167 lines
19 KiB
XML
167 lines
19 KiB
XML
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<mods:title id="F46C40260852AD15904544C40E8A4C20">Review of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) in the Solomon Islands with description of two new species</mods:title>
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<mods:namePart id="168F3DB7705D8A9AB12B293F9D753B15">Minkina, Łukasz</mods:namePart>
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<mods:affiliation id="DC32077C6D7B35E6BFF2B9F9F274B776">os. Polana Szaflarska 4 / 39, 34 - 400 Nowy Targ, Poland.</mods:affiliation>
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<mods:namePart id="A04B667277E1E3C4D8D6FC80616C37A9">Skelley, Paul</mods:namePart>
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<mods:affiliation id="6C85ECC0D1FE9D9051159A553587F328">Florida State Collection of Arthropods, 1911 SW 34 th Street, Gainesville, FL, USA.</mods:affiliation>
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<mods:namePart id="DEE16DA1C609B106CB3D20A80E4B553F">Jákl, Stanislav</mods:namePart>
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<mods:affiliation id="E2B05756C21C5F34ED0F21E9BE8218AE">Geologická 1218 / 2 C, CZ- 152 00 Praha 5, Czechia.</mods:affiliation>
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<mods:namePart id="14E0447B08AF960C0BD50175EF1CB1A7">Král, David</mods:namePart>
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<mods:affiliation id="4A1880C650CADE8F11CA86F0F35D0BF3">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czechia.</mods:affiliation>
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<mods:namePart id="CF808FDAD8AC01F0949CFA37F9106D78">Li, Xiao-Feng</mods:namePart>
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<mods:nameIdentifier id="DC908203393C4BEA44EE7006AF3A0A96" type="email">yansi3241@gmail.com</mods:nameIdentifier>
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<mods:title id="627322285923110081414861B96F3F73">Zootaxa</mods:title>
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<mods:date id="C93640AADF3EA0FB7C83A1E1FEEBCF29">2025</mods:date>
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Key to the species of the genus
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<taxonomicName id="2CDD4D3E2354045F5509620CFD29F98D" ID-CoL="79QS" authority="Westwood, 1845" authorityName="Westwood" authorityYear="1845" box="[538,655,1590,1617]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="genus">
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<emphasis id="D9A9EAAF2354045F5509620CFD29F98D" bold="true" box="[538,655,1590,1617]" italics="true" pageId="9" pageNumber="572">Rhyparus</emphasis>
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from the
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<collectingCountry id="93CA762D2354045F5419620CFC7FF98C" box="[778,985,1590,1616]" name="Solomon Islands" pageId="9" pageNumber="572">Solomon Islands</collectingCountry>
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</emphasis>
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<keyLead id="502C95882354045F57846241FA3AF935" pageId="9" pageNumber="572">
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1. Pygidium of females distinctly bent inwards (lateral view), with last abdominal ventrite shortened medially to receive pygidium (
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<figureCitation id="73E62A382354045F57FA62A2FE96F972" box="[233,304,1688,1710]" captionStart="FIGURES 4" captionStartId="8.[151,264,1820,1844]" captionTargetBox="[156,1431,601,1792]" captionTargetId="figure-195@8.[151,1435,597,1797]" captionTargetPageId="8" captionText="FIGURES 4A–G. Details of morphology. A, Rhyparus guadalcanalensis, new species, male, holotype, head; B, Rhyparus guadalcanalensis, new species, male, holotype, metatibia; C, Rhyparus helophoroides Fairmaire, 1893, male, metatibia; D, Rhyparus malaitaensis, new species, female, holotype, head; E, Rhyparus malaitaensis, new species, female, holotype, pygidium and last abdominal ventrite; F, Rhyparus argopuroensis Minkina & Jákl, 2024, female, holotype, pygidium and last abdominal ventrite; G, Rhyparus bacanensis Minkina, Anichtchenko, Vasiljeva & Skelley, 2022, female, holotype, pygidium and last abdominal ventrite." figureDoi="http://doi.org/10.5281/zenodo.14745471" httpUri="https://zenodo.org/record/14745471/files/figure.png" pageId="9" pageNumber="572">Fig. 4E</figureCitation>
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). Body length more than
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. Lateral lobes of pronotum weakly developed. Caudal bulbs of eltyra lacking sinuation between median and lateral bulbs. Third intercostae of elytra with three distinct rows of medium sized punctures.......................................................................................
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<taxonomicName id="2CDD4D3E2354045F539A62EEFABBF934" authorityName="Minkina & Skelley & Jákl & Král & Li" authorityYear="2025" box="[1161,1309,1747,1769]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="malaitaensis" status="new species">
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<emphasis id="D9A9EAAF2354045F539A62EEFB3EF935" box="[1161,1176,1748,1769]" italics="true" pageId="9" pageNumber="572">R</emphasis>
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.
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<emphasis id="D9A9EAAF2354045F53B762E9FABBF934" box="[1188,1309,1747,1768]" italics="true" pageId="9" pageNumber="572">malaitaensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="C29A57D42354045F523A62EEFA3AF935" box="[1321,1436,1747,1769]" pageId="9" pageNumber="572" rank="species">new species</taxonomicNameLabel>
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<paragraph id="EB6236BD2354045F578462CBFA3AF89C" blockId="9.[151,1437,1659,2004]" pageId="9" pageNumber="572">
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<keyLead id="502C95882354045F578462CBFA3AF89C" pageId="9" pageNumber="572">
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- Pygidium and last abdominal ventrite in females normally developed. Body length more than
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<quantity id="2C259B582354045F534762CBFB39F8DA" box="[1108,1183,1777,1798]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="572" unit="mm" value="3.4">3.4 mm</quantity>
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. Lateral lobes of pronotum distinctly developed. Caudal bulbs of elytra with sinuation between median and lateral bulbs more or less distinct. Third intercostae of elytra with two rows of punctures or with a third median row of much finer punctures than lateral rows...... 2
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<paragraph id="EB6236BD2354045F57846373FA3AF80F" blockId="9.[151,1437,1659,2004]" pageId="9" pageNumber="572">
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<keyLead id="502C95882354045F57846373FA3AF80F" pageId="9" pageNumber="572">
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2. Larger species; body length more than
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.Anterior lateral lobes of pronotum more prominent than median lobe. Pronotum with posterior half of median intercosta always with dense punctation. Third intercostae of elytra usually with two distinct rows of punctures, some with a third median row with much finer punctures. Caudal bulbs of elytra with sinuation between median and lateral bulbs always distinct. Meso- and metatibiae proportionally longer, distinctly longer than all tarsomeres together. Meso- and metatarsi densely pilose ventrally, surface of tarsomeres not visible.................
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<taxonomicName id="2CDD4D3E2354045F538E6384FA3AF80F" ID-CoL="4SV45" authority="Paulian, 1984" authorityName="Paulian" authorityYear="1984" box="[1181,1436,1982,2004]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="breviceps">
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<emphasis id="D9A9EAAF2354045F538E6384FB0AF80F" box="[1181,1196,1982,2003]" italics="true" pageId="9" pageNumber="572">R</emphasis>
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.
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<emphasis id="D9A9EAAF2354045F53A46384FAB4F80F" box="[1207,1298,1982,2003]" italics="true" pageId="9" pageNumber="572">breviceps</emphasis>
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<bibRefCitation id="8F4C4B4C2354045F520B6384FA3AF80F" author="Paulian, R." box="[1304,1436,1982,2004]" pageId="9" pageNumber="572" pagination="471 - 474" refId="ref8066" refString="Paulian, R. (1984) Notes sur les Coleopteres Scarabaeidae du Museum de Geneve, I. Revue Suisse de Zoologie, 91, 471 - 474. https: // doi. org / 10.5962 / bhl. part. 81890" type="journal article" year="1984">Paulian, 1984</bibRefCitation>
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<paragraph id="EB6236BD2357045C578464A2FA3AFEFE" blockId="10.[151,1437,151,467]" pageId="10" pageNumber="573">
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- Smaller species; body length
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. Anterior lateral lobes of pronotum lower than posterior. Pronotum with posterior half of median intercosta with or without punctation. Third intercostae of elytra always with two rows of punctures. Caudal bulbs of elytra with sinuation between median and lateral caudal bulbs indistinct or very weakly visible. Meso- and metatibiae proportionally shorter; similar in length to all tarsomeres together. Meso- and metatarsi not densely pilose ventrally, surface of tarsomeres visible..................................................................................... 3
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</paragraph>
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<paragraph id="EB6236BD2357045C57846510FA3AFEA6" blockId="10.[151,1437,151,467]" pageId="10" pageNumber="573">
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<keyLead id="502C95882357045C57846510FA3AFEA6" pageId="10" pageNumber="573">
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3. Pronotum with posterior third of median intercosta usually bearing some dense punctures. Elytral intercostae rarely with visible setae between rows of punctures. Metatibia of male (
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<figureCitation id="73E62A382357045C55C6657DFCB9FE81" box="[725,799,327,349]" captionStart="FIGURES 4" captionStartId="8.[151,264,1820,1844]" captionTargetBox="[156,1431,601,1792]" captionTargetId="figure-195@8.[151,1435,597,1797]" captionTargetPageId="8" captionText="FIGURES 4A–G. Details of morphology. A, Rhyparus guadalcanalensis, new species, male, holotype, head; B, Rhyparus guadalcanalensis, new species, male, holotype, metatibia; C, Rhyparus helophoroides Fairmaire, 1893, male, metatibia; D, Rhyparus malaitaensis, new species, female, holotype, head; E, Rhyparus malaitaensis, new species, female, holotype, pygidium and last abdominal ventrite; F, Rhyparus argopuroensis Minkina & Jákl, 2024, female, holotype, pygidium and last abdominal ventrite; G, Rhyparus bacanensis Minkina, Anichtchenko, Vasiljeva & Skelley, 2022, female, holotype, pygidium and last abdominal ventrite." figureDoi="http://doi.org/10.5281/zenodo.14745471" httpUri="https://zenodo.org/record/14745471/files/figure.png" pageId="10" pageNumber="573">Fig. 4C</figureCitation>
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) with inner margin swollen at basal third, then flattened or concave to apex which lacks an inner apical tooth.........................................
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<taxonomicName id="2CDD4D3E2357045C5349655FFA3AFEA6" ID-CoL="4SV4K" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[1114,1436,357,379]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="species" species="helophoroides">
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<emphasis id="D9A9EAAF2357045C5349655FFBCFFEA6" box="[1114,1129,357,378]" italics="true" pageId="10" pageNumber="573">R</emphasis>
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.
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<emphasis id="D9A9EAAF2357045C5367655FFB5AFEA6" box="[1140,1276,357,378]" italics="true" pageId="10" pageNumber="573">helophoroides</emphasis>
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<bibRefCitation id="8F4C4B4C2357045C5210655FFA3AFEA6" author="Fairmaire, L." box="[1283,1436,357,379]" pageId="10" pageNumber="573" pagination="144 - 146" refId="ref7731" refString="Fairmaire, L. (1893) Description de quatre especes nouvelles du genre Rhyparus, Westw. (Scarabaeidae: Coprini). Note XVII. Notes from the Leyden Museum, 15, 144 - 146." type="journal article" year="1893">Fairmaire, 1893</bibRefCitation>
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</taxonomicName>
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</paragraph>
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- Pronotum with basal third of median intercosta lacking punctures. Elytral intercostae with macrosetae between rows of punctures. Metatibia of male (
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<figureCitation id="73E62A382357045C551065A5FDEBFE69" box="[515,589,415,437]" captionStart="FIGURES 4" captionStartId="8.[151,264,1820,1844]" captionTargetBox="[156,1431,601,1792]" captionTargetId="figure-195@8.[151,1435,597,1797]" captionTargetPageId="8" captionText="FIGURES 4A–G. Details of morphology. A, Rhyparus guadalcanalensis, new species, male, holotype, head; B, Rhyparus guadalcanalensis, new species, male, holotype, metatibia; C, Rhyparus helophoroides Fairmaire, 1893, male, metatibia; D, Rhyparus malaitaensis, new species, female, holotype, head; E, Rhyparus malaitaensis, new species, female, holotype, pygidium and last abdominal ventrite; F, Rhyparus argopuroensis Minkina & Jákl, 2024, female, holotype, pygidium and last abdominal ventrite; G, Rhyparus bacanensis Minkina, Anichtchenko, Vasiljeva & Skelley, 2022, female, holotype, pygidium and last abdominal ventrite." figureDoi="http://doi.org/10.5281/zenodo.14745471" httpUri="https://zenodo.org/record/14745471/files/figure.png" pageId="10" pageNumber="573">Fig. 4B</figureCitation>
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) with inner margin regularly rounded entire length to a large inner apical tooth................................................................................
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<taxonomicName id="2CDD4D3E2357045C53496587FABBFE0E" authorityName="Minkina & Skelley & Jákl & Král & Li" authorityYear="2025" box="[1114,1309,445,466]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="573" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="new species">
|
||
<emphasis id="D9A9EAAF2357045C53496587FBCFFE0E" box="[1114,1129,445,466]" italics="true" pageId="10" pageNumber="573">R</emphasis>
|
||
.
|
||
<emphasis id="D9A9EAAF2357045C53666587FABBFE0E" box="[1141,1309,445,466]" italics="true" pageId="10" pageNumber="573">guadalcanalensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="C29A57D42357045C523A6587FA3AFE0F" box="[1321,1436,445,467]" pageId="10" pageNumber="573" rank="species">new species</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |