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<document id="C777021068E9F197FACFC199470C2946" ID-DOI="10.11646/zootaxa.4803.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3909077" ID-ZooBank="7BEC4D1E-A021-4A10-8E17-E7732E5193A4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593158411237" checkinUser="plazi" docAuthor="Al-Kandari, Manal, Anker, Arthur, Hussain, Sumaiah, Al-Yassen, Sharifa, Sattari, Zainab &amp; Grave, Sammy De" docDate="2020" docId="631687EAFF9FFFA2E2C27B1AF249FD02" docLanguage="en" docName="zootaxa.4803.2.2.pdf" docOrigin="Zootaxa 4803 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alpheus macrodactylus Ortmann 1890" docType="treatment" docVersion="5" lastPageNumber="256" masterDocId="9F2FFF92FF9BFFA7E2557C2FF317FFCD" masterDocTitle="New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda)" masterLastPageNumber="280" masterPageNumber="251" pageNumber="255" updateTime="1698776443936" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D9B974575D91BCBAF2CF7DE14112C893">New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda)</mods:title>
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<mods:namePart id="16DA5271DD3678AE4B9521B9B996845F">Al-Kandari, Manal</mods:namePart>
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<mods:namePart id="A4EF047BF841441D3772378CB53022EC">Anker, Arthur</mods:namePart>
<mods:affiliation id="8888AFEDA2CAB26C02D67FBD95E1656B">Universidade Federal de Goiás, Campus Samambaia, Instituto de Ciências Biológicas-ICB- 5. Av. Esperança, s / n., 74690 - 900, Goiâ-</mods:affiliation>
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<mods:namePart id="3ED309B3E5A1FE542E16C8CFA31290A6">Hussain, Sumaiah</mods:namePart>
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<mods:namePart id="DBD047E1131540CF2E26E1DF9AC3D0D5">Al-Yassen, Sharifa</mods:namePart>
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<mods:namePart id="E0979E1AF1C29C19BE102C04DE6E2C1F">Sattari, Zainab</mods:namePart>
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<mods:namePart id="FF2DB81407768BF384E6735B723C32DD">Grave, Sammy De</mods:namePart>
<mods:affiliation id="0860E2F2FD5E2D1A5FABE1B77886447A">Oxford University Museum of Natural History, Parks Road, Oxford, OX 1 3 PW, United Kingdom https: // orcid. org / 0000 - 0002 - 2437 - 2445</mods:affiliation>
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<mods:date id="A098C815C271A8F1B63CA465FAF6804E">2020</mods:date>
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<mods:number id="23D78F345E2DC1FCFBDE4291CD1F1116">4803</mods:number>
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<heading id="B0488190FF9FFFA3E2C27B1AF16FF882" bold="true" box="[151,632,1845,1872]" fontSize="11" level="1" pageId="4" pageNumber="255" reason="1">
<taxonomicName id="2CBF4D7FFF9FFFA3E2C27B1AF16FF882" ID-CoL="85LL5" authority="Ortmann, 1890" authorityName="Ortmann" authorityYear="1890" box="[151,632,1845,1872]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="macrodactylus">
<emphasis id="D9CBEAEEFF9FFFA3E2C27B1AF16FF882" bold="true" box="[151,632,1845,1872]" pageId="4" pageNumber="255">
<emphasis id="D9CBEAEEFF9FFFA3E2C27B1AF2A5F89D" bold="true" box="[151,434,1845,1872]" italics="true" pageId="4" pageNumber="255">Alpheus macrodactylus</emphasis>
<bibRefCitation id="8F2E4B0DFF9FFFA3E3EC7B1AF16FF882" author="Ortmann, A." box="[441,632,1845,1871]" pageId="4" pageNumber="255" pagination="437 - 542" refId="ref16180" refString="Ortmann, A. (1890) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung dr von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. I. Theil. Die Unterordnung Natantia (Boas) (Abtheilungen: Penaeidae und Eucyphidea = Caridae der Autoren). Zoologische Jahrbucher. Abtheilung fur Systematik, Geographie und Biologie der Thiere, 5, 437 - 542." type="journal article" year="1890">Ortmann, 1890</bibRefCitation>
</emphasis>
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(
<figureCitation id="73842A79FF9FFFA3E2F57B76F3F5F8BE" box="[160,226,1881,1907]" captionStart="FIGURE 3" captionStartId="5.[151,250,1586,1611]" captionTargetBox="[151,1435,741,1561]" captionTargetId="figure@5.[151,1435,741,1561]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus macrodactylus Ortmann, 1890: male (missing minor cheliped) from Failaka Island, Kuwait (OUMNH.ZC. 2020.01.019), dorsal view. Photograph by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.3909083" httpUri="https://zenodo.org/record/3909083/files/figure.png" pageId="4" pageNumber="255">Fig. 3</figureCitation>
)
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<treatmentCitation id="6A1E10EDFF9FFFA3E2C27BB2F147F87B" author="Ortmann, A." box="[151,592,1949,1974]" page="473" pageId="4" pageNumber="255" year="1890">
<taxonomicName id="2CBF4D7FFF9FFFA3E2C27BB2F147F87B" ID-CoL="85LL5" authority="Ortmann 1890: 473" authorityName="Ortmann" authorityPageNumber="473" authorityYear="1890" box="[151,592,1949,1974]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="macrodactylus">
<emphasis id="D9CBEAEEFF9FFFA3E2C27BB2F294F878" box="[151,387,1949,1973]" italics="true" pageId="4" pageNumber="255">Alpheus macrodactylus</emphasis>
<bibRefCitation id="8F2E4B0DFF9FFFA3E3DC7BB1F147F87B" author="Ortmann, A." box="[393,592,1950,1974]" pageId="4" pageNumber="255" pagination="437 - 542" refId="ref16180" refString="Ortmann, A. (1890) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung dr von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. I. Theil. Die Unterordnung Natantia (Boas) (Abtheilungen: Penaeidae und Eucyphidea = Caridae der Autoren). Zoologische Jahrbucher. Abtheilung fur Systematik, Geographie und Biologie der Thiere, 5, 437 - 542." type="journal article" year="1890">Ortmann 1890: 473</bibRefCitation>
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, pl. 36, fig.
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;
<treatmentCitation id="6A1E10EDFF9FFFA3E14A7BB1F0C6F87B" author="Pearson, J." box="[799,977,1949,1974]" page="87" pageId="4" pageNumber="255" year="1905">
<bibRefCitation id="8F2E4B0DFF9FFFA3E14A7BB1F0C6F87B" author="Pearson, J." box="[799,977,1949,1974]" pageId="4" pageNumber="255" pagination="65 - 92" refId="ref16460" refString="Pearson, J. (1905). Report on the Macrura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.), Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with supplementary reports upon the marine biology of Ceylon. The Royal Society, London, pp. 65 - 92." type="book chapter" year="1905">Pearson 1905: 87</bibRefCitation>
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;
<treatmentCitation id="6A1E10EDFF9FFFA3E18E7BB1F7EDF87B" author="Banner, D. M. &amp; Banner, A. H." box="[987,1274,1950,1974]" page="210" pageId="4" pageNumber="255" year="1982">
<bibRefCitation id="8F2E4B0DFF9FFFA3E18E7BB1F7EDF87B" author="Banner, D. M. &amp; Banner, A. H." box="[987,1274,1950,1974]" pageId="4" pageNumber="255" pagination="1 - 357" refId="ref12237" refString="Banner, D. M. &amp; Banner, A. H. (1982) The Alpheid Shrimp of Australia, Part III: The remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982: 210</bibRefCitation>
</treatmentCitation>
, fig. 65;
<treatmentCitation id="6A1E10EDFF9FFFA3E7067BB1F29AF819" author="Banner, A. H. &amp; Banner, D. M." page="44" pageId="4" pageNumber="255" year="1983">
<bibRefCitation id="8F2E4B0DFF9FFFA3E7067BB1F29AF819" author="Banner, A. H. &amp; Banner, D. M." pageId="4" pageNumber="255" pagination="1 - 164" refId="ref12098" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983: 44</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6A1E10EDFF9FFFA3E3C27B94F1D6F819" ID-DOI="http://doi.org/10.5281/zenodo.5255509" ID-Zenodo-Dep="5255509" author="Naderloo, R. &amp; Turkay, M." box="[407,705,1979,2004]" httpUri="http://treatment.plazi.org/id/E177585FC60F1B3AFF3CF9E3FCE0F805" page="10" pageId="4" pageNumber="255" year="2012">
<bibRefCitation id="8F2E4B0DFF9FFFA3E3C27B94F1D6F819" author="Naderloo, R. &amp; Turkay, M." box="[407,705,1979,2004]" pageId="4" pageNumber="255" pagination="1 - 67" refId="ref15707" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012: 10</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="A3A56577FF9EFFA2E2C27CB7F734FE46" pageId="5" pageNumber="256" type="materials_examined">
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<emphasis id="D9CBEAEEFF9EFFA2E2C27CB7F26DFF7E" bold="true" box="[151,378,152,179]" pageId="5" pageNumber="256">Material examined</emphasis>
.
<materialsCitation id="5BD73CA1FF9EFFA2E3DD7CB6F18CFF36" collectingDate="2019-10-03" collectionCode="OUMNH, ZC" collectorName="A. Anker &amp; S. De Grave" country="Kuwait" latitude="29.39015" location="Failaka Island" longLatPrecision="1" longitude="48.398785" pageId="5" pageNumber="256" specimenCount="1" specimenCount-male="1">
<collectingCountry id="93A8766CFF9EFFA2E3DD7CB6F2C8FF7E" box="[392,479,153,179]" name="Kuwait" pageId="5" pageNumber="256">Kuwait</collectingCountry>
:
<specimenCount id="FDB9FD75FF9EFFA2E3BC7CB6F123FF7E" box="[489,564,153,179]" pageId="5" pageNumber="256" type="male">1 male</specimenCount>
(minor cheliped missing),
<collectionCode id="8DAEAE39FF9EFFA2E1317CB6F0C3FF7E" box="[868,980,153,179]" pageId="5" pageNumber="256">OUMNH</collectionCode>
.
<collectionCode id="8DAEAE39FF9EFFA2E1827CB6F0EBFF7E" box="[983,1020,153,179]" country="Austria" name="Zoological Collection, University of Vienna" pageId="5" pageNumber="256" type="University or college">ZC</collectionCode>
. 2020.01.019,
<location id="EE606027FF9EFFA2E6F47CB6F656FF7E" LSID="urn:lsid:plazi:treatment:631687EAFF9FFFA2E2C27B1AF249FD02:EE606027FF9EFFA2E6F47CB6F656FF7E" box="[1185,1345,153,179]" country="Kuwait" latitude="29.39015" longLatPrecision="1" longitude="48.398785" name="Failaka Island" pageId="5" pageNumber="256">Failaka Island</location>
, eastern side,
<geoCoordinate id="8E8B503BFF9EFFA2E2807C92F271FF1A" box="[213,358,189,215]" degrees="29" direction="north" minutes="23.409" orientation="latitude" pageId="5" pageNumber="256" precision="1" value="29.39015">29°23.409N</geoCoordinate>
<geoCoordinate id="8E8B503BFF9EFFA2E3257C92F11DFF1A" box="[368,522,189,215]" degrees="48" direction="east" minutes="23.927" orientation="longitude" pageId="5" pageNumber="256" precision="1" value="48.398785">48° 23.927E</geoCoordinate>
, mixed rocky-muddy intertidal, burrows in mud, suction (yabby) pump, leg.
<collectorName id="464A532AFF9EFFA2E7D47C92F3CAFF36" pageId="5" pageNumber="256">A. Anker</collectorName>
&amp;
<collectorName id="464A532AFF9EFFA2E3547CCEF29AFF36" box="[257,397,225,251]" pageId="5" pageNumber="256">S. De Grave</collectorName>
,
<date id="9F01103CFF9EFFA2E3CC7CCEF11FFF36" box="[409,520,225,251]" pageId="5" pageNumber="256" value="2019-10-03">
<collectingDate id="8F45E9D4FF9EFFA2E3CC7CCEF11FFF36" box="[409,520,225,251]" pageId="5" pageNumber="256" value="2019-10-03">03.x.2019</collectingDate>
</date>
[KUW-073]
</materialsCitation>
.
</paragraph>
<paragraph id="EB0036FCFF9EFFA2E2927D2BF734FE46" blockId="5.[151,1437,152,719]" pageId="5" pageNumber="256">
<emphasis id="D9CBEAEEFF9EFFA2E2927D2BF2A8FED2" bold="true" box="[199,447,260,287]" pageId="5" pageNumber="256">General distribution</emphasis>
. Apparently widely distributed in the Indo-West Pacific, with records from the Arabi- an Gulf (
<collectingCountry id="93A8766CFF9EFFA2E2AA7D06F241FE8E" box="[255,342,297,323]" name="Kuwait" pageId="5" pageNumber="256">Kuwait</collectingCountry>
,
<collectingCountry id="93A8766CFF9EFFA2E3367D06F283FE8E" box="[355,404,297,323]" name="Iran" pageId="5" pageNumber="256">Iran</collectingCountry>
), East Africa (
<collectingCountry id="93A8766CFF9EFFA2E06F7D06F18DFE8E" box="[570,666,297,323]" name="Somalia" pageId="5" pageNumber="256">Somalia</collectingCountry>
,
<collectingCountry id="93A8766CFF9EFFA2E0FD7D06F1E2FE8E" box="[680,757,297,323]" name="Kenya" pageId="5" pageNumber="256">Kenya</collectingCountry>
),
<collectingCountry id="93A8766CFF9EFFA2E15F7D06F084FE8E" box="[778,915,297,323]" name="Madagascar" pageId="5" pageNumber="256">Madagascar</collectingCountry>
,
<collectingCountry id="93A8766CFF9EFFA2E1F47D06F703FE8E" box="[929,1044,297,323]" name="Sri Lanka" pageId="5" pageNumber="256">Sri Lanka</collectingCountry>
,
<collectingCountry id="93A8766CFF9EFFA2E6747D06F786FE8E" box="[1057,1169,297,323]" name="Indonesia" pageId="5" pageNumber="256">Indonesia</collectingCountry>
,
<collectingCountry id="93A8766CFF9EFFA2E6C87D06F7EBFE8E" box="[1181,1276,297,323]" name="Vietnam" pageId="5" pageNumber="256">Vietnam</collectingCountry>
and
<collectingCountry id="93A8766CFF9EFFA2E7637D06F68AFE8E" box="[1334,1437,297,323]" name="Australia" pageId="5" pageNumber="256">Australia</collectingCountry>
(
<collectingRegion id="297BF81EFF9EFFA2E2CA7D62F23FFEAA" box="[159,296,333,359]" country="Australia" name="Queensland" pageId="5" pageNumber="256">Queensland</collectingRegion>
,
<collectingRegion id="297BF81EFF9EFFA2E3617D62F2E8FEA5" box="[308,511,333,360]" country="Australia" name="New South Wales" pageId="5" pageNumber="256">New South Wales</collectingRegion>
) (
<bibRefCitation id="8F2E4B0DFF9EFFA2E0407D62F1AAFEAA" author="Ortmann, A." box="[533,701,333,359]" pageId="5" pageNumber="256" pagination="437 - 542" refId="ref16180" refString="Ortmann, A. (1890) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung dr von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. I. Theil. Die Unterordnung Natantia (Boas) (Abtheilungen: Penaeidae und Eucyphidea = Caridae der Autoren). Zoologische Jahrbucher. Abtheilung fur Systematik, Geographie und Biologie der Thiere, 5, 437 - 542." type="journal article" year="1890">Ortmann 1890</bibRefCitation>
; De Man 1898;
<bibRefCitation id="8F2E4B0DFF9EFFA2E1207D62F706FEAA" author="Pearson, J." box="[885,1041,332,359]" pageId="5" pageNumber="256" pagination="65 - 92" refId="ref16460" refString="Pearson, J. (1905). Report on the Macrura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.), Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with supplementary reports upon the marine biology of Ceylon. The Royal Society, London, pp. 65 - 92." type="book chapter" year="1905">Pearson 1905</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF9EFFA2E6487D62F630FEAA" author="Banner, D. M. &amp; Banner, A. H." box="[1053,1319,333,359]" pageId="5" pageNumber="256" pagination="1 - 357" refId="ref12237" refString="Banner, D. M. &amp; Banner, A. H. (1982) The Alpheid Shrimp of Australia, Part III: The remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF9EFFA2E7617D62F250FE46" author="Naderloo, R. &amp; Turkay, M." pageId="5" pageNumber="256" pagination="1 - 67" refId="ref15707" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012</bibRefCitation>
; exact range uncertain due to confusion with closely related taxa.
</paragraph>
</subSubSection>
<subSubSection id="A3A56577FF9EFFA2E2927DBAF267FE1E" pageId="5" pageNumber="256" type="biology_ecology">
<paragraph id="EB0036FCFF9EFFA2E2927DBAF267FE1E" blockId="5.[151,1437,152,719]" pageId="5" pageNumber="256">
<emphasis id="D9CBEAEEFF9EFFA2E2927DBAF231FE62" bold="true" box="[199,294,405,431]" pageId="5" pageNumber="256">Ecology</emphasis>
. Intertidal and shallow subtidal mud flats, sometimes close to mixed rocky-muddy beaches, also in mangrove swamps.
</paragraph>
</subSubSection>
<subSubSection id="A3A56577FF9EFFA2E2927DF2F249FD02" pageId="5" pageNumber="256" type="discussion">
<paragraph id="EB0036FCFF9EFFA2E2927DF2F249FD02" blockId="5.[151,1437,152,719]" pageId="5" pageNumber="256">
<emphasis id="D9CBEAEEFF9EFFA2E2927DF2F223FE3A" bold="true" box="[199,308,477,503]" pageId="5" pageNumber="256">Remarks</emphasis>
.
<taxonomicName id="2CBF4D7FFF9EFFA2E3147DF2F152FE3A" authorityName="Ortmann" authorityYear="1890" box="[321,581,477,503]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="macrodactylus">
<emphasis id="D9CBEAEEFF9EFFA2E3147DF2F152FE3A" box="[321,581,477,503]" italics="true" pageId="5" pageNumber="256">Alpheus macrodactylus</emphasis>
</taxonomicName>
is part of the larger
<taxonomicName id="2CBF4D7FFF9EFFA2E14A7DF2F799FE3A" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[799,1166,476,503]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="malabaricus">
<emphasis id="D9CBEAEEFF9EFFA2E14A7DF2F0DEFE3A" box="[799,969,477,503]" italics="true" pageId="5" pageNumber="256">A. malabaricus</emphasis>
(
<bibRefCitation id="8F2E4B0DFF9EFFA2E1837DF2F791FE3A" author="Fabricius, J. C." box="[982,1158,476,503]" pageId="5" pageNumber="256" refId="ref13793" refString="Fabricius, J. C. (1775) Systema Entomologiae, sistens Insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. In Officina Libraria Kortii, Flensbergi et Lipsiae, 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510" type="book" year="1775">Fabricius, 1775</bibRefCitation>
)
</taxonomicName>
species complex, which needs a thorough taxonomic revision (
<bibRefCitation id="8F2E4B0DFF9EFFA2E0687E2EF1C1FDD1" author="Anker, A." box="[573,726,513,540]" pageId="5" pageNumber="256" pagination="57 - 72" refId="ref11322" refString="Anker, A. (2001 b) Two new species of snapping shrimps from the Indo-Pacific, with remarks on colour patterns and sibling species in Alpheidae (Crustacea: Caridea). Raffles Bulletin of Zoology, 49, 57 - 72." type="journal article" year="2001">Anker 2001b</bibRefCitation>
). The species was previously known from the Arabian Gulf only by a record based on a single specimen from
<collectingCountry id="93A8766CFF9EFFA2E0F77E0AF056FDF2" box="[674,833,549,575]" name="Qeshm Island" pageId="5" pageNumber="256">Qeshm Island</collectingCountry>
,
<collectingCountry id="93A8766CFF9EFFA2E11B7E0AF06CFDF2" box="[846,891,549,575]" name="Iran" pageId="5" pageNumber="256">Iran</collectingCountry>
(
<bibRefCitation id="8F2E4B0DFF9EFFA2E1D97E0AF7A4FDF2" author="Naderloo, R. &amp; Turkay, M." box="[908,1203,549,575]" pageId="5" pageNumber="256" pagination="1 - 67" refId="ref15707" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012</bibRefCitation>
). The identity of the Kuwaiti specimen (
<figureCitation id="73842A79FF9EFFA2E3257E66F2A6FDAE" box="[368,433,585,611]" captionStart="FIGURE 3" captionStartId="5.[151,250,1586,1611]" captionTargetBox="[151,1435,741,1561]" captionTargetId="figure@5.[151,1435,741,1561]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus macrodactylus Ortmann, 1890: male (missing minor cheliped) from Failaka Island, Kuwait (OUMNH.ZC. 2020.01.019), dorsal view. Photograph by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.3909083" httpUri="https://zenodo.org/record/3909083/files/figure.png" pageId="5" pageNumber="256">Fig. 3</figureCitation>
) is somewhat uncertain for (1) it is missing the diagnostic minor cheliped; (2) both second pereiopods appear to have regenerated and therefore present an abnormal ratio of the carpal subdivisions; (3) the proportions of the major chela are somewhat different from those of the Australian specimen illustrated by
<bibRefCitation id="8F2E4B0DFF9EFFA2E71E7EBEF24EFD02" author="Banner, D. M. &amp; Banner, A. H." pageId="5" pageNumber="256" pagination="1 - 357" refId="ref12237" refString="Banner, D. M. &amp; Banner, A. H. (1982) The Alpheid Shrimp of Australia, Part III: The remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner (1982)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>