygidium and last abdominal ventrite." figureDoi="http://doi.org/10.5281/zenodo.14745471" httpUri="https://zenodo.org/record/14745471/files/figure.png" pageId="6" pageNumber="569">Fig. 4E</figureCitation>
) triangular with acute apex, distinctly bent inwards in lateral view (not perpendicular to body axis), with dense, irregularly spaced punctation, with weak longitudinal carina in the middle and deep excision on each side.
</paragraph>
<caption id="BFA26635235A045157846300FAC8F88F" ID-DOI="http://doi.org/10.5281/zenodo.14745469" ID-Zenodo-Dep="14745469" box="[151,1390,1850,1875]" httpUri="https://zenodo.org/record/14745469/files/figure.png" pageId="7" pageNumber="570" startId="7.[151,264,1850,1874]" targetBox="[156,1431,186,1825]" targetPageId="7" targetType="figure">
<paragraph id="EB6236BD235A045157846300FAC8F88F" blockId="7.[151,1390,1850,1875]" box="[151,1390,1850,1875]" pageId="7" pageNumber="570">
<emphasis id="D9A9EAAF235A045157846300FEF5F88F" bold="true" box="[151,339,1850,1875]" pageId="7" pageNumber="570">FIGURES 3A–C.</emphasis>
<taxonomicName id="2CDD4D3E235A0451564A6301FDE6F88F" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[345,576,1851,1875]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="570" phylum="Arthropoda" rank="species" species="malaitaensis" status="sp. nov.">
<emphasis id="D9A9EAAF235A0451564A6301FDE6F88F" box="[345,576,1851,1875]" italics="true" pageId="7" pageNumber="570">Rhyparus malaitaensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D4235A0451555E6301FD6FF88F" box="[589,713,1851,1875]" pageId="7" pageNumber="570" rank="species">new species</taxonomicNameLabel>
, female, holotype. A, dorsal view; B, ventral view; C, lateral view.
</paragraph>
</caption>
<paragraph id="EB6236BD235A045E57D4639CFE1AFE55" blockId="7.[151,1436,1957,2020]" lastBlockId="8.[151,1437,151,573]" lastPageId="8" lastPageNumber="571" pageId="7" pageNumber="570">
Venter (
<figureCitation id="73E62A38235A04515633639FFED1F863" box="[288,375,1957,1983]" captionStart="FIGURES 3" captionStartId="7.[151,264,1850,1874]" captionTargetBox="[156,1431,186,1825]" captionTargetId="figure-21@7.[151,1435,181,1827]" captionTargetPageId="7" captionText="FIGURES 3A–C. Rhyparus malaitaensis, new species, female, holotype. A, dorsal view; B, ventral view; C, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14745469" httpUri="https://zenodo.org/record/14745469/files/figure.png" pageId="7" pageNumber="570">Fig. 3B</figureCitation>
) moderately shiny. Metaventral plate flattened in the middle; with distinct, narrow, quite deep longitudinal furrow on posterior half; punctation of plate dense, quite regularly spaced, slightly irregular in size; all punctures bearing short macrosetae. Abdominal ventrites weakly shiny, on sides with rows of punctures; anteriorly with an additional punctured furrow. Last abdominal ventrite shortened medially for reception of pygidium, with dense punctures, which are about one and half time larger than on penultimate ventrite; anterior half with dense, very deep punctures (pits) separated by narrow longitudinal ridges; posterior part in the middle with very deep groove, located very slightly below apex of pygidium. Meso- and metafemora with two very indistinct tubercles on posterior border; all femora shiny, with regular, very distinct, rather small, very dense punctation; all punctures bearing small macrosetae.
</paragraph>
<paragraph id="EB6236BD2355045E57D465A8FAB0FE29" blockId="8.[151,1437,151,573]" pageId="8" pageNumber="571">
<emphasis id="D9A9EAAF2355045E57D465A8FE9EFE70" bold="true" box="[199,312,402,428]" pageId="8" pageNumber="571">Variation</emphasis>
. Length
<quantity id="2C259B582355045E568A65A9FD87FE71" box="[409,545,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="8.6" metricValueMin="6.2" pageId="8" pageNumber="571" unit="mm" value="7.4" valueMax="8.6" valueMin="6.2">6.2–8.6 mm</quantity>
, width 2.00–
<quantity id="2C259B582355045E55A365A8FCB3FE71" box="[688,789,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" pageId="8" pageNumber="571" unit="mm" value="2.65">2.65 mm</quantity>
. Punctation of pronotum can be more or less dense. Sexual dimorphism evident in the shape of meso- and metatibiae, which are weakly sinuate on inner edge and are distinctly inwardly hooked at apex; additionally last abdominal ventrite and pygidium in males are not modified.
</paragraph>
</subSubSection>
<subSubSection id="A3C765362355045E57D465C5FEA2FDE1" pageId="8" pageNumber="571" type="etymology">
<paragraph id="EB6236BD2355045E57D465C5FEA2FDE1" blockId="8.[151,1437,151,573]" pageId="8" pageNumber="571">
<emphasis id="D9A9EAAF2355045E57D465C5FEE8FDC5" bold="true" box="[199,334,511,537]" pageId="8" pageNumber="571">Etymology.</emphasis>
Toponymic; an adjective derived from the name of
<collectingRegion id="2919F85F2355045E54D465C5FBBBFDC5" box="[967,1053,511,537]" country="Solomon Islands" name="Malaita" pageId="8" pageNumber="571">Malaita</collectingRegion>
Island, where the
<typeStatus id="3466881F2355045E521165C5FAC2FDC5" box="[1282,1380,511,537]" pageId="8" pageNumber="571" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<caption id="BFA266352355045E57846326FE14F834" ID-DOI="http://doi.org/10.5281/zenodo.14745471" ID-Zenodo-Dep="14745471" httpUri="https://zenodo.org/record/14745471/files/figure.png" pageId="8" pageNumber="571" startId="8.[151,264,1820,1844]" targetBox="[156,1431,601,1792]" targetPageId="8" targetType="figure">
<paragraph id="EB6236BD2355045E57846326FE14F834" blockId="8.[151,1437,1820,2024]" pageId="8" pageNumber="571">
<emphasis id="D9A9EAAF2355045E57846326FEF1F8E8" bold="true" box="[151,343,1820,1844]" pageId="8" pageNumber="571">FIGURES 4A–G.</emphasis>
Details of morphology. A,
<taxonomicName id="2CDD4D3E2355045E55696326FC33F8E8" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[634,917,1820,1844]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. nov.">
<emphasis id="D9A9EAAF2355045E55696326FC33F8E8" box="[634,917,1820,1844]" italics="true" pageId="8" pageNumber="571">Rhyparus guadalcanalensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42355045E54B66327FB83F8E8" box="[933,1061,1820,1844]" pageId="8" pageNumber="571" rank="species">new species</taxonomicNameLabel>
, male, holotype, head; B,
<taxonomicName id="2CDD4D3E2355045E52286326FEEEF884" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. nov.">
<emphasis id="D9A9EAAF2355045E52286326FEEEF884" italics="true" pageId="8" pageNumber="571">Rhyparus guadalcanalensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42355045E5648637BFE7BF884" box="[347,477,1856,1880]" pageId="8" pageNumber="571" rank="species">new species</taxonomicNameLabel>
, male, holotype, metatibia; C,
<taxonomicName id="2CDD4D3E2355045E5423637AFB45F885" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[816,1251,1856,1881]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="helophoroides">
<emphasis id="D9A9EAAF2355045E5423637AFB8BF884" box="[816,1069,1856,1880]" italics="true" pageId="8" pageNumber="571">Rhyparus helophoroides</emphasis>
<bibRefCitation id="8F4C4B4C2355045E532A637AFB45F885" author="Fairmaire, L." box="[1081,1251,1856,1881]" pageId="8" pageNumber="571" pagination="144 - 146" refId="ref7731" refString="Fairmaire, L. (1893) Description de quatre especes nouvelles du genre Rhyparus, Westw. (Scarabaeidae: Coprini). Note XVII. Notes from the Leyden Museum, 15, 144 - 146." type="journal article" year="1893">Fairmaire, 1893</bibRefCitation>
</taxonomicName>
, male, metatibia; D,
<taxonomicName id="2CDD4D3E2355045E57A9635EFE02F8A0" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[186,420,1892,1916]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="malaitaensis" status="sp. nov.">
<emphasis id="D9A9EAAF2355045E57A9635EFE02F8A0" box="[186,420,1892,1916]" italics="true" pageId="8" pageNumber="571">Rhyparus malaitaensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42355045E56A6635FFD93F8A0" box="[437,565,1892,1916]" pageId="8" pageNumber="571" rank="species">new species</taxonomicNameLabel>
, female, holotype, head; E,
<taxonomicName id="2CDD4D3E2355045E5473635EFBEDF8A0" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[864,1099,1892,1916]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="malaitaensis" status="sp. nov.">
<emphasis id="D9A9EAAF2355045E5473635EFBEDF8A0" box="[864,1099,1892,1916]" italics="true" pageId="8" pageNumber="571">Rhyparus malaitaensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42355045E5348635FFB7DF8A0" box="[1115,1243,1892,1916]" pageId="8" pageNumber="571" rank="species">new species</taxonomicNameLabel>
, female, holotype, pygidium and last abdominal ventrite; F,
<taxonomicName id="2CDD4D3E2355045E552E63B2FB87F87D" authority="Minkina &amp; Jakl, 2024" authorityName="Minkina &amp; Jakl" authorityYear="2024" box="[573,1057,1928,1953]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="argopuroensis">
<emphasis id="D9A9EAAF2355045E552E63B2FC91F87C" box="[573,823,1928,1952]" italics="true" pageId="8" pageNumber="571">Rhyparus argopuroensis</emphasis>
<bibRefCitation id="8F4C4B4C2355045E542E63B2FB87F87D" author="Minkina, L. &amp; JAkl, S." box="[829,1057,1928,1953]" pageId="8" pageNumber="571" pagination="131 - 152" refId="ref7932" refString="Minkina, L. &amp; JAkl, S. (2024) Seven new Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) species. Studies and Reports, Taxonomic Series, 20 (1), 131 - 152." type="journal article" year="2024">Minkina &amp; Jákl, 2024</bibRefCitation>
</taxonomicName>
, female, holotype, pygidium and last abdominal ventrite; G,
<taxonomicName id="2CDD4D3E2355045E56946396FBD3F819" authority="Minkina, Anichtchenko, Vasiljeva &amp; Skelley, 2022" authorityName="Minkina, Anichtchenko, Vasiljeva &amp; Skelley" authorityYear="2022" box="[391,1141,1964,1989]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="571" phylum="Arthropoda" rank="species" species="bacanensis">
<emphasis id="D9A9EAAF2355045E56946396FDC7F818" box="[391,609,1964,1988]" italics="true" pageId="8" pageNumber="571">Rhyparus bacanensis</emphasis>
<bibRefCitation id="8F4C4B4C2355045E557B6396FBD3F819" author="Minkina, L. &amp; Anichtchenko, A. &amp; Vasiljeva, A. &amp; Skelley, P." box="[616,1141,1964,1989]" pageId="8" pageNumber="571" pagination="81 - 101" refId="ref7869" refString="Minkina, L., Anichtchenko, A., Vasiljeva, A. &amp; Skelley, P. (2022) Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology, 22 (1), 81 - 101." type="journal article" year="2022">Minkina, Anichtchenko, Vasiljeva &amp; Skelley, 2022</bibRefCitation>
</taxonomicName>
, female, holotype, pygidium and last abdominal ventrite.
</paragraph>
</caption>
<subSubSection id="A3C765362354045F57D464ACFE9CFB18" pageId="9" pageNumber="572" type="diagnosis">
<paragraph id="EB6236BD2354045F57D464ACFAA4FEB8" blockId="9.[151,1437,150,1220]" pageId="9" pageNumber="572">
<emphasis id="D9A9EAAF2354045F57D464ACFE6AFF6C" bold="true" box="[199,460,150,176]" pageId="9" pageNumber="572">Differential diagnosis.</emphasis>
<taxonomicName id="2CDD4D3E2354045F56C264ACFD69FF6C" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[465,719,150,176]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="malaitaensis" status="sp. nov.">
<emphasis id="D9A9EAAF2354045F56C264ACFD69FF6C" box="[465,719,150,176]" italics="true" pageId="9" pageNumber="572">Rhyparus malaitaensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42354045F55C864ADFCC6FF6C" box="[731,864,150,177]" pageId="9" pageNumber="572" rank="species">new species</taxonomicNameLabel>
belongs to the group of species with females having a modified pygidium and last abdominal ventrite. Due to this modification, females are always easily distinguishable from other
<taxonomicName id="2CDD4D3E2354045F560464E4FE44FF24" box="[279,482,222,248]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D9A9EAAF2354045F560464E4FE24FF24" box="[279,386,222,248]" italics="true" pageId="9" pageNumber="572">Rhyparus</emphasis>
species.
</taxonomicName>
Males of this group are not so easily distinguishable and for this reason we selected a female as the
<typeStatus id="3466881F2354045F565A6538FE09FEC0" box="[329,431,258,284]" pageId="9" pageNumber="572" type="holotype">holotype</typeStatus>
. Members of this group can be distinguished from other
<taxonomicName id="2CDD4D3E2354045F53216538FB3BFEC0" authorityName="Westwood" authorityYear="1845" box="[1074,1181,258,284]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="D9A9EAAF2354045F53216538FB3BFEC0" box="[1074,1181,258,284]" italics="true" pageId="9" pageNumber="572">Rhyparus</emphasis>
</taxonomicName>
by the relatively large body (for a representatives of the genus), with a body length in nearly all cases more than 6.0 mm, elongate body, relatively weakly developed, rounded lateral lobes of pronotum, and weakly developed caudal bulbs.
</paragraph>
<paragraph id="EB6236BD2354045F57D46554FA89FCE5" blockId="9.[151,1437,150,1220]" pageId="9" pageNumber="572">
The pygidium of female
<taxonomicName id="2CDD4D3E2354045F56F66555FD32FE54" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[485,660,366,392]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="malaitaensis" status="new species">
<emphasis id="D9A9EAAF2354045F56F66555FD32FE54" box="[485,660,366,392]" italics="true" pageId="9" pageNumber="572">R. malaitaensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42354045F55B06555FC8AFE54" box="[675,812,366,393]" pageId="9" pageNumber="572" rank="species">new species</taxonomicNameLabel>
has the apex acute, shaped like a distinct triangle (not truncate). Of all species in the group with a modified female pygidium, only
<taxonomicName id="2CDD4D3E2354045F54FF65A9FE16FE0C" authority="Minkina, Anichtchenko, Vasiljeva &amp; Skelley 2022" authorityName="Minkina, Anichtchenko, Vasiljeva &amp; Skelley" authorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="bacanensis">
<emphasis id="D9A9EAAF2354045F54FF65A9FB21FE70" box="[1004,1159,402,428]" italics="true" pageId="9" pageNumber="572">R. bacanensis</emphasis>
<bibRefCitation id="8F4C4B4C2354045F539E65A8FE16FE0C" author="Minkina, L. &amp; Anichtchenko, A. &amp; Vasiljeva, A. &amp; Skelley, P." pageId="9" pageNumber="572" pagination="81 - 101" refId="ref7869" refString="Minkina, L., Anichtchenko, A., Vasiljeva, A. &amp; Skelley, P. (2022) Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology, 22 (1), 81 - 101." type="journal article" year="2022">Minkina, Anichtchenko, Vasiljeva &amp; Skelley 2022</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="2CDD4D3E2354045F56F7658DFC3EFE0C" authority="Minkina &amp; Jakl, 2024" authorityName="Minkina &amp; Jakl" authorityYear="2024" box="[484,920,438,464]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="argopuroensis">
<emphasis id="D9A9EAAF2354045F56F7658DFD07FE0C" box="[484,673,439,464]" italics="true" pageId="9" pageNumber="572">R. argopuroensis</emphasis>
<bibRefCitation id="8F4C4B4C2354045F55B4658CFC3EFE0C" author="Minkina, L. &amp; JAkl, S." box="[679,920,438,464]" pageId="9" pageNumber="572" pagination="131 - 152" refId="ref7932" refString="Minkina, L. &amp; JAkl, S. (2024) Seven new Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) species. Studies and Reports, Taxonomic Series, 20 (1), 131 - 152." type="journal article" year="2024">Minkina &amp; Jákl, 2024</bibRefCitation>
</taxonomicName>
have a similar pygidial apex (see figs
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<bibRefCitation id="8F4C4B4C2354045F578465E0FED6FE28" author="Minkina, L. &amp; Anichtchenko, A. &amp; Vasiljeva, A. &amp; Skelley, P." box="[151,368,474,500]" pageId="9" pageNumber="572" pagination="81 - 101" refId="ref7869" refString="Minkina, L., Anichtchenko, A., Vasiljeva, A. &amp; Skelley, P. (2022) Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology, 22 (1), 81 - 101." type="journal article" year="2022">
Minkina
<emphasis id="D9A9EAAF2354045F57EF65E1FE95FE28" box="[252,307,474,500]" italics="true" pageId="9" pageNumber="572">et al.</emphasis>
2022
</bibRefCitation>
and figs 1–3, 22,
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<bibRefCitation id="8F4C4B4C2354045F556365E0FCF8FE28" author="Minkina, L. &amp; JAkl, S." box="[624,862,474,500]" pageId="9" pageNumber="572" pagination="131 - 152" refId="ref7932" refString="Minkina, L. &amp; JAkl, S. (2024) Seven new Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) species. Studies and Reports, Taxonomic Series, 20 (1), 131 - 152." type="journal article" year="2024">Minkina &amp; Jákl 2024</bibRefCitation>
), but they have different last abdominal ventrites. In
<taxonomicName id="2CDD4D3E2354045F578465C5FEE4FDC4" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[151,322,510,536]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="malaitaensis" status="new species">
<emphasis id="D9A9EAAF2354045F578465C5FEE4FDC4" box="[151,322,510,536]" italics="true" pageId="9" pageNumber="572">R. malaitaensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42354045F565E65C5FE77FDC4" box="[333,465,510,537]" pageId="9" pageNumber="572" rank="species">new species</taxonomicNameLabel>
the pits in the anterior furrow are about 1.5–2.5 times longer than wider; in
<taxonomicName id="2CDD4D3E2354045F521065C5FA3AFDC4" authorityName="Minkina, Anichtchenko, Vasiljeva &amp; Skelley" authorityYear="2022" box="[1283,1436,510,536]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="bacanensis">
<emphasis id="D9A9EAAF2354045F521065C5FA3AFDC4" box="[1283,1436,510,536]" italics="true" pageId="9" pageNumber="572">R. bacanensis</emphasis>
</taxonomicName>
the pits are about 3.5–5.0 times longer than wider; while
<taxonomicName id="2CDD4D3E2354045F540D6619FC78FDE0" box="[798,990,547,572]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="argopuorensis">
<emphasis id="D9A9EAAF2354045F540D6619FC78FDE0" box="[798,990,547,572]" italics="true" pageId="9" pageNumber="572">R. argopuorensis</emphasis>
</taxonomicName>
, has much smaller pits, similar to large punctures.
<taxonomicName id="2CDD4D3E2354045F5607667CFDA3FDBC" authorityName="Minkina, Anichtchenko, Vasiljeva &amp; Skelley" authorityYear="2022" box="[276,517,582,608]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="bacanensis">
<emphasis id="D9A9EAAF2354045F5607667CFDA3FDBC" box="[276,517,582,608]" italics="true" pageId="9" pageNumber="572">Rhyparus bacanensis</emphasis>
</taxonomicName>
have apical part of last abdominal ventrite located distinctly below the pygidial apex;
<taxonomicName id="2CDD4D3E2354045F57CE6651FE2DFD58" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[221,395,618,644]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="malaitaensis" status="new species and r.">
<emphasis id="D9A9EAAF2354045F57CE6651FE2DFD58" box="[221,395,618,644]" italics="true" pageId="9" pageNumber="572">R. malaitaensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42354045F56886651FDD3FD58" box="[411,629,618,645]" pageId="9" pageNumber="572" rank="subSpecies">
new species and
<taxonomicName id="2CDD4D3E2354045F554E6651FDD3FD58" authorityName="Minkina &amp; Jakl" authorityYear="2024" box="[605,629,619,644]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="argopuroensis">
<emphasis id="D9A9EAAF2354045F554E6651FDD3FD58" box="[605,629,619,644]" italics="true" pageId="9" pageNumber="572">R.</emphasis>
</taxonomicName>
</taxonomicNameLabel>
<emphasis id="D9A9EAAF2354045F556D6651FCB8FD58" box="[638,798,619,644]" italics="true" pageId="9" pageNumber="572">argopuroensis</emphasis>
have apical part of last abdominal ventrite located very slightly below the pygidial apex. Female pygidia of these three species are not easy to distinguish without having specimens of all species for comparison: proportions and prominence (especially median carina of pygidium) are slightly different. Another feature that distinguishes females of
<taxonomicName id="2CDD4D3E2354045F547166EDFBB6FD2C" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[866,1040,726,752]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="malaitaensis" status="new species and r.">
<emphasis id="D9A9EAAF2354045F547166EDFBB6FD2C" box="[866,1040,726,752]" italics="true" pageId="9" pageNumber="572">R. malaitaensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42354045F530D66EDFB53FD2C" box="[1054,1269,726,753]" pageId="9" pageNumber="572" rank="subSpecies">
new species and
<taxonomicName id="2CDD4D3E2354045F53CE66EDFB53FD2C" authorityName="Minkina &amp; Jakl" authorityYear="2024" box="[1245,1269,727,752]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="argopuroensis">
<emphasis id="D9A9EAAF2354045F53CE66EDFB53FD2C" box="[1245,1269,727,752]" italics="true" pageId="9" pageNumber="572">R.</emphasis>
</taxonomicName>
</taxonomicNameLabel>
<emphasis id="D9A9EAAF2354045F53EF66EDFA3AFD2C" box="[1276,1436,727,752]" italics="true" pageId="9" pageNumber="572">argopuroensis</emphasis>
from
<taxonomicName id="2CDD4D3E2354045F57C766C1FEC8FCC8" authorityName="Minkina, Anichtchenko, Vasiljeva &amp; Skelley" authorityYear="2022" box="[212,366,762,788]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="bacanensis">
<emphasis id="D9A9EAAF2354045F57C766C1FEC8FCC8" box="[212,366,762,788]" italics="true" pageId="9" pageNumber="572">R. bacanensis</emphasis>
</taxonomicName>
is the shape of metaventral plate:
<taxonomicName id="2CDD4D3E2354045F55FA66C0FC71FCC8" authorityName="Minkina, Anichtchenko, Vasiljeva &amp; Skelley" authorityYear="2022" box="[745,983,762,788]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="bacanensis">
<emphasis id="D9A9EAAF2354045F55FA66C0FC71FCC8" box="[745,983,762,788]" italics="true" pageId="9" pageNumber="572">Rhyparus bacanensis</emphasis>
</taxonomicName>
have distinct concavities on each side of the median longitudinal furrow, which are lacking on
<taxonomicName id="2CDD4D3E2354045F55E16725FC39FCE4" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[754,927,798,824]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="malaitaensis" status="new species and r.">
<emphasis id="D9A9EAAF2354045F55E16725FC39FCE4" box="[754,927,798,824]" italics="true" pageId="9" pageNumber="572">R. malaitaensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42354045F54BF6725FB27FCE4" box="[940,1153,798,825]" pageId="9" pageNumber="572" rank="subSpecies">
new species and
<taxonomicName id="2CDD4D3E2354045F537A6725FB27FCE4" authorityName="Minkina &amp; Jakl" authorityYear="2024" box="[1129,1153,799,824]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="argopuroensis">
<emphasis id="D9A9EAAF2354045F537A6725FB27FCE4" box="[1129,1153,799,824]" italics="true" pageId="9" pageNumber="572">R.</emphasis>
</taxonomicName>
</taxonomicNameLabel>
<emphasis id="D9A9EAAF2354045F539B6725FA8EFCE4" box="[1160,1320,799,824]" italics="true" pageId="9" pageNumber="572">argopuroensis</emphasis>
.
</paragraph>
<paragraph id="EB6236BD2354045F57D46778FD9DFBA1" blockId="9.[151,1437,150,1220]" pageId="9" pageNumber="572">
Males of this group with modified female pygidia are difficult to distinguish. Males of
<taxonomicName id="2CDD4D3E2354045F53BC6779FAF8FC80" authorityName="Minkina &amp; Skelley &amp; Jákl &amp; Král &amp; Li" authorityYear="2025" box="[1199,1374,834,861]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="malaitaensis" status="new species">
<emphasis id="D9A9EAAF2354045F53BC6779FB66FC80" box="[1199,1216,835,860]" italics="true" pageId="9" pageNumber="572">R</emphasis>
.
<emphasis id="D9A9EAAF2354045F53C36778FAF8FC80" box="[1232,1374,834,860]" italics="true" pageId="9" pageNumber="572">malaitaensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C29A57D42354045F527D6779FF4EFC5C" pageId="9" pageNumber="572" rank="species">new species</taxonomicNameLabel>
can be recognised by comparing the following combination of characters with those presented in Table 1 and Table 2 of
<bibRefCitation id="8F4C4B4C2354045F567467B0FDC0FC78" author="Minkina, L. &amp; Anichtchenko, A. &amp; Vasiljeva, A. &amp; Skelley, P." box="[359,614,906,932]" pageId="9" pageNumber="572" pagination="81 - 101" refId="ref7869" refString="Minkina, L., Anichtchenko, A., Vasiljeva, A. &amp; Skelley, P. (2022) Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology, 22 (1), 81 - 101." type="journal article" year="2022">
Minkina
<emphasis id="D9A9EAAF2354045F56C267B1FDABFC78" box="[465,525,906,932]" italics="true" pageId="9" pageNumber="572">et al.</emphasis>
(2022)
</bibRefCitation>
: all costae shiny, intercostae of pronotum weakly shiny, intercostae of elytra matt; body distinctly elongate, lateral lobes of pronotum weakly developed, sinuation between them not so distinct, anterior lateral lobe of pronotum triangular, very slightly higher than posteriorly; posterior lateral lobe of pronotum widely, regularly rounded; punctation of median intercostae of pronotum dense, regularly spaced, distinctly concentrate in the middle into longitudinal line, punctures coarse here; elytral costae distinctly convex, relatively narrow; some additional fine punctures are observed on all intercostae; third intercostae with three distinct rows of moderately coarse punctures.
</paragraph>
<paragraph id="EB6236BD2354045F57D460BCFE9CFB18" blockId="9.[151,1437,150,1220]" pageId="9" pageNumber="572">
Additionally it can be identified from other members of the genus
<taxonomicName id="2CDD4D3E2354045F54AD60BCFB8FFB7C" authorityName="Westwood" authorityYear="1845" box="[958,1065,1158,1184]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="D9A9EAAF2354045F54AD60BCFB8FFB7C" box="[958,1065,1158,1184]" italics="true" pageId="9" pageNumber="572">Rhyparus</emphasis>
</taxonomicName>
from the
<collectingCountry id="93CA762D2354045F538960BCFAF1FB7C" box="[1178,1367,1158,1184]" name="Solomon Islands" pageId="9" pageNumber="572">Solomon Islands</collectingCountry>
using the key below.
</paragraph>
</subSubSection>
</treatment>
</document>
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