163 lines
14 KiB
XML
163 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.33.5078" ID-GBIF-Taxon="182249313" ID-Pensoft-Pub="1314-2607-33-1" ID-Pensoft-UUID="5F20FFE2E95AAC21FFD1FFA0D8160112" ID-Zenodo-Dep="574825" ID-ZooBank="88E03F8FD9B947F1B6E90456D47B1E37" ModsDocID="1314-2607-33-1" checkinTime="1621553083643" checkinUser="pensoft" docAuthor="Kimsey, Lynn S. & Wasbauer, Marius S." docDate="2013" docId="6300B4D650F2B36C9B1EBDE3D2890E4B" docLanguage="en" docOrigin="Journal of Hymenoptera Research 33" docPubDate="2013-08-01" docSource="http://dx.doi.org/10.3897/jhr.33.5078" docTitle="Colocistis pilosa Krombein 1942" docType="treatment" docVersion="2" id="5F20FFE2E95AAC21FFD1FFA0D8160112" lastPageId="15" lastPageNumber="16" masterDocId="5F20FFE2E95AAC21FFD1FFA0D8160112" masterDocTitle="Revision of the brachycistidine genus Colocistis Krombein, 1942 (Hymenoptera, Tiphiidae)" masterLastPageNumber="24" masterPageNumber="1" pageId="14" pageNumber="15" updateTime="1643536911076" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the brachycistidine genus Colocistis Krombein, 1942 (Hymenoptera, Tiphiidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kimsey, Lynn S.</mods:namePart>
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<mods:affiliation>Bohart Museum of Entomology, Department of Entomology, University of California, Davis, 95616 USA</mods:affiliation>
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<mods:nameIdentifier type="email">lskimsey@ucdavis.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wasbauer, Marius S.</mods:namePart>
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<mods:affiliation>Bohart Museum of Entomology, Department of Entomology, University of California, Davis, 95616 USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-08-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>33</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>24</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.33.5078</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.33.5078</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-33-1</mods:identifier>
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<mods:identifier type="ZooBank">88E03F8FD9B947F1B6E90456D47B1E37</mods:identifier>
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<mods:identifier type="Pensoft-UUID">5F20FFE2E95AAC21FFD1FFA0D8160112</mods:identifier>
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<mods:identifier type="Zenodo-Dep">574825</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182249313" LSID="urn:lsid:plazi:treatment:6300B4D650F2B36C9B1EBDE3D2890E4B" httpUri="http://treatment.plazi.org/id/6300B4D650F2B36C9B1EBDE3D2890E4B" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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<subSubSection pageId="14" pageNumber="15" type="nomenclature">
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName LSID="http://species-id.net/wiki/Colocistis_pilosa" authority="Krombein" authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Colocistis pilosa" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pilosa">Colocistis pilosa Krombein</taxonomicName>
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<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="14" pageNumber="15">Figures 7</figureCitation>
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<figureCitation captionStart="Figures 25–32" captionStartId="F3" captionText="Figures 25 - 32. 25 - 28. Males. 29 - 32 Females. 25, 27, 29, 31 Lateral view 26, 28, 30, 32 Dorsal view 25 - 30 are images of the holotypes." httpUri="https://binary.pensoft.net/fig/11789" pageId="14" pageNumber="15">, 27, 28</figureCitation>
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<figureCitation captionStart="Figures 37–40" captionStartId="F5" captionText="Figures 37 - 40. Distribution maps Colocistis species." httpUri="https://binary.pensoft.net/fig/11791" pageId="14" pageNumber="15">, 37</figureCitation>
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<figureCitation captionStart="Figures 41–48" captionStartId="F6" captionText="Figures 41 - 48. Ventral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11792" pageId="14" pageNumber="15">, 47</figureCitation>
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<figureCitation captionStart="Figures 49–56" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="14" pageNumber="15">, 55</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="15" type="reference_group">
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pilosa">Colocistis pilosa</taxonomicName>
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Krombein, 1942:66. Holotype male; USA: California, San Diego County, San Diego, June 22, 1890 (USNM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="15" type="diagnosis">
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<paragraph pageId="14" pageNumber="15">Diagnosis.</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pilosa">
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<emphasis italics="true" pageId="14" pageNumber="15">Colocistis pilosa</emphasis>
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</taxonomicName>
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is only one of two species, including some specimens of
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<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="castanea">
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<emphasis italics="true" pageId="14" pageNumber="15">Colocistis castanea</emphasis>
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</taxonomicName>
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, with black body coloration. However, it shares with
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<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="crassa">
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<emphasis italics="true" pageId="14" pageNumber="15">Colocistis crassa</emphasis>
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</taxonomicName>
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the distinctive modification of metasomal sternum II, with the basal area elevated and separated from the rest of the sternum by a transverse declivity. These two species can be separated by the clypeus without a medial tubercle in
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<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pilosa">
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<emphasis italics="true" pageId="14" pageNumber="15">Colocistis pilosa</emphasis>
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</taxonomicName>
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(well-developed in
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<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="castanea">
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<emphasis italics="true" pageId="14" pageNumber="15">Colocistis castanea</emphasis>
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</taxonomicName>
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) and small compound eyes and ocelli (large in
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<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="castanea">
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<emphasis italics="true" pageId="14" pageNumber="15">Colocistis castanea</emphasis>
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</taxonomicName>
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), and long, dense, whitish pubescence.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
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<paragraph pageId="14" pageNumber="15">Description.</paragraph>
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<paragraph pageId="14" pageNumber="15">
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Male (
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<figureCitation captionStart="Figures 25–32" captionStartId="F3" captionText="Figures 25 - 32. 25 - 28. Males. 29 - 32 Females. 25, 27, 29, 31 Lateral view 26, 28, 30, 32 Dorsal view 25 - 30 are images of the holotypes." httpUri="https://binary.pensoft.net/fig/11789" pageId="14" pageNumber="15">Figs 27, 28</figureCitation>
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).
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<emphasis italics="true" pageId="14" pageNumber="15">Body length</emphasis>
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11-12 mm.
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<emphasis italics="true" pageId="14" pageNumber="15">Head</emphasis>
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(
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<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="14" pageNumber="15">Fig. 7</figureCitation>
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). Compound eyes not enlarged, barely extending beyond temples, inner margins slightly evenly curved; ocelli small, diameter of lateral ocellus 0.5
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<normalizedToken originalValue="×">x</normalizedToken>
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OOL; without impressed line between lateral ocelli; antennal socket rim slightly thickened ventrolaterally; clypeus with median tubercle visible in lateral view, apex bounded laterally by strong teeth; gular carina not raised; LID 0.85
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<normalizedToken originalValue="×">x</normalizedToken>
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UID; flagellomere I length 2.4
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<normalizedToken originalValue="×">x</normalizedToken>
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breadth; flagellomeres II-X missing; occipital carina strong but incomplete ventrally, with carina extending from foramen magnum to hypostomal carina medially; mandible narrowest submedially.
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</paragraph>
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<paragraph pageId="15" pageNumber="16">
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<emphasis italics="true" pageId="15" pageNumber="16">
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<pageBreakToken pageId="15" pageNumber="16" start="start">Mesosoma</pageBreakToken>
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</emphasis>
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. Pronotum nearly vertical anteriorly, sloping posteriorly, punctures deep, largely contiguous, shoulders well-developed, not produced laterally; forefemur without stout spines; foretibia without spines, except apically; basitarsus with three spines on outer surface; tibial spur 0.9
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<normalizedToken originalValue="×">x</normalizedToken>
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length of basitarsus; hindcoxa without longitudinal dorsal carina tooth-like basally; mesonotal and mesopleural punctures contiguous; scutellum strongly arched dorsally much above level of propodeum, punctures 0.5-1.5 puncture diameters apart, with small, polished impunctate area medially; postscutellum punctures large, contiguous; propodeum dorsal surface coarsely rugulose, separated from posterior face by distinct carina, posterior face punctures contiguous laterally, becoming more widely separated toward midline, except for small dorsomedial impunctate area, punctures becoming small toward petiole, laterally punctures 0.5 puncture diameter apart to contiguous; forewing with 3 submarginal cells, costa extending along wing margin beyond stigma.
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</paragraph>
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<paragraph pageId="15" pageNumber="16">
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<emphasis italics="true" pageId="15" pageNumber="16">Metasoma</emphasis>
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. Tergum I without transverse carina; sternum II with anterior portion slightly raised and separated from remainder of sternum by transverse declivity.
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</paragraph>
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<paragraph pageId="15" pageNumber="16">
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<emphasis italics="true" pageId="15" pageNumber="16">Genitalia</emphasis>
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(
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<figureCitation captionStart="Figures 41–48" captionStartId="F6" captionText="Figures 41 - 48. Ventral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11792" pageId="15" pageNumber="16">Figs 47</figureCitation>
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,
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<figureCitation captionStart="Figures 49–56" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="15" pageNumber="16">55</figureCitation>
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). Gonostylus slightly emarginate ventrally before tip in lateral view; aedeagus evenly curved, simple at apex in lateral view.
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</paragraph>
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<paragraph pageId="15" pageNumber="16">
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<emphasis italics="true" pageId="15" pageNumber="16">Color</emphasis>
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. Head and body unicolorous, black, with extensive long, erect pale, yellowish setae, setae more than half as long as eye height.
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</paragraph>
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<paragraph pageId="15" pageNumber="16">Female unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="16" type="distribution">
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<paragraph pageId="15" pageNumber="16">Distribution</paragraph>
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<paragraph pageId="15" pageNumber="16">
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(
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<figureCitation captionStart="Figures 37–40" captionStartId="F5" captionText="Figures 37 - 40. Distribution maps Colocistis species." httpUri="https://binary.pensoft.net/fig/11791" pageId="15" pageNumber="16">Fig. 37</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="16" type="discussion">
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<paragraph pageId="15" pageNumber="16">Discussion.</paragraph>
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<paragraph pageId="15" pageNumber="16">
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<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pilosa">
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<emphasis italics="true" pageId="15" pageNumber="16">Colocistis pilosa</emphasis>
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</taxonomicName>
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may be extinct; no specimens have been seen since the type was collected. The San Diego region where the type was collected has become increasingly urbanized since 1942 and is now highly urbanized with commercial and residential developments.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |