286 lines
29 KiB
XML
286 lines
29 KiB
XML
<document id="0BB4A7F4F783CE7B85AD19DDFA72A2E3" ID-CLB-Dataset="35540" ID-DOI="http://dx.doi.org/10.3897/zookeys.350.6127" ID-GBIF-Dataset="d6cfb7e6-c895-4063-b928-f2f01e407a59" ID-PMC="PMC3837497" ID-Pensoft-Pub="1313-2970-350-21" ID-PubMed="24294082" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-350-21" ModsDocOrigin="ZooKeys 350" ModsDocTitle="Taxonomic revision of Madagascan Rhantus (Coleoptera, Dytiscidae, Colymbetinae) with an emphasis on Manjakatompo as a conservation priority" checkinTime="1451246682682" checkinUser="pensoft" docAuthor="Hjalmarsson, Anna Emilia, Bukontaite, Rasa, Ranarilalatiana, Tolotra, Randriamihaja, Jacquelin Herisahala & Bergsten, Johannes" docDate="2013" docId="636DDD6BB8F248A5FC36B2E42DCB1BCA" docLanguage="en" docName="ZooKeys 350: 21-45" docOrigin="ZooKeys 350" docSource="http://dx.doi.org/10.3897/zookeys.350.6127" docTitle="Rhantus manjakatompo Pederzani & Rocchi 2009" docType="treatment" docVersion="5" lastPageNumber="28" masterDocId="164B5B6F0B649731770BFF989706FFDB" masterDocTitle="Taxonomic revision of Madagascan Rhantus (Coleoptera, Dytiscidae, Colymbetinae) with an emphasis on Manjakatompo as a conservation priority" masterLastPageNumber="45" masterPageNumber="21" pageNumber="24" updateTime="1732810536443" updateUser="ExternalLinkService">
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<mods:titleInfo id="E5DE64020219799E294BB36E376D7D19">
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<mods:title id="01398AD80526F5872BC1E653A63C45DC">Taxonomic revision of Madagascan Rhantus (Coleoptera, Dytiscidae, Colymbetinae) with an emphasis on Manjakatompo as a conservation priority</mods:title>
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<mods:namePart id="6445D61A957B381A0176D81716426B13">Hjalmarsson, Anna Emilia</mods:namePart>
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<mods:namePart id="F69AC9C5F76179F1CA0C547E604F2794">Bukontaite, Rasa</mods:namePart>
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<mods:namePart id="38409649B828894A599B99A6E922C91E">Ranarilalatiana, Tolotra</mods:namePart>
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<mods:namePart id="260050ADA2446B7364D896DCB8CF7D66">Randriamihaja, Jacquelin Herisahala</mods:namePart>
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<mods:namePart id="31998BDA7DCB20258A619082815EB111">Bergsten, Johannes</mods:namePart>
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<mods:title id="3AEC5EE639F719D86D1300FA9E7173AA">ZooKeys</mods:title>
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<mods:date id="D3AFAB6E86F81ABE8E4933F3D4D30883">2013</mods:date>
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<mods:number id="0A36585EFFABEB285F5B368E8DE360A1">350</mods:number>
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<mods:url id="4AFFE7632019F6CAC4BA28757ED4FF54">http://dx.doi.org/10.3897/zookeys.350.6127</mods:url>
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<mods:classification id="E6E444FACC82247B494E9E75F09C4FAF">journal article</mods:classification>
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<mods:identifier id="4C32964CA3BB912CAC0C3D517D3AC3E3" type="DOI">http://dx.doi.org/10.3897/zookeys.350.6127</mods:identifier>
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<mods:identifier id="097B7CCD398882AB25212C44A936FBB2" type="Pensoft-Pub">1313-2970-350-21</mods:identifier>
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<treatment id="636DDD6BB8F248A5FC36B2E42DCB1BCA" ID-GBIF-Taxon="152049549" LSID="urn:lsid:plazi:treatment:636DDD6BB8F248A5FC36B2E42DCB1BCA" httpUri="http://treatment.plazi.org/id/636DDD6BB8F248A5FC36B2E42DCB1BCA" lastPageId="7" lastPageNumber="28" pageId="3" pageNumber="24" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Colymbetinae">
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<subSubSection id="89122B9C1F08A24236AADF192FF6F501" pageId="3" pageNumber="24" type="nomenclature">
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<paragraph id="79460CDF59CAD930BF275901EA8B3C85" pageId="3" pageNumber="24">
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<taxonomicName id="2AEA0CF72EA43DDE6984D616FE5B6F1B" ID-CoL="4S3NM" LSID="http://species-id.net/wiki/Rhantus_manjakatompo" authority="Pederzani & Rocchi, 2009" authorityName="Pederzani & Rocchi" authorityYear="2009" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo Pederzani & Rocchi, 2009</taxonomicName>
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Figures 2
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<normalizedToken id="D67482FB5B84FEA69FAE254FF7C0D5DD" originalValue="a–b">a-b</normalizedToken>
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, 3
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<normalizedToken id="1C2D2BB75F217A9C048E4C389AFFF6BE" originalValue="a–c">a-c</normalizedToken>
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, 4a, 5
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<normalizedToken id="874B5A6B04144FE327DD4AC3D47BF93E" originalValue="a–b">a-b</normalizedToken>
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, 6
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<normalizedToken id="F0521AF5D7C31D096B8E07CF8A25118E" originalValue="a–b">a-b</normalizedToken>
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, 7
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<normalizedToken id="9DCB6A2C6AE1130E981F6DC5E14F88F3" originalValue="a–b">a-b</normalizedToken>
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, g
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</paragraph>
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</subSubSection>
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<subSubSection id="04C15ACD922A49D3BFA03BBAD65A59B9" pageId="3" pageNumber="24" type="reference_group">
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<paragraph id="F00E176EA5A294AF6E50C1C9BF22A2EC" pageId="3" pageNumber="24">
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<taxonomicName id="6536AA24B3C4764CF81DD2C27574643B" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
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Pederzani & Rocchi, 2009: 88
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</paragraph>
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</subSubSection>
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<subSubSection id="9F356449C66A5C284C5A418A35574A70" lastPageId="4" lastPageNumber="25" pageId="3" pageNumber="24" type="type locality">
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<paragraph id="AA10714046B9FD00AD5A8C9586A68C28" pageId="3" pageNumber="24">Type locality.</paragraph>
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<paragraph id="428CA28F81CD5F5A145829BF0F9062CC" pageId="3" pageNumber="24">Madagascar, Antananarivo province, Ankaratra mountains, Manjakatompo reserve.</paragraph>
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<paragraph id="4B95AAC5EA55E833ADA2C7854E91FA23" pageId="3" pageNumber="24">Type material (NMW). Holotype ♂ "Madagascar, Ankaratra (Antananarivo), Reserve Manjakatompo, spring stream exp. SE. N deviation to Analamitana (left affl. MD 107) m 1850 asl, 8.x.2001, 16.0°C, 0.003 mS/cm. Gerecke & Goldschmidt collectors BMNH (E) 2004-46", "Holotype Rhantus manjakatompo Pederzani & Rocchi 2008", "Data in NHRS-JLKB 000030412" "DNA Voucher".</paragraph>
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<paragraph id="AE5924363C219EB2E1E8808600E0A376" pageId="4" pageNumber="25">
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<pageBreakToken id="BA19008FE6AF11FBA1CA3E5F3A495946" pageId="4" pageNumber="25" start="start">Additional</pageBreakToken>
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material studied (NHRS, BMNH, DEUA, see Appendix): Sofia region (former provinces: Mahajanga): 2♂, 2♀ (Cat. No. BMNH-672731, 672730, 672726, 672725): Tsaratanana [Antetykalambazaha Camp],
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<geoCoordinate id="78D9128053808D1F91E75EF3EA3643DD" direction="south" orientation="latitude" precision="5" value="-14.1824">14.1824S</geoCoordinate>
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,
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<geoCoordinate id="59F4E1C1A61957090200F8C4D62785CB" direction="east" orientation="longitude" precision="5" value="48.9448">48.9448E</geoCoordinate>
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, 1700m, 20-24.xii.2004, Malaisetrap, Leg. Lees, Ranaivosolo.
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</paragraph>
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<paragraph id="ACC300915E198CB6AB343E8213FC14DC" pageId="4" pageNumber="25">
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Vakinankaratra region (former provinces: Antananarivo): 4spp. (Cat. No. NHRS-JLKB000021018) Manjakatompo [Analamitana] [swamp near stream][MJK12-02],
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<geoCoordinate id="DB760E54B03FEF08C8758DA16E5F603A" direction="south" orientation="latitude" precision="5" value="-19.364">19.3640S</geoCoordinate>
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,
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<geoCoordinate id="32EB213020AEAD27A945C04A35B1C8F8" direction="east" orientation="longitude" precision="5" value="47.2991">47.2991E</geoCoordinate>
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, 1760m, 22.i.2012, Leg. Ranarilalatiana, Randriamihaja, 1sp. (Cat. No. NHRS-JLKB000021019) Manjakatompo [Tavolatara][swamp near stream][MJK12-08],
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<geoCoordinate id="8EBC118CBD672E6B449391DD9727621F" direction="south" orientation="latitude" precision="5" value="-19.3491">19.3491S</geoCoordinate>
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,
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<geoCoordinate id="D0C6C0730D75FF234DF849B9DE2CCB58" direction="east" orientation="longitude" precision="5" value="47.2784">47.2784E</geoCoordinate>
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, 2050m, 24.i.2012, Leg. Ranarilalatiana, Randriamihaja, 5spp. (Cat. No. NHRS-JLKB000021020) Manjakatompo [Tavolatara][pool near stream][MJK12-09],
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<geoCoordinate id="5FAACC3450B3C6527D075BBAC72402B0" direction="south" orientation="latitude" precision="5" value="-19.3491">19.3491S</geoCoordinate>
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,
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<geoCoordinate id="23C958A22A815F32EC0989BCAD7FE1FB" direction="east" orientation="longitude" precision="5" value="47.278">47.2780E</geoCoordinate>
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, 2050m, 24.i.2012, Leg. Ranarilalatiana, Randriamihaja, 11spp. (Cat. No. NHRS-JLKB000021021) Manjakatompo [Tavolatara][swamp near source][MJK12-10],
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<geoCoordinate id="04FCA92AA46D8DBED9ECBD809C3C05D3" direction="south" orientation="latitude" precision="5" value="-19.3496">19.3496S</geoCoordinate>
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,
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<geoCoordinate id="7DFAA9B86DCD3B60046557DBC5DDAB29" direction="east" orientation="longitude" precision="5" value="47.2779">47.2779E</geoCoordinate>
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, 2050m, 24.i.2012, Leg. Ranarilalatiana, Randriamihaja, 3spp. (Cat. No. NHRS-JLKB000021022) Manjakatompo [Anosiarivo][lake near source][MJK12-13],
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<geoCoordinate id="1A75C3797D5768A40F42F24128D65066" direction="south" orientation="latitude" precision="5" value="-19.3449">19.3449S</geoCoordinate>
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,
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<geoCoordinate id="F4EA9A6879A0410AADFCCF9B65682724" direction="east" orientation="longitude" precision="5" value="47.3041">47.3041E</geoCoordinate>
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, 2070m, 24.i.2012, Leg. Ranarilalatiana, Randriamihaja, 1spp. (Cat. No. NHRS-JLKB000021023) Manjakatompo [Andongolongo][pool near source][MJK12-12],
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<geoCoordinate id="892790B58D98FDE58470797909FAA37A" direction="south" orientation="latitude" precision="5" value="-19.3536">19.3536S</geoCoordinate>
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,
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<geoCoordinate id="70812CFC8DFA8555F3D24692DADDB5DD" direction="east" orientation="longitude" precision="5" value="47.3006">47.3006E</geoCoordinate>
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, 1900m, 24.i.2012, Leg. Ranarilalatiana, Randriamihaja.
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</paragraph>
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</subSubSection>
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<subSubSection id="609D4F52E2CA7319746E9757FF694A74" pageId="5" pageNumber="26" type="diagnosis">
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<paragraph id="9F66E02A61B7414A93096B1F561BECBF" pageId="5" pageNumber="26">
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<pageBreakToken id="F887F2928D721C7F1893BA4491E1B8D8" pageId="5" pageNumber="26" start="start">Diagnosis</pageBreakToken>
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.
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</paragraph>
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<paragraph id="4291BCDB21F89DC293F3CD77159F9D5F" pageId="5" pageNumber="26">
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Pronotum entirely testaceous or with two small dark spots medially (Figure 2
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<normalizedToken id="945F05AA95CE828D0B2AD65BE0A9B90E" originalValue="a–b">a-b</normalizedToken>
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). Black irrorations of elytra somewhat confluent subapically (Figure 2
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<normalizedToken id="79325EAC3CF4B875D0956381791FC9D9" originalValue="a–b">a-b</normalizedToken>
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). Male protarsal claws equally long, evenly curved, apex acute (Figure 5
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<normalizedToken id="8914FECA16DBB82C33B06CD11F2962B1" originalValue="a–b">a-b</normalizedToken>
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). Male mesotarsal claws equally long but posterior claw distinctly thicker than anterior claw (Figure 6
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<normalizedToken id="B5FA35EC7DD3D2580B4AC4E21D324DF4" originalValue="a–b">a-b</normalizedToken>
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). Penis short and robust (Figure 7
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<normalizedToken id="29806BA16E6EAE8C0DFDB122A5A06234" originalValue="a–b">a-b</normalizedToken>
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), Parameres evenly curved and tapering to apex (Figure 7g). 11.5-12.5 mm long.
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</paragraph>
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<caption id="4FBBC568B370E9A22D83FF61C864A21C" pageId="5" pageNumber="26">
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<paragraph id="E68572CF67092822CE6FBB14EEA028D4" pageId="5" pageNumber="26">
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Figure 2. Habitus, dorsal view: A
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<taxonomicName id="751837DDDDCD01AE254E3CD4FFD141A9" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
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holotype (Ankaratra mountains) B
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<taxonomicName id="9457F860B6628E273CE952327DA12014" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
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(Tsaratanana mountains) C
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<taxonomicName id="AF9F83D24AC7630B9A3C9678117CFD6D" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bouvieri" order="Coleoptera" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="bouvieri">Rhantus bouvieri</taxonomicName>
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D
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<taxonomicName id="664342CD8F01ADA516A9E8707594F612" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus latus" order="Coleoptera" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="latus">Rhantus latus</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="7BB8745A35BA865AEB5C1677EE5CAE13" lastPageId="6" lastPageNumber="27" pageId="5" pageNumber="26" type="description">
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<paragraph id="6B7498137FD9EA05A2E6FDEB597CCA7E" pageId="5" pageNumber="26">Description.</paragraph>
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<paragraph id="F584E2135C08066BE60C9AE717308C3B" pageId="5" pageNumber="26">Size: ML 11.5-12.5 mm; MW 5.7-5.9 mm; Lp 1.4 mm; Epb 4.7-4.9 mm; Le 8.4-9.0 mm (n = 5).</paragraph>
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<paragraph id="38D0D5C7BB37C6BF58C7C558531F197B" pageId="5" pageNumber="26">
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Head. Testaceous with black areas posteriorly and inside eyes. Interocular black pattern narrowly separated medially (Figure 2
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<normalizedToken id="49F92B6C9602758366B2584F518080F9" originalValue="a–b">a-b</normalizedToken>
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). Dense reticulation of well impressed meshes, very fine punctuation within and at edges of meshes.
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</paragraph>
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<paragraph id="301D64813F07CD3AFC80986171AB42BB" pageId="5" pageNumber="26">Pronotum. Testaceous to ferrugineous with two small dark spots medially (Figure 2b), which may be absent (Figure 2a). Rim at lateral margin clearly visible to indistinct. Dense reticulation of well impressed meshes, very fine punctuation within and at edges of meshes (Figure 3a).</paragraph>
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<paragraph id="A6BC2CF1A73246D0CC73C11BA6AAD05B" pageId="6" pageNumber="27">
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<pageBreakToken id="F9EBB867A2E545E88952319DFA9FA5CF" pageId="6" pageNumber="27" start="start">Elytra</pageBreakToken>
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. Testaceous with black irrorations, leaving paler sides and yellow sutural lines (Figure 2
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<normalizedToken id="67C1A98C136C77BEB00AA61A39BDE3BC" originalValue="a–b">a-b</normalizedToken>
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). Black irrorations somewhat confluent subapically. Reticulation of polygonal meshes simple anteriorly and double at middle and posteriorly, meshes are somewhat less impressed than on pronotum and larger posteriorly. Very fine punctuation within and at edges of meshes (Figure 3
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<normalizedToken id="0FC6EAA208CE17A69D3E69E19219A7A7" originalValue="b–c">b-c</normalizedToken>
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).
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</paragraph>
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<caption id="3BF0D2E8EB0F9139E918F564435D4758" pageId="6" pageNumber="27">
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<paragraph id="8DE55B21CA9B13B2A0092408D5278F5D" pageId="6" pageNumber="27">
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Figure 3. SEM images of pronotal and elytral microstructure:
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<normalizedToken id="A47B7DF83E0C9A042D30E3FBE307649A" originalValue="A–C">A-C</normalizedToken>
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<taxonomicName id="C538272579AF5A77F7F62F48F138A04E" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
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pronotum (A) anterior part of elytron (B) and posterior part of elytron (C)
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<normalizedToken id="7568CC51E68EB18246C323D0A1353245" originalValue="D–F">D-F</normalizedToken>
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<taxonomicName id="B1DFDB01F0CDCDF2EAA1096636CE6027" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bouvieri" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="bouvieri">Rhantus bouvieri</taxonomicName>
|
||
pronotum (D) anterior part of elytron (E) and posterior part of elytron (F)
|
||
<normalizedToken id="3B4B1AA420D17E69DE2DB75B6FFA1748" originalValue="G–I">G-I</normalizedToken>
|
||
<taxonomicName id="9A41CE2618F792FCF469E0DC9B9E0A76" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus latus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="latus">Rhantus latus</taxonomicName>
|
||
pronotum (G) anterior part of elytron (H) and posterior part of elytron (I).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C1D45CA57DBB411214196E63B0BEA60A" pageId="6" pageNumber="27">Ventral side. Dark brown to black. Epiplura testaceous. Metafemora infuscated (Figure 4a).</paragraph>
|
||
<caption id="0F674140E63CDAA9ACF4124C39066B1F" pageId="6" pageNumber="27">
|
||
<paragraph id="282FD5517A29B48A3FF83E3355E46735" pageId="6" pageNumber="27">
|
||
Figure 4. Habitus, ventral view: A
|
||
<taxonomicName id="9478537F57AB394B3149E7FACFA2EB68" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
|
||
holotype B
|
||
<taxonomicName id="C3167A4EBF47B5D80EDE03293E219267" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bouvieri" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="bouvieri">Rhantus bouvieri</taxonomicName>
|
||
C
|
||
<taxonomicName id="00822B7C3AB29C110D543D931709EAFB" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus latus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="latus">Rhantus latus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B6B3E53D639EE87CB0067C55E66D1424" pageId="6" pageNumber="27">
|
||
Male. Protarsal claws equally long, evenly curved, apex acute (Figure 5
|
||
<normalizedToken id="13E1F5B3BCE9306B5C838A071534C8EB" originalValue="a–b">a-b</normalizedToken>
|
||
). Mesotarsal claws equally long but posterior claw distinctly thicker than anterior claw (Figure 6
|
||
<normalizedToken id="5273E36AEF00F7873E796FDB639E086D" originalValue="a–b">a-b</normalizedToken>
|
||
). Penis short and robust, shape as Figure 7
|
||
<normalizedToken id="BA06771AD2BE30FD1F4F1A02A0942AE1" originalValue="a–b">a-b</normalizedToken>
|
||
. Parameres evenly curved and tapering to apex (Figure 7g).
|
||
</paragraph>
|
||
<caption id="8A9F323B2F6F9809D145D62B36E0D7C6" pageId="6" pageNumber="27">
|
||
<paragraph id="2C7E6D78E4F85AEF7FD70816EE0510F1" pageId="6" pageNumber="27">
|
||
Figure 5. Left protarsal claws:
|
||
<normalizedToken id="F64F2586E4ADFD5BA4FF719C2A4F98A1" originalValue="A–B">A-B</normalizedToken>
|
||
<taxonomicName id="EFB976FF2114F197669AA15722876AB5" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
|
||
,lateral (A) and dorsal (B) view
|
||
<normalizedToken id="684D75C3D68FAC98029195EC63955159" originalValue="C–D">C-D</normalizedToken>
|
||
<taxonomicName id="7F8936668F4BE0A1B36963BDB095C490" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bouvieri" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="bouvieri">Rhantus bouvieri</taxonomicName>
|
||
, lateral (C) and dorsal (D) view
|
||
<normalizedToken id="9BA06BEE63A4AD4705ACFAD729DFC1A1" originalValue="E–F">E-F</normalizedToken>
|
||
<taxonomicName id="4559082A3B5568F4B229954911C1C96C" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus latus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="latus">Rhantus latus</taxonomicName>
|
||
, lateral (E) and dorsal (F) view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="55C872D9A0A3F0EF911240233D618637" pageId="6" pageNumber="27">
|
||
<paragraph id="F970C115DCD04BA203299DE6965C0D04" pageId="6" pageNumber="27">
|
||
Figure 6. Left mesotarsal claws:
|
||
<normalizedToken id="F9944AB509ECEBDBE45964B65AE9772B" originalValue="A–B">A-B</normalizedToken>
|
||
<taxonomicName id="26595D66AA0C8B21B5EE2B412B6E71CC" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
|
||
,lateral (A) and dorsal (B) view
|
||
<normalizedToken id="8ED608B2CD52CF053FF5BE7C2EC6808B" originalValue="C–D">C-D</normalizedToken>
|
||
<taxonomicName id="7F200CE706784AEACF3F20E3CEDFADBE" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bouvieri" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="bouvieri">Rhantus bouvieri</taxonomicName>
|
||
, lateral (C) and dorsal (D) view
|
||
<normalizedToken id="F8ED169B69D93E1C126207336B59858A" originalValue="E–F">E-F</normalizedToken>
|
||
<taxonomicName id="ACD465BFAD7B9F5FE86C4ECCCAAF361F" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus latus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="latus">Rhantus latus</taxonomicName>
|
||
, lateral (E) and dorsal (F) view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="28871E74C80017533B8596DF363353C6" pageId="6" pageNumber="27">
|
||
<paragraph id="C990130F78B87CAFF7D483D3322C8822" pageId="6" pageNumber="27">
|
||
Figure 7. Male genitals, penis in lateral and dorsal view:
|
||
<normalizedToken id="D059534C99B05EA47A60D3C7B64CE518" originalValue="A–B">A-B</normalizedToken>
|
||
<taxonomicName id="AD861D1529414122F293B979904B8CE3" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
|
||
<normalizedToken id="246A4A9A543EF5E29C22222F8A7365C5" originalValue="C–D">C-D</normalizedToken>
|
||
<taxonomicName id="4EED1B52BD4AC3F028D23B50AB2EBB25" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bouvieri" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="bouvieri">Rhantus bouvieri</taxonomicName>
|
||
<normalizedToken id="DDD21C032B2B9B840B127ADD95AD45DA" originalValue="E–F">E-F</normalizedToken>
|
||
<taxonomicName id="DF91C67E10A8DDCCC78FC67073CFE73C" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus latus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="latus">Rhantus latus</taxonomicName>
|
||
. Parameres in lateral view G
|
||
<taxonomicName id="3BA12FAF655F5FCEAF04BBCAF968FE0F" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
|
||
H
|
||
<taxonomicName id="0A7305A0A6A07275A62F57B9F0C7F109" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bouvieri" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="bouvieri">Rhantus bouvieri</taxonomicName>
|
||
I
|
||
<taxonomicName id="E201FDD2A5B801C3B38033EC0020BD99" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus latus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="latus">Rhantus latus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="821BFE7BB5F9A3694FC9509BD4449D4F" pageId="6" pageNumber="27" type="remarks">
|
||
<paragraph id="7C4AE64A4C90A40C685C221D445BAB86" pageId="6" pageNumber="27">Remarks.</paragraph>
|
||
<paragraph id="DD660EFF89552896655B5FC709098DA0" pageId="6" pageNumber="27">
|
||
In 2004 David Lees and Ravomiarana Ranaivosolo collected material of what seemed to be a new
|
||
<taxonomicName id="ADE9E7DBCD8990A5A93F9120CC07EADF" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="genus">Rhantus</taxonomicName>
|
||
species from the mountain massif of Tsaratanana, North Madagascar.
|
||
<taxonomicName id="5DE944DEC58F87063E9496729538C0CC" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
|
||
was described in 2009 based on a single male specimen from Ankaratra mountains, 70km south of Antananarivo in central Madagascar. Despite variation in color and impression of a lateral rim of pronotum, molecular data indicate that the studied material from Ankaratra and Tsaratanana are conspecific or at least not an old enough divergence to be recognised based on COI sequence (genetic distance 0.008-0.011). As also male tarsal claws, aedeagus and parameres were identical, we consider the specimens from Tsaratanana mountains conspecific with the holotype of
|
||
<taxonomicName id="71CC3FDF8A936D45A8E5A9A5DA24E12E" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus manjakatompo" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="manjakatompo">Rhantus manjakatompo</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04B7C870C1270F570DE09473B4F926A1" pageId="6" pageNumber="27" type="habitat">
|
||
<paragraph id="F5A43F9ED9D9CC522BAB18ED07ED2CB4" pageId="6" pageNumber="27">Habitat.</paragraph>
|
||
<paragraph id="D434BC83D8AFA139D170C958922698A4" pageId="6" pageNumber="27">Associated with sources and streams, and surrounding water assemblages like nearby pools and marshes at altitudes between 1700 to 2070 m a.s.l. In Manjakatompo, the species was most numerous at elevations above 2000 m.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E233D6A748AAC2E2CF1ACC5C3B053FA6" pageId="7" pageNumber="28" type="distribution">
|
||
<paragraph id="BED4133C402DB3E691464A8318F4B7C8" pageId="7" pageNumber="28">
|
||
<pageBreakToken id="909752299B6103B0DE6E8216F89D2066" pageId="7" pageNumber="28" start="start">Distribution</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C2EED7EED18C23996A3DBE43699C2F2E" pageId="7" pageNumber="28">Endemic to Madagascar. Only known from Tsaratanana mountains and Manjakatompo forestry station in the Ankaratra mountains (Figure 8).</paragraph>
|
||
<caption id="AC90D1520A54FF1A3423AE9B95A1CEBF" pageId="7" pageNumber="28">
|
||
<paragraph id="B3826DF45804440FE613DBEC6708DBD8" pageId="7" pageNumber="28">
|
||
Figure 8. Distribution and all known records of the three
|
||
<taxonomicName id="C3D0EEA23BB78815A87E375FC3E3CE1C" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus" order="Coleoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="genus">Rhantus</taxonomicName>
|
||
species of Madagascar with a special emphasis on Manjakatompo. Inset map of Manjakatompo adapted from
|
||
<bibRefCitation id="2E01448DD883B7F8207800F7E27B5968" pageId="7" pageNumber="28">FTM (1995)</bibRefCitation>
|
||
Extent of natural forest left may have changed since the map was made.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |