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<document id="1C39A9F7A9FE66181856D0BD67E8698F" ID-DOI="10.11646/zootaxa.4803.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3909077" ID-ZooBank="7BEC4D1E-A021-4A10-8E17-E7732E5193A4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593158411237" checkinUser="plazi" docAuthor="Al-Kandari, Manal, Anker, Arthur, Hussain, Sumaiah, Al-Yassen, Sharifa, Sattari, Zainab &amp; Grave, Sammy De" docDate="2020" docId="631687EAFF93FFAFE2C27E42F72CFA80" docLanguage="en" docName="zootaxa.4803.2.2.pdf" docOrigin="Zootaxa 4803 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Synalpheus quinquedens Tattersall 1921" docType="treatment" docVersion="5" lastPageNumber="259" masterDocId="9F2FFF92FF9BFFA7E2557C2FF317FFCD" masterDocTitle="New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda)" masterLastPageNumber="280" masterPageNumber="251" pageNumber="259" updateTime="1698776443936" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0731F464A6F5BA2110E4052DF2D258B7">New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda)</mods:title>
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<mods:namePart id="89E2775D227C8FCE363B606A3431588A">Al-Kandari, Manal</mods:namePart>
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<mods:namePart id="589199888723A8BECBE4EA21E809B817">Anker, Arthur</mods:namePart>
<mods:affiliation id="5F9913ABAA045A1859B7690D99058A15">Universidade Federal de Goiás, Campus Samambaia, Instituto de Ciências Biológicas-ICB- 5. Av. Esperança, s / n., 74690 - 900, Goiâ-</mods:affiliation>
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<mods:namePart id="7454F14369F61DE0444F83D3EE8E8D1F">Sattari, Zainab</mods:namePart>
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<taxonomicName id="2CBF4D7FFF93FFAFE2C27E42F199FD4A" ID-CoL="85YYL" authority="Tattersall, 1921" authorityName="Tattersall" authorityYear="1921" box="[151,654,621,648]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="quinquedens">
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<emphasis id="D9CBEAEEFF93FFAFE2C27E42F2D1FD45" bold="true" box="[151,454,621,648]" italics="true" pageId="8" pageNumber="259">Synalpheus quinquedens</emphasis>
<bibRefCitation id="8F2E4B0DFF93FFAFE3997E42F199FD4A" author="Tattersall, W. M." box="[460,654,621,647]" pageId="8" pageNumber="259" pagination="345 - 398" refId="ref17650" refString="Tattersall, W. M. (1921) Report on the Stomatopoda and macrurous Decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. The Journal of the Linnean Society, Zoology, 34, 345 - 398. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersall, 1921</bibRefCitation>
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<taxonomicName id="2CBF4D7FFF93FFAFE2C27E9EF171FD04" ID-CoL="85YYL" authority="Tattersall 1921: 376" authorityName="Tattersall" authorityPageNumber="376" authorityYear="1921" box="[151,614,688,713]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="D9CBEAEEFF93FFAFE2C27E9EF285FD04" box="[151,402,689,713]" italics="true" pageId="8" pageNumber="259">Synalpheus quinquedens</emphasis>
<bibRefCitation id="8F2E4B0DFF93FFAFE3CD7E9FF171FD04" author="Tattersall, W. M." box="[408,614,688,713]" pageId="8" pageNumber="259" pagination="345 - 398" refId="ref17650" refString="Tattersall, W. M. (1921) Report on the Stomatopoda and macrurous Decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. The Journal of the Linnean Society, Zoology, 34, 345 - 398. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersall 1921: 376</bibRefCitation>
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, pl. 28, figs. 15;
<treatmentCitation id="6A1E10EDFF93FFAFE1757E9FF0F4FD04" author="Ramadan, M. M." box="[800,995,688,713]" page="19" pageId="8" pageNumber="259" year="1936">
<bibRefCitation id="8F2E4B0DFF93FFAFE1757E9FF0F4FD04" author="Ramadan, M. M." box="[800,995,688,713]" pageId="8" pageNumber="259" pagination="1 - 43" refId="ref16580" refString="Ramadan, M. M. (1936) Report on a collection of Stomatopoda and Decapoda from Ghardaga, Red Sea. Bulletin of the Faculty of Science, The Egyptian University, 6, 1 - 43." type="journal article" year="1936">Ramadan 1936: 19</bibRefCitation>
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;
<treatmentCitation id="6A1E10EDFF93FFAFE1BA7E9DF611FD04" author="Banner, D. M. &amp; Banner, A. H." box="[1007,1286,689,713]" page="74" pageId="8" pageNumber="259" year="1981">
<bibRefCitation id="8F2E4B0DFF93FFAFE1BA7E9DF611FD04" author="Banner, D. M. &amp; Banner, A. H." box="[1007,1286,689,713]" pageId="8" pageNumber="259" pagination="1 - 99" refId="ref12195" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981: 74</bibRefCitation>
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, 92;
<treatmentCitation id="6A1E10EDFF93FFAFE76D7E9DF297FD25" author="Banner, A. H. &amp; Banner, D. M." page="105" pageId="8" pageNumber="259" year="1983">
<bibRefCitation id="8F2E4B0DFF93FFAFE76D7E9DF297FD25" author="Banner, A. H. &amp; Banner, D. M." pageId="8" pageNumber="259" pagination="1 - 164" refId="ref12098" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983: 105</bibRefCitation>
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;
<treatmentCitation id="6A1E10EDFF93FFAFE3DF7EFFF126FD25" author="Jones, D. A." box="[394,561,719,744]" page="153" pageId="8" pageNumber="259" year="1986">
<bibRefCitation id="8F2E4B0DFF93FFAFE3DF7EFFF126FD25" author="Jones, D. A." box="[394,561,719,744]" pageId="8" pageNumber="259" refId="ref14747" refString="Jones, D. A. (1986) A field guide to the sea shores of Kuwait and the Arabian Gulf. Blandford Press, University of Kuwait, Kuwait City, 192 pp." type="book" year="1986">Jones 1986: 153</bibRefCitation>
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, pl. 43;
<treatmentCitation id="6A1E10EDFF93FFAFE0D67EFFF0BBFD25" author="De Grave, S. &amp; Ashelby, C. W." box="[643,940,719,744]" page="3" pageId="8" pageNumber="259" year="2011">
<bibRefCitation id="8F2E4B0DFF93FFAFE0D67EFFF0BBFD25" author="De Grave, S. &amp; Ashelby, C. W." box="[643,940,719,744]" pageId="8" pageNumber="259" pagination="1 - 7" refId="ref13141" refString="De Grave, S. &amp; Ashelby, C. W. (2011) Notes on some shrimp species (Decapoda: Caridea) from Qatar, with an annotated checklist of marine caridean species known from the Persian Gulf. Marine Biodiversity Records, 4 (e 20), 1 - 7. https: // doi. org / 10.1017 / S 1755267211000224" type="journal article" year="2011">De Grave &amp; Ashelby 2011: 3</bibRefCitation>
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;
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<bibRefCitation id="8F2E4B0DFF93FFAFE1E27EE0F7C3FD25" author="Naderloo, R. &amp; Turkay, M." box="[951,1236,719,744]" pageId="8" pageNumber="259" pagination="1 - 67" refId="ref15707" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012: 5</bibRefCitation>
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;
<treatmentCitation id="6A1E10EDFF93FFAFE68A7EFFF68FFD25" author="George, J. D." box="[1247,1432,719,744]" page="285" pageId="8" pageNumber="259" year="2012">
<bibRefCitation id="8F2E4B0DFF93FFAFE68A7EFFF68FFD25" author="George, J. D." box="[1247,1432,719,744]" pageId="8" pageNumber="259" pagination="253 - 308" refId="ref13998" refString="George, J. D. (2012) Reef-associated macroinvertebrates of the SE Gulf. In: Riegl, B. M &amp; Purkis, S. J. (Eds.), Co ral reefs of the Gulf: Adaptation to climatic extremes. Coral Reefs of the World, 3, pp. 253 - 308. https: // doi. org / 10.1007 / 978 - 94 - 007 - 3008 - 3 _ 13" type="journal article" year="2012">George 2012: 285</bibRefCitation>
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.
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<emphasis id="D9CBEAEEFF93FFAFE2C27F3AF26CFCFD" bold="true" box="[151,379,789,816]" pageId="8" pageNumber="259">Material examined</emphasis>
.
<materialsCitation id="5BD73CA1FF93FFAFE3D97F39F12BFC99" collectingDate="2014-12-24" collectionCode="OUMNH, ZC" collectorName="M. Al-Kandari" country="Kuwait" location="Failaka Island" pageId="8" pageNumber="259" specimenCount="1" specimenCount-female="1">
<collectingCountry id="93A8766CFF93FFAFE3D97F39F2F4FCFD" box="[396,483,790,816]" name="Kuwait" pageId="8" pageNumber="259">Kuwait</collectingCountry>
:
<specimenCount id="FDB9FD75FF93FFAFE3BA7F39F142FCFD" box="[495,597,790,816]" pageId="8" pageNumber="259" type="female">1 female</specimenCount>
,
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.
<collectionCode id="8DAEAE39FF93FFAFE0807F39F1EDFCFD" box="[725,762,790,816]" country="Austria" name="Zoological Collection, University of Vienna" pageId="8" pageNumber="259" type="University or college">ZC</collectionCode>
. 2020.01.022,
<location id="EE606027FF93FFAFE1F67F39F751FCFD" LSID="urn:lsid:plazi:treatment:631687EAFF93FFAFE2C27E42F72CFA80:EE606027FF93FFAFE1F67F39F751FCFD" box="[931,1094,790,816]" country="Kuwait" name="Failaka Island" pageId="8" pageNumber="259">Failaka Island</location>
, eastern side, rocky substrate, leg.
<collectorName id="464A532AFF93FFAFE2927F14F27AFC99" box="[199,365,826,853]" pageId="8" pageNumber="259">M. Al-Kandari</collectorName>
<emphasis id="D9CBEAEEFF93FFAFE3267F14F2B3FC99" box="[371,420,826,852]" italics="true" pageId="8" pageNumber="259">et al</emphasis>
.,
<date id="9F01103CFF93FFAFE3EC7F15F12BFC99" box="[441,572,826,852]" pageId="8" pageNumber="259" value="2014-12-24">
<collectingDate id="8F45E9D4FF93FFAFE3EC7F15F12BFC99" box="[441,572,826,852]" pageId="8" pageNumber="259" value="2014-12-24">24.xii.2014</collectingDate>
</date>
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.
</paragraph>
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<emphasis id="D9CBEAEEFF93FFAFE2927F72F2AAFCB5" bold="true" box="[199,445,861,888]" pageId="8" pageNumber="259">General distribution</emphasis>
. Indo-West Pacific, with records from the Red Sea (e.g. Suez, Gulf of Aqaba, Ghardaqa, Khor Dongonab, Suakin), Arabian Gulf (
<collectingCountry id="93A8766CFF93FFAFE0087FADF19AFC51" box="[605,653,898,924]" name="Iran" pageId="8" pageNumber="259">Iran</collectingCountry>
,
<collectingCountry id="93A8766CFF93FFAFE0CC7FADF1C0FC51" box="[665,727,898,924]" name="Qatar" pageId="8" pageNumber="259">Qatar</collectingCountry>
,
<collectingCountry id="93A8766CFF93FFAFE0B67FADF02FFC50" box="[739,824,898,925]" name="Kuwait" pageId="8" pageNumber="259">Kuwait</collectingCountry>
), East Africa (
<collectingCountry id="93A8766CFF93FFAFE1827FADF733FC51" box="[983,1060,898,924]" name="Kenya" pageId="8" pageNumber="259">Kenya</collectingCountry>
),
<collectingCountry id="93A8766CFF93FFAFE6627FADF7ACFC51" box="[1079,1211,898,924]" name="Madagascar" pageId="8" pageNumber="259">Madagascar</collectingCountry>
and
<collectingCountry id="93A8766CFF93FFAFE6A67FADF64AFC51" box="[1267,1373,898,924]" name="Indonesia" pageId="8" pageNumber="259">Indonesia</collectingCountry>
(e.g.,
<bibRefCitation id="8F2E4B0DFF93FFAFE2C27F89F22AFC0D" author="Tattersall, W. M." box="[151,317,934,960]" pageId="8" pageNumber="259" pagination="345 - 398" refId="ref17650" refString="Tattersall, W. M. (1921) Report on the Stomatopoda and macrurous Decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. The Journal of the Linnean Society, Zoology, 34, 345 - 398. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersall 1921</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF93FFAFE31D7F89F2E6FC0D" author="Ramadan, M. M." box="[328,497,934,960]" pageId="8" pageNumber="259" pagination="1 - 43" refId="ref16580" refString="Ramadan, M. M. (1936) Report on a collection of Stomatopoda and Decapoda from Ghardaga, Red Sea. Bulletin of the Faculty of Science, The Egyptian University, 6, 1 - 43." type="journal article" year="1936">Ramadan 1936</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF93FFAFE3A97F88F1EAFC0D" author="Banner, D. M. &amp; Banner, A. H." box="[508,765,934,961]" pageId="8" pageNumber="259" pagination="1 - 99" refId="ref12195" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
,
<bibRefCitation id="8F2E4B0DFF93FFAFE15D7F89F055FC0D" author="Banner, A. H. &amp; Banner, D. M." box="[776,834,934,960]" pageId="8" pageNumber="259" pagination="1 - 164" refId="ref12098" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">1983</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF93FFAFE1187F88F0DCFC0D" author="Jones, D. A." box="[845,971,934,961]" pageId="8" pageNumber="259" refId="ref14747" refString="Jones, D. A. (1986) A field guide to the sea shores of Kuwait and the Arabian Gulf. Blandford Press, University of Kuwait, Kuwait City, 192 pp." type="book" year="1986">Jones 1986</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF93FFAFE1837F89F770FC0D" author="George, J. D." box="[982,1127,934,960]" pageId="8" pageNumber="259" pagination="253 - 308" refId="ref13998" refString="George, J. D. (2012) Reef-associated macroinvertebrates of the SE Gulf. In: Riegl, B. M &amp; Purkis, S. J. (Eds.), Co ral reefs of the Gulf: Adaptation to climatic extremes. Coral Reefs of the World, 3, pp. 253 - 308. https: // doi. org / 10.1007 / 978 - 94 - 007 - 3008 - 3 _ 13" type="journal article" year="2012">George 2012</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF93FFAFE6277F89F69BFC0D" author="Naderloo, R. &amp; Turkay, M." box="[1138,1420,934,960]" pageId="8" pageNumber="259" pagination="1 - 67" refId="ref15707" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A3A56577FF93FFAFE2927FE5F670FC28" box="[199,1383,970,997]" pageId="8" pageNumber="259" type="biology_ecology">
<paragraph id="EB0036FCFF93FFAFE2927FE5F670FC28" blockId="8.[151,1437,789,1357]" box="[199,1383,970,997]" pageId="8" pageNumber="259">
<emphasis id="D9CBEAEEFF93FFAFE2927FE5F231FC29" bold="true" box="[199,294,970,996]" pageId="8" pageNumber="259">Ecology</emphasis>
. Lower intertidal to
<quantity id="2C479B19FF93FFAFE0587FE5F151FC28" box="[525,582,970,997]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="8" pageNumber="259" unit="m" value="64.0">64 m</quantity>
(
<bibRefCitation id="8F2E4B0DFF93FFAFE0017FE5F048FC29" author="Banner, A. H. &amp; Banner, D. M." box="[596,863,970,997]" pageId="8" pageNumber="259" pagination="1 - 164" refId="ref12098" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983</bibRefCitation>
); possibly associated with encrusting sponges.
</paragraph>
</subSubSection>
<subSubSection id="A3A56577FF93FFAFE2927FC1F72CFA80" pageId="8" pageNumber="259" type="discussion">
<paragraph id="EB0036FCFF93FFAFE2927FC1F28FFBB9" blockId="8.[151,1437,789,1357]" pageId="8" pageNumber="259">
<emphasis id="D9CBEAEEFF93FFAFE2927FC1F223FBC5" bold="true" box="[199,308,1006,1032]" pageId="8" pageNumber="259">Remarks</emphasis>
. The species was previously known from the Arabian
<collectingRegion id="297BF81EFF93FFAFE1FC7FC1F0CBFBC5" box="[937,988,1006,1032]" country="Papua New Guinea" name="Gulf" pageId="8" pageNumber="259">Gulf</collectingRegion>
based on unspecified material from an unknown locality collected by the “Danish Persian
<collectingRegion id="297BF81EFF93FFAFE098783DF017FBE1" box="[717,768,1042,1068]" country="Papua New Guinea" name="Gulf" pageId="8" pageNumber="259">Gulf</collectingRegion>
Expedition” (
<bibRefCitation id="8F2E4B0DFF93FFAFE1CE783DF7B5FBE1" author="Banner, D. M. &amp; Banner, A. H." box="[923,1186,1042,1069]" pageId="8" pageNumber="259" pagination="1 - 99" refId="ref12195" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
). Since this expedition was conducted mainly along the Iranian coast,
<bibRefCitation id="8F2E4B0DFF93FFAFE0F67819F0C7FB9D" author="Naderloo, R. &amp; Turkay, M." box="[675,976,1078,1104]" pageId="8" pageNumber="259" pagination="1 - 67" refId="ref15707" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay (2012)</bibRefCitation>
included
<taxonomicName id="2CBF4D7FFF93FFAFE66B7818F7F0FB9D" authorityName="Tattersall" authorityYear="1921" box="[1086,1255,1078,1105]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="D9CBEAEEFF93FFAFE66B7818F7F0FB9D" box="[1086,1255,1078,1105]" italics="true" pageId="8" pageNumber="259">S. quinquedens</emphasis>
</taxonomicName>
in the list of the Iranian decapod fauna.
</paragraph>
<paragraph id="EB0036FCFF93FFAFE2927851F72CFA80" blockId="8.[151,1437,789,1357]" pageId="8" pageNumber="259">
<taxonomicName id="2CBF4D7FFF93FFAFE2927851F2C9FB55" authorityName="Tattersall" authorityYear="1921" box="[199,478,1150,1176]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="D9CBEAEEFF93FFAFE2927851F2C9FB55" box="[199,478,1150,1176]" italics="true" pageId="8" pageNumber="259">Synalpheus quinquedens</emphasis>
</taxonomicName>
was also included in the popular field-guide to the intertidal macrofauna of
<collectingCountry id="93A8766CFF93FFAFE71F7851F68BFB55" box="[1354,1436,1150,1176]" name="Kuwait" pageId="8" pageNumber="259">Kuwait</collectingCountry>
(
<bibRefCitation id="8F2E4B0DFF93FFAFE2CA788DF236FB70" author="Jones, D. A." box="[159,289,1186,1213]" pageId="8" pageNumber="259" refId="ref14747" refString="Jones, D. A. (1986) A field guide to the sea shores of Kuwait and the Arabian Gulf. Blandford Press, University of Kuwait, Kuwait City, 192 pp." type="book" year="1986">Jones 1986</bibRefCitation>
), with a short text accompanied by a highly diagrammatic, inaccurate illustration. Importantly, the diagnostic characters (presence of five strong rounded or subacute teeth on the posterior margin of the sixth pleonite) of
<taxonomicName id="2CBF4D7FFF93FFAFE2C278C4F256FAC9" authorityName="Tattersall" authorityYear="1921" box="[151,321,1258,1285]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="D9CBEAEEFF93FFAFE2C278C4F256FAC9" box="[151,321,1258,1285]" italics="true" pageId="8" pageNumber="259">S. quinquedens</emphasis>
</taxonomicName>
cannot be seen in the figure of the general habitus provided by
<bibRefCitation id="8F2E4B0DFF93FFAFE65C78C4F78CFAC8" author="Jones, D. A." box="[1033,1179,1258,1285]" pageId="8" pageNumber="259" refId="ref14747" refString="Jones, D. A. (1986) A field guide to the sea shores of Kuwait and the Arabian Gulf. Blandford Press, University of Kuwait, Kuwait City, 192 pp." type="book" year="1986">Jones (1986)</bibRefCitation>
. Therefore, the present specimen represents the first formal record of
<taxonomicName id="2CBF4D7FFF93FFAFE0F77920F05BFAE5" authorityName="Tattersall" authorityYear="1921" box="[674,844,1294,1321]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="quinquedens">
<emphasis id="D9CBEAEEFF93FFAFE0F77920F05BFAE5" box="[674,844,1294,1321]" italics="true" pageId="8" pageNumber="259">S. quinquedens</emphasis>
</taxonomicName>
from
<collectingCountry id="93A8766CFF93FFAFE1C77921F0F0FAE5" box="[914,999,1294,1320]" name="Kuwait" pageId="8" pageNumber="259">Kuwait</collectingCountry>
, confirming the previous, rather vague records of the species from the Arabian Gulf (
<bibRefCitation id="8F2E4B0DFF93FFAFE0CD791DF0B6FA81" author="Banner, D. M. &amp; Banner, A. H." box="[664,929,1330,1357]" pageId="8" pageNumber="259" pagination="1 - 99" refId="ref12195" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF93FFAFE1F8791CF739FA80" author="Jones, D. A." box="[941,1070,1330,1357]" pageId="8" pageNumber="259" refId="ref14747" refString="Jones, D. A. (1986) A field guide to the sea shores of Kuwait and the Arabian Gulf. Blandford Press, University of Kuwait, Kuwait City, 192 pp." type="book" year="1986">Jones 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>