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<document id="E9B2ED044879D250C15E9233663691B7" ID-DOI="10.11646/zootaxa.4803.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3909077" ID-ZooBank="7BEC4D1E-A021-4A10-8E17-E7732E5193A4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593158411237" checkinUser="plazi" docAuthor="Al-Kandari, Manal, Anker, Arthur, Hussain, Sumaiah, Al-Yassen, Sharifa, Sattari, Zainab &amp; Grave, Sammy De" docDate="2020" docId="631687EAFF95FFB7E2C27A65F626FF36" docLanguage="en" docName="zootaxa.4803.2.2.pdf" docOrigin="Zootaxa 4803 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Upogebia carinicauda" docType="treatment" docVersion="5" lastPageNumber="266" masterDocId="9F2FFF92FF9BFFA7E2557C2FF317FFCD" masterDocTitle="New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda)" masterLastPageNumber="280" masterPageNumber="251" pageNumber="265" updateTime="1698776443936" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B14CF6322E0111A89D0107F44F959947">New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda)</mods:title>
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<mods:namePart id="CB2DDDF65C09A73FD5E4264821C79F5E">Al-Kandari, Manal</mods:namePart>
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<mods:namePart id="E2F55E99310080E794A79085E9336A53">Anker, Arthur</mods:namePart>
<mods:affiliation id="3B955ADBF2D1161C929804FF1FBD0A11">Universidade Federal de Goiás, Campus Samambaia, Instituto de Ciências Biológicas-ICB- 5. Av. Esperança, s / n., 74690 - 900, Goiâ-</mods:affiliation>
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<mods:namePart id="CE6A1335D59C0CCC4E9712FE143057A8">Hussain, Sumaiah</mods:namePart>
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<mods:namePart id="BDC613C2CFDE1BF811CA16F77E8491E0">Al-Yassen, Sharifa</mods:namePart>
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<mods:namePart id="82493E032F1E2158E670F28E0593A69F">Sattari, Zainab</mods:namePart>
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<mods:namePart id="FC423B6C28E5E17280E3BF5BBB8686D7">Grave, Sammy De</mods:namePart>
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<mods:date id="3A4F3958CBE7F022C73794BB2C43E442">2020</mods:date>
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<heading id="B0488190FF95FFA9E2C27A65F16AF9A9" bold="true" box="[151,637,1610,1637]" fontSize="11" level="1" pageId="14" pageNumber="265" reason="1">
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<emphasis id="D9CBEAEEFF95FFA9E2C27A65F16AF9A9" bold="true" box="[151,637,1610,1637]" pageId="14" pageNumber="265">
<emphasis id="D9CBEAEEFF95FFA9E2C27A65F2B4F9A8" bold="true" box="[151,419,1610,1637]" italics="true" pageId="14" pageNumber="265">Upogebia carinicauda</emphasis>
(
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(
<figureCitation id="73842A79FF95FFA9E2F57A41F3E6F944" box="[160,241,1646,1673]" captionStart="FIGURE 10" captionStartId="15.[151,250,1778,1803]" captionTargetBox="[151,1435,272,1752]" captionTargetId="figure@15.[151,1435,272,1752]" captionTargetPageId="15" captionText="FIGURE 10. Upogebia carinicauda (Stimpson, 1860): A—female (carapace slightly damaged) from Failaka Island, Kuwait (OUMNH.ZC. 2020.01.034), lateral view; B—same, sixth pleonite and tail fan, dorsal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.3909097" httpUri="https://zenodo.org/record/3909097/files/figure.png" pageId="14" pageNumber="265">Fig. 10</figureCitation>
)
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<emphasis id="D9CBEAEEFF95FFA9E2C27A9CF242F906" box="[151,341,1715,1739]" italics="true" pageId="14" pageNumber="265">Gebia carinicauda</emphasis>
<bibRefCitation id="8F2E4B0DFF95FFA9E3097A9DF132F906" author="Stimpson, W." box="[348,549,1714,1739]" pageId="14" pageNumber="265" pagination="22 - 47" refId="ref17592" refString="Stimpson, W. (1860) Prodromus descriptionis animalium evertebratorum, quae in Expeditione as Oceanum Pacificum Septentrionalem, a Republic Federata missa, Cadwaladore Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VIII, Crustacea Macrura. Proceedings of the Academy of Natural Sciences of Philadelphia, 1860, 22 - 47." type="journal article" year="1860">Stimpson, 1860: 23</bibRefCitation>
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.
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<treatmentCitationGroup id="CBAF11D2FF95FFA9E2C27AFEF0EAF8C5" pageId="14" pageNumber="265">
<emphasis id="D9CBEAEEFF95FFA9E2C27AFEF294F924" box="[151,387,1745,1769]" italics="true" pageId="14" pageNumber="265">
<taxonomicName id="2CBF4D7FFF95FFA9E2C27AFEF297F924" ID-CoL="301c9808-a854-47c3-b755-e6ae0fcce05d" baseAuthorityName="Stimpson" baseAuthorityYear="1860" box="[151,384,1745,1769]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="carinicauda">Upogebia carinicauda</taxonomicName>
.
</emphasis>
<treatmentCitation id="6A1E10EDFF95FFA9E3C97AFEF1BBF927" author="Sakai, K. &amp; Takeda, M." box="[412,684,1745,1770]" page="204" pageId="14" pageNumber="265" year="1995">
<bibRefCitation id="8F2E4B0DFF95FFA9E3C97AFEF1BBF927" author="Sakai, K. &amp; Takeda, M." box="[412,684,1745,1770]" pageId="14" pageNumber="265" pagination="203 - 210" refId="ref16796" refString="Sakai, K. &amp; Takeda, M. (1995) New records of two species of decapod crustaceans from Amami-Oshima Island, the Northern Ryukyu Islands, Japan. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21, 203 - 210." type="journal article" year="1995">Sakai &amp; Takeda 1995: 204</bibRefCitation>
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, figs. 13;
<treatmentCitation id="6A1E10EDFF95FFA9E14B7AFEF0AEF927" author="Sakai, K." box="[798,953,1745,1770]" page="98" pageId="14" pageNumber="265" year="2006">
<bibRefCitation id="8F2E4B0DFF95FFA9E14B7AFEF0AEF927" author="Sakai, K." box="[798,953,1745,1770]" pageId="14" pageNumber="265" pagination="1 - 185" refId="ref16726" refString="Sakai, K. (2006) Upogebiidae of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 6, 1 - 185." type="journal article" year="2006">Sakai 2006: 98</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6A1E10EDFF95FFA9E1917AFEF7C8F927" author="Sepahvand, V. &amp; Sari, A." box="[964,1247,1745,1770]" page="44" pageId="14" pageNumber="265" year="2010">
<bibRefCitation id="8F2E4B0DFF95FFA9E1917AFEF7C8F927" author="Sepahvand, V. &amp; Sari, A." box="[964,1247,1745,1770]" pageId="14" pageNumber="265" pagination="43 - 52" refId="ref17257" refString="Sepahvand, V. &amp; Sari, A. (2010) Taxonomy and geographical distribution of intertidal thalassinidean shrimps (Crustacean: Decapoda) from the Qeshm Island, Persian Gulf. Journal of Science University of Tehran, 36, 43 - 52. [in Persian]" type="journal article" year="2010">Sepahvand &amp; Sari 2010: 44</bibRefCitation>
</treatmentCitation>
, fig. 2;
<treatmentCitation id="6A1E10EDFF95FFA9E7787AFEF258F8C5" author="Sepahvand, V. &amp; Sari, A. &amp; Salehi, H. &amp; Nabavi, S. M. B. &amp; Ghorbanzadeh, S. G." page="5" pageId="14" pageNumber="265" year="2013">
<bibRefCitation id="8F2E4B0DFF95FFA9E7787AFEF258F8C5" author="Sepahvand, V. &amp; Sari, A. &amp; Salehi, H. &amp; Nabavi, S. M. B. &amp; Ghorbanzadeh, S. G." pageId="14" pageNumber="265" pagination="999 - 1008" refId="ref17380" refString="Sepahvand, V., Sari, A., Salehi, H., Nabavi, S. M. B. &amp; Ghorbanzadeh, S. G. (2013) Littoral mud shrimps (Decapoda: Gebiidea &amp; Axiidea) of the Persian Gulf and Gulf of Oman, Iran. Journal of the Marine Biological Association of the United Kingdom, 93, 999 - 1008. https: // doi. org / 10.1017 / S 0025315412001361" type="journal article" year="2013">
Sepahvand
<emphasis id="D9CBEAEEFF95FFA9E2927ADEF3EDF8C5" box="[199,250,1776,1800]" italics="true" pageId="14" pageNumber="265">et al.</emphasis>
2013: 5
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;
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<bibRefCitation id="8F2E4B0DFF95FFA9E30F7AC0F193F8C5" author="Naderloo, R. &amp; Turkay, M." box="[346,644,1775,1800]" pageId="14" pageNumber="265" pagination="1 - 67" refId="ref15707" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012: 18</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6A1E10EDFF95FFA9E0DA7AC0F0B2F8C5" author="Sepahvand, V. &amp; Sari, A. &amp; Tudge, C. &amp; Bolouki, M." box="[655,933,1775,1800]" page="138" pageId="14" pageNumber="265" year="2014">
<bibRefCitation id="8F2E4B0DFF95FFA9E0DA7AC0F0B2F8C5" author="Sepahvand, V. &amp; Sari, A. &amp; Tudge, C. &amp; Bolouki, M." box="[655,933,1775,1800]" pageId="14" pageNumber="265" pagination="137 - 144" refId="ref17462" refString="Sepahvand, V., Sari, A., Tudge, C. &amp; Bolouki, M. (2014) A study of burrow morphology in representative axiidean and gebiidean mud shrimps, from the Persian Gulf and Gulf of Oman, Iran. Nauplius, 22, 137 - 144. https: // doi. org / 10.1590 / S 0104 - 64972014000200007" type="journal article" year="2014">
Sepahvand
<emphasis id="D9CBEAEEFF95FFA9E1517ADEF020F8C5" box="[772,823,1776,1800]" italics="true" pageId="14" pageNumber="265">et al.</emphasis>
2014: 138
</bibRefCitation>
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, fig. 2B.
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<emphasis id="D9CBEAEEFF95FFA9E2C27B19F26FF89C" bold="true" box="[151,376,1846,1873]" pageId="14" pageNumber="265">Material examined</emphasis>
.
<materialsCitation id="5BD73CA1FF95FFA9E3D07B19F25BF854" collectingDate="2019-10-03" collectionCode="OUMNH, ZC" collectorName="A. Anker" country="Kuwait" elevation="3" latitude="29.39015" location="Failaka Island" longLatPrecision="1" longitude="48.398785" pageId="14" pageNumber="265" specimenCount="1" specimenCount-female="1">
<collectingCountry id="93A8766CFF95FFA9E3D07B19F2CBF89D" box="[389,476,1846,1872]" name="Kuwait" pageId="14" pageNumber="265">Kuwait</collectingCountry>
:
<specimenCount id="FDB9FD75FF95FFA9E3B17B18F150F89D" box="[484,583,1846,1873]" pageId="14" pageNumber="265" type="female">1 female</specimenCount>
,
<collectionCode id="8DAEAE39FF95FFA9E0047B18F1D6F89C" box="[593,705,1847,1873]" pageId="14" pageNumber="265">OUMNH</collectionCode>
.
<collectionCode id="8DAEAE39FF95FFA9E0967B18F1FFF89C" box="[707,744,1847,1873]" country="Austria" name="Zoological Collection, University of Vienna" pageId="14" pageNumber="265" type="University or college">ZC</collectionCode>
. 2020.01.034,
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,
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, shallow subtidal pool (
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at low tide), muddy bottom close to rocky platform, in burrow, leg.
<collectorName id="464A532AFF95FFA9E6E27B74F637F8B9" box="[1207,1312,1882,1909]" pageId="14" pageNumber="265">A. Anker</collectorName>
,
<date id="9F01103CFF95FFA9E7797B74F68BF8B8" box="[1324,1436,1883,1909]" pageId="14" pageNumber="265" value="2019-10-03">
<collectingDate id="8F45E9D4FF95FFA9E7797B74F68BF8B8" box="[1324,1436,1883,1909]" pageId="14" pageNumber="265" value="2019-10-03">03.x.2019</collectingDate>
</date>
[fcn KUW-061]
</materialsCitation>
.
</paragraph>
<paragraph id="EB0036FCFF95FFA9E2927B8DF050F82C" blockId="14.[151,1436,1846,2017]" pageId="14" pageNumber="265">
<emphasis id="D9CBEAEEFF95FFA9E2927B8DF2ABF870" bold="true" box="[199,444,1954,1981]" pageId="14" pageNumber="265">General distribution</emphasis>
. Indo-West Pacific from the Red Sea, Arabian Gulf and
<collectingCountry id="93A8766CFF95FFA9E67E7B8DF7A5F871" box="[1067,1202,1954,1980]" name="Madagascar" pageId="14" pageNumber="265">Madagascar</collectingCountry>
to
<collectingCountry id="93A8766CFF95FFA9E6807B8CF60FF870" box="[1237,1304,1955,1981]" name="Japan" pageId="14" pageNumber="265">Japan</collectingCountry>
,
<collectingCountry id="93A8766CFF95FFA9E7777B8CF679F870" box="[1314,1390,1955,1981]" name="Samoa" pageId="14" pageNumber="265">Samoa</collectingCountry>
and
<collectingCountry id="93A8766CFF95FFA9E2C27BE9F3E9F82D" box="[151,254,1990,2016]" name="Australia" pageId="14" pageNumber="265">Australia</collectingCountry>
(
<bibRefCitation id="8F2E4B0DFF95FFA9E3597BE9F298F82D" author="Sakai, K." box="[268,399,1990,2016]" pageId="14" pageNumber="265" pagination="1 - 185" refId="ref16726" refString="Sakai, K. (2006) Upogebiidae of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 6, 1 - 185." type="journal article" year="2006">Sakai 2006</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF95FFA9E3CF7BE9F181F82C" author="Sepahvand, V. &amp; Sari, A. &amp; Salehi, H. &amp; Nabavi, S. M. B. &amp; Ghorbanzadeh, S. G." box="[410,662,1990,2017]" pageId="14" pageNumber="265" pagination="999 - 1008" refId="ref17380" refString="Sepahvand, V., Sari, A., Salehi, H., Nabavi, S. M. B. &amp; Ghorbanzadeh, S. G. (2013) Littoral mud shrimps (Decapoda: Gebiidea &amp; Axiidea) of the Persian Gulf and Gulf of Oman, Iran. Journal of the Marine Biological Association of the United Kingdom, 93, 999 - 1008. https: // doi. org / 10.1017 / S 0025315412001361" type="journal article" year="2013">
Sepahvand
<emphasis id="D9CBEAEEFF95FFA9E0497BE8F143F82D" box="[540,596,1990,2016]" italics="true" pageId="14" pageNumber="265">et al.</emphasis>
2013
</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="A3A56577FF94FFA8E2927CB6F3F2FF36" pageId="15" pageNumber="266" type="biology_ecology">
<paragraph id="EB0036FCFF94FFA8E2927CB6F3F2FF36" blockId="15.[151,1436,152,251]" pageId="15" pageNumber="266">
<emphasis id="D9CBEAEEFF94FFA8E2927CB6F231FF7E" bold="true" box="[199,294,153,179]" pageId="15" pageNumber="266">Ecology</emphasis>
. Largely intertidal and shallow subtidal; mixed rocky-sandy shores with mud and shell fragments; typically, in complex ramified burrows in clay, mud/sand or limestone rocks (
<bibRefCitation id="8F2E4B0DFF94FFA8E1A97C93F7EEFF1A" author="Sepahvand, V. &amp; Sari, A. &amp; Salehi, H. &amp; Nabavi, S. M. B. &amp; Ghorbanzadeh, S. G." box="[1020,1273,188,215]" pageId="15" pageNumber="266" pagination="999 - 1008" refId="ref17380" refString="Sepahvand, V., Sari, A., Salehi, H., Nabavi, S. M. B. &amp; Ghorbanzadeh, S. G. (2013) Littoral mud shrimps (Decapoda: Gebiidea &amp; Axiidea) of the Persian Gulf and Gulf of Oman, Iran. Journal of the Marine Biological Association of the United Kingdom, 93, 999 - 1008. https: // doi. org / 10.1017 / S 0025315412001361" type="journal article" year="2013">
Sepahvand
<emphasis id="D9CBEAEEFF94FFA8E62A7C92F7A6FF1B" box="[1151,1201,188,214]" italics="true" pageId="15" pageNumber="266">et al</emphasis>
. 2013
</bibRefCitation>
,
<bibRefCitation id="8F2E4B0DFF94FFA8E7507C92F656FF1A" author="Sepahvand, V. &amp; Sari, A. &amp; Tudge, C. &amp; Bolouki, M." box="[1285,1345,189,215]" pageId="15" pageNumber="266" pagination="137 - 144" refId="ref17462" refString="Sepahvand, V., Sari, A., Tudge, C. &amp; Bolouki, M. (2014) A study of burrow morphology in representative axiidean and gebiidean mud shrimps, from the Persian Gulf and Gulf of Oman, Iran. Nauplius, 22, 137 - 144. https: // doi. org / 10.1590 / S 0104 - 64972014000200007" type="journal article" year="2014">2014</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<caption id="BFC06674FF94FFA8E2C27ADDF65DF8E2" ID-DOI="http://doi.org/10.5281/zenodo.3909097" ID-Zenodo-Dep="3909097" httpUri="https://zenodo.org/record/3909097/files/figure.png" pageId="15" pageNumber="266" startId="15.[151,250,1778,1803]" targetBox="[151,1435,272,1752]" targetPageId="15">
<paragraph id="EB0036FCFF94FFA8E2C27ADDF65DF8E2" blockId="15.[151,1437,1778,1839]" pageId="15" pageNumber="266">
<emphasis id="D9CBEAEEFF94FFA8E2C27ADDF20CF8C6" bold="true" box="[151,283,1778,1803]" pageId="15" pageNumber="266">FIGURE 10</emphasis>
.
<taxonomicName id="2CBF4D7FFF94FFA8E37F7ADDF1D9F8C6" authority="(Stimpson, 1860)" baseAuthorityName="Stimpson" baseAuthorityYear="1860" box="[298,718,1778,1803]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="carinicauda">
<emphasis id="D9CBEAEEFF94FFA8E37F7ADDF107F8C7" box="[298,528,1778,1802]" italics="true" pageId="15" pageNumber="266">Upogebia carinicauda</emphasis>
(
<bibRefCitation id="8F2E4B0DFF94FFA8E04A7ADDF1D1F8C6" author="Stimpson, W." box="[543,710,1778,1803]" pageId="15" pageNumber="266" pagination="22 - 47" refId="ref17592" refString="Stimpson, W. (1860) Prodromus descriptionis animalium evertebratorum, quae in Expeditione as Oceanum Pacificum Septentrionalem, a Republic Federata missa, Cadwaladore Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VIII, Crustacea Macrura. Proceedings of the Academy of Natural Sciences of Philadelphia, 1860, 22 - 47." type="journal article" year="1860">Stimpson, 1860</bibRefCitation>
)
</taxonomicName>
: A—female (carapace slightly damaged) from Failaka Island, Kuwait (OUMNH.ZC. 2020.01.034), lateral view; B—same, sixth pleonite and tail fan, dorsal view. Photographs by A. Anker.
</paragraph>
</caption>
<subSubSection id="A3A56577FF94FFB7E2927B72F626FF36" lastPageId="16" lastPageNumber="267" pageId="15" pageNumber="266" type="discussion">
<paragraph id="EB0036FCFF94FFB7E2927B72F626FF36" blockId="15.[151,1437,1885,2020]" lastBlockId="16.[151,1436,153,251]" lastPageId="16" lastPageNumber="267" pageId="15" pageNumber="266">
<emphasis id="D9CBEAEEFF94FFA8E2927B72F223F8BA" bold="true" box="[199,308,1885,1911]" pageId="15" pageNumber="266">Remarks</emphasis>
. The single specimen from
<collectingCountry id="93A8766CFF94FFA8E03A7B72F1D6F8BA" box="[623,705,1885,1911]" name="Kuwait" pageId="15" pageNumber="266">Kuwait</collectingCountry>
corresponds well to
<taxonomicName id="2CBF4D7FFF94FFA8E1FB7B71F74FF8BA" baseAuthorityName="Stimpson" baseAuthorityYear="1860" box="[942,1112,1885,1911]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="carinicauda">
<emphasis id="D9CBEAEEFF94FFA8E1FB7B71F74FF8BA" box="[942,1112,1885,1911]" italics="true" pageId="15" pageNumber="266">U. carinicauda</emphasis>
</taxonomicName>
, as defined by
<bibRefCitation id="8F2E4B0DFF94FFA8E7507B72F68EF8B5" author="Sakai, K." box="[1285,1433,1885,1912]" pageId="15" pageNumber="266" pagination="1 - 185" refId="ref16726" refString="Sakai, K. (2006) Upogebiidae of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 6, 1 - 185." type="journal article" year="2006">Sakai (2006)</bibRefCitation>
. The main diagnostic features of the species are the dorsal surface of the telson bearing an elevated, sharp, transverse carina that connects to two blunt, lateral, longitudinal carinae (
<figureCitation id="73842A79FF94FFA8E1317B8AF0DAF872" box="[868,973,1957,1983]" captionStart="FIGURE 10" captionStartId="15.[151,250,1778,1803]" captionTargetBox="[151,1435,272,1752]" captionTargetId="figure@15.[151,1435,272,1752]" captionTargetPageId="15" captionText="FIGURE 10. Upogebia carinicauda (Stimpson, 1860): A—female (carapace slightly damaged) from Failaka Island, Kuwait (OUMNH.ZC. 2020.01.034), lateral view; B—same, sixth pleonite and tail fan, dorsal view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.3909097" httpUri="https://zenodo.org/record/3909097/files/figure.png" pageId="15" pageNumber="266">Fig. 10B</figureCitation>
); the presence of four teeth on the distal margin of the rostrum (one of them obsolete in the Kuwaiti specimen) and a single infra-rostral tooth; the presence of a conspicuous sharp tooth on the ventromesial surface of the cheliped propodus; and the dactylus being almost twice as long as the propodus. However, we believe that some synonymies proposed by
<bibRefCitation id="8F2E4B0DFF8BFFB7E62F7C92F61BFF15" author="Sakai, K." box="[1146,1292,189,216]" pageId="16" pageNumber="267" pagination="1 - 185" refId="ref16726" refString="Sakai, K. (2006) Upogebiidae of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 6, 1 - 185." type="journal article" year="2006">Sakai (2006)</bibRefCitation>
, for instance that of
<taxonomicName id="2CBF4D7FFF8BFFB7E2B37CCDF113FF36" authority="Komai, 2005" authorityName="Komai" authorityYear="2005" box="[230,516,224,251]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="rupicola">
<emphasis id="D9CBEAEEFF8BFFB7E2B37CCDF27FFF36" box="[230,360,225,251]" italics="true" pageId="16" pageNumber="267">U. rupicola</emphasis>
<bibRefCitation id="8F2E4B0DFF8BFFB7E33A7CCEF113FF36" author="Komai, T." box="[367,516,224,251]" pageId="16" pageNumber="267" pagination="259 - 268" refId="ref14899" refString="Komai, T. (2005) A new species of Upogebia (Crustacea: Decapoda: Thalassinidea: Upogebiidae) from Okinawa Island, Ryukyu Islands. Species Diversity, 10, 259 - 268. https: // doi. org / 10.12782 / specdiv. 10.259" type="journal article" year="2005">Komai, 2005</bibRefCitation>
</taxonomicName>
, may need a re-assessment (see also Ngoc-Ho 2008;
<bibRefCitation id="8F2E4B0DFF8BFFB7E6037CCEF632FF36" author="Komai, T. &amp; Ravinesh, R. &amp; Riyas, A. &amp; Biju Kumar, A." box="[1110,1317,225,251]" pageId="16" pageNumber="267" pagination="477 - 494" refId="ref15013" refString="Komai, T., Ravinesh, R., Riyas, A. &amp; Biju Kumar, A. (2020) New records of two sponge-associated species of Upogebia Leach, 1814 from southern India (Decapoda: Gebiidea: Upogebiidae): U. hexaceras (Ortmann, 1894) and U. nithyanandan (Sakai, Tu ¨ rkay &amp; Al Aidaroos, 2015) resurrected from the synonymy of U. balmaorum Ngoc-Ho, 1990. Zootaxa, 4747 (3), 477 - 494." type="journal article" year="2020">
Komai
<emphasis id="D9CBEAEEFF8BFFB7E6FC7CCDF7F6FF36" box="[1193,1249,225,251]" italics="true" pageId="16" pageNumber="267">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>