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<document id="92188F467F19B052DFADE424D4AA02FC" ID-DOI="10.11646/zootaxa.5551.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14389841" ID-ZooBank="BEBB4D36-855E-4AD3-829D-A3B2B17B81B7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1733924247133" checkinUser="plazi" docAuthor="Brown, John W." docDate="2024" docId="626F651B187BFF95FF3812E6FB0AFF33" docLanguage="en" docName="zootaxa.5551.1.2.pdf" docOrigin="Zootaxa 5551 (1)" docSource="https://doi.org/10.11646/zootaxa.5551.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Argentiliana Brown 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="53" masterDocId="9E561D63187AFF97FFAF1134FFAFFFE6" masterDocTitle="New genera and species of tortricid moths from Chile and Argentina (Lepidoptera: Tortricidae)" masterLastPageNumber="90" masterPageNumber="51" pageNumber="52" updateTime="1733925089986" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B0EA156C4A10F32E5ABF3E458EC28330">New genera and species of tortricid moths from Chile and Argentina (Lepidoptera: Tortricidae)</mods:title>
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<mods:namePart id="08FA70E676BD270AF29B69DBF78C29E6">Brown, John W.</mods:namePart>
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<mods:date id="A5BF3A75712A382EE33DA5E0375DED28">2024</mods:date>
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<treatment id="626F651B187BFF95FF3812E6FB0AFF33" LSID="urn:lsid:plazi:treatment:626F651B187BFF95FF3812E6FB0AFF33" httpUri="http://treatment.plazi.org/id/626F651B187BFF95FF3812E6FB0AFF33" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
<subSubSection id="A2DC8786187BFF96FF3812E6FC44FBF7" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="EA79D40D187BFF96FF3812E6FDB3FC0B" blockId="1.[151,1003,978,1041]" box="[151,540,978,1005]" pageId="1" pageNumber="52">
<heading id="B1316361187BFF96FF3812E6FDB3FC0B" bold="true" box="[151,540,978,1005]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
<emphasis id="D8B2081F187BFF96FF3812E6FDB3FC0B" bold="true" box="[151,540,978,1005]" pageId="1" pageNumber="52">
<taxonomicName id="2DC6AF8E187BFF96FF3812E6FE21FC0B" authority="Brown" authorityName="Brown" authorityYear="2024" box="[151,398,978,1005]" class="Insecta" family="Tortricidae" genus="Argentiliana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D8B2081F187BFF96FF3812E6FE9FFC0B" bold="true" box="[151,304,978,1005]" italics="true" pageId="1" pageNumber="52">Argentiliana</emphasis>
Brown
</taxonomicName>
,
<taxonomicNameLabel id="C381B564187BFF96FE3412E7FDB3FC0B" box="[411,540,979,1005]" pageId="1" pageNumber="52" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
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LSIDurn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="A2DC8786187BFF96FF38150BFD47FBBF" box="[151,744,1086,1113]" pageId="1" pageNumber="52" type="type_taxon">
<paragraph id="EA79D40D187BFF96FF38150BFD47FBBF" blockId="1.[151,1437,1086,2013]" box="[151,744,1086,1113]" pageId="1" pageNumber="52">
<emphasis id="D8B2081F187BFF96FF38150BFE84FBBE" bold="true" box="[151,299,1086,1113]" pageId="1" pageNumber="52">
<typeStatus id="357D6AAF187BFF96FF38150BFF7DFBBF" box="[151,210,1087,1113]" pageId="1" pageNumber="52">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="2DC6AF8E187BFF96FE95150BFDE3FBBF" authorityName="Brown" authorityYear="2024" box="[314,588,1087,1113]" class="Insecta" family="Tortricidae" genus="Argentiliana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="basipallida" status="sp. nov.">
<emphasis id="D8B2081F187BFF96FE95150BFDE3FBBF" box="[314,588,1087,1113]" italics="true" pageId="1" pageNumber="52">Argentiliana basipallida</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C381B564187BFF96FDF5150BFD4CFBBF" box="[602,739,1087,1113]" pageId="1" pageNumber="52" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="A2DC8786187BFF96FF681556FF44FA7B" pageId="1" pageNumber="52" type="diagnosis">
<paragraph id="EA79D40D187BFF96FF681556FF44FA7B" blockId="1.[151,1437,1086,2013]" pageId="1" pageNumber="52">
<emphasis id="D8B2081F187BFF96FF681556FE95FB9A" bold="true" box="[199,314,1122,1148]" pageId="1" pageNumber="52">Diagnosis</emphasis>
.
<taxonomicName id="2DC6AF8E187BFF96FEE41557FE77FB9B" authorityName="Brown" authorityYear="2024" box="[331,472,1123,1149]" class="Insecta" family="Tortricidae" genus="Argentiliana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="D8B2081F187BFF96FEE41557FE77FB9B" box="[331,472,1123,1149]" italics="true" pageId="1" pageNumber="52">Argentiliana</emphasis>
</taxonomicName>
can be distinguished from all other cochyline genera by facies alone. The forewing pattern includes a pale region in the basal 0.33, with a contrasting, dark brown, linear dash along the costa (
<figureCitation id="72FDC888187BFF96FAE615B3FA25FB47" box="[1353,1418,1159,1185]" captionStart="FIGURES 18" captionStartId="3.[151,264,1592,1616]" captionTargetBox="[151,1436,171,1572]" captionTargetId="figure-191@3.[151,1436,171,1572]" captionTargetPageId="3" captionText="FIGURES 18.Adults of tortricid species from Chile andArgentina.1. Argentiliana basipallidana, paratype female. 2. Terifrona fronteria, holotype male. 3. Transtilliana biloba, paratype male. 4. Transtilliana biloba, paratype male. 5. Vargasia karsholti, holotype male. 6. Hamiamia trancasiana, Chile, Ñuble Province. 7. Varifula fulvaria, Chile, Aconcagua Province. 8. Varifula geomops, holotype male." figureDoi="http://doi.org/10.5281/zenodo.14389843" httpUri="https://zenodo.org/record/14389843/files/figure.png" pageId="1" pageNumber="52">Fig. 1</figureCitation>
); the remainder of the wing features shades of brown and gray with a pair of narrow, dark brown lines bordering an irregular median fascia (
<figureCitation id="72FDC888187BFF96FE0615FBFE45FB0F" box="[425,490,1231,1257]" captionStart="FIGURES 18" captionStartId="3.[151,264,1592,1616]" captionTargetBox="[151,1436,171,1572]" captionTargetId="figure-191@3.[151,1436,171,1572]" captionTargetPageId="3" captionText="FIGURES 18.Adults of tortricid species from Chile andArgentina.1. Argentiliana basipallidana, paratype female. 2. Terifrona fronteria, holotype male. 3. Transtilliana biloba, paratype male. 4. Transtilliana biloba, paratype male. 5. Vargasia karsholti, holotype male. 6. Hamiamia trancasiana, Chile, Ñuble Province. 7. Varifula fulvaria, Chile, Aconcagua Province. 8. Varifula geomops, holotype male." figureDoi="http://doi.org/10.5281/zenodo.14389843" httpUri="https://zenodo.org/record/14389843/files/figure.png" pageId="1" pageNumber="52">Fig. 1</figureCitation>
). The overall aspect of the single included species is somewhat reminiscent of some
<taxonomicName id="2DC6AF8E187BFF96FF3815C7FE81FAEB" authorityName="Walsingham" authorityYear="1897" box="[151,302,1267,1293]" class="Insecta" family="Carposinidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="family">Carposinidae</taxonomicName>
, but the presence of ocelli, chaetosemata, and characteristic tortricoid ovipositor lobes (i.e., large, flattened papillae anales) confirm its familial assignment to
<taxonomicName id="2DC6AF8E187BFF96FCE21423FC64FAD7" box="[845,971,1303,1329]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="family">Tortricidae</taxonomicName>
. The female genitalia are simple, with a broad antrum and a long ductus bursae that broadens slightly to a pear-shaped corpus bursae lacking a signum. In the absence of males, the plesiomorphic female genitalia provide few clues to which genera
<taxonomicName id="2DC6AF8E187BFF96FB0A146BFA9DFA9F" authorityName="Brown" authorityYear="2024" box="[1189,1330,1375,1401]" class="Insecta" family="Tortricidae" genus="Argentiliana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="D8B2081F187BFF96FB0A146BFA9DFA9F" box="[1189,1330,1375,1401]" italics="true" pageId="1" pageNumber="52">Argentiliana</emphasis>
</taxonomicName>
might be related.
</paragraph>
</subSubSection>
<subSubSection id="A2DC8786187BFF96FF681492FBCCF83B" pageId="1" pageNumber="52" type="description">
<paragraph id="EA79D40D187BFF96FF681492FCA0F993" blockId="1.[151,1437,1086,2013]" pageId="1" pageNumber="52">
<emphasis id="D8B2081F187BFF96FF681492FEFEFA26" bold="true" box="[199,337,1446,1472]" pageId="1" pageNumber="52">Description</emphasis>
. Head: Vertex rough-scaled, scales directed anteriorly, posterior region of vertex with a pair of large, subdorsal, fan-like scale patches; scaling on frons shorter, more appressed; antenna ca. 0.5 length of forewing, non-serrate, with one row of scales per flagellomere, sensory setae extremely short, less than 0.25 width of flagellomere; ocellus well developed; chaetosemata present; labial palpus relatively slender, scaling only slightly expanded distally, third segment exposed, combined length of all segments ca. 1.25 times diameter of compound eye. Haustellum well developed, presumably functional.
</paragraph>
<paragraph id="EA79D40D187BFF96FF68174BFC17F8E3" blockId="1.[151,1437,1086,2013]" pageId="1" pageNumber="52">
Thorax: Dorsum smooth-scaled, metathoracic tuft present. Legs unmodified, without hairpencil or other secondary scaling. Forewing length 2.4 times width; costa weakly and evenly arched throughout; termen straight, oblique; all veins present and separate, chorda and M-stem absent, R
<subScript id="7642D648187BFF96FC2317EDFC3BF901" attach="left" box="[908,916,1753,1767]" fontSize="6" pageId="1" pageNumber="52">4</subScript>
to costa, R
<subScript id="7642D648187BFF96FBA117EDFBB9F901" attach="left" box="[1038,1046,1753,1767]" fontSize="6" pageId="1" pageNumber="52">5</subScript>
to termen. Hindwing with all veins present and separate; CuP weak at margin; frenulum with three bristles.
</paragraph>
<paragraph id="EA79D40D187BFF96FF68163BFBCCF83B" blockId="1.[151,1437,1086,2013]" pageId="1" pageNumber="52">
Abdomen: Male genitalia unknown. Female genitalia (
<figureCitation id="72FDC888187BFF96FC84163BFCD3F8CF" box="[811,892,1807,1833]" captionStart="FIGURES 3944" captionStartId="27.[151,264,1948,1972]" captionTargetBox="[151,1436,520,1925]" captionTargetId="figure-170@27.[151,1436,520,1928]" captionTargetPageId="27" captionText="FIGURES 3944. Female genitalia of tortricid species from Chile and Argentina. 39. Argentiliana basipallida, USNM slide 154,800. 40. Terifrona fronteria, USNM slide 145,518. 41. Transtilliana biloba, USNM slide 154,847. 42. Vargasia karsholti, JWBrown slide 1216. 43. Hamiamia trancasiana, USNM slide 92491. 44. Varifula fulvaria, USNM slide 88,471." figureDoi="http://doi.org/10.5281/zenodo.14389861" httpUri="https://zenodo.org/record/14389861/files/figure.png" pageId="1" pageNumber="52">Fig. 39</figureCitation>
) with papilla analis rather slender, ca. 4 times as long as wide; apophyses slender, posteriores ca. 0.75 as long as anteriores; sterigma membranous, with transverse band bearing a pair of short, narrow, lateral, anteriorly-directed, triangular “tails” from anterior margin; ostium wide; antrum wide, gradually narrowing to ductus bursae; ductus bursae nearly uniform in width anterad of antrum, broadening slightly to junction with corpus bursae; ductus seminalis arising from ductus bursae ca. 0.33 distance from ostium; corpus bursae pear-shaped, lacking signum and conspicuous scobination.
</paragraph>
</subSubSection>
<subSubSection id="A2DC87861878FF95FF6811A3FB0AFF33" pageId="2" pageNumber="53" type="etymology">
<paragraph id="EA79D40D1878FF95FF6811A3FB0AFF33" blockId="2.[151,1436,151,213]" pageId="2" pageNumber="53">
<emphasis id="D8B2081F1878FF95FF6811A3FEE7FF57" bold="true" box="[199,328,151,177]" pageId="2" pageNumber="53">Etymology</emphasis>
. The generic name is a combination of
<taxonomicName id="2DC6AF8E1878FF95FCB511A3FC36FF57" authorityName="Brown" authorityYear="1991" box="[794,921,151,177]" class="Insecta" family="Tortricidae" genus="Cuproxena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="argentina">“Argentina</taxonomicName>
” and “Gentili,” the latter the surname of the noted
<collectingCountry id="92D1949D1878FF95FF74118FFEE5FF33" box="[219,330,187,213]" name="Argentina" pageId="2" pageNumber="53">Argentine</collectingCountry>
lepidopterist Mario Gentili, the collector of the
<typeStatus id="357D6AAF1878FF95FCCA118FFC68FF33" box="[869,967,187,213]" pageId="2" pageNumber="53" type="holotype">holotype</typeStatus>
of the type species.
</paragraph>
</subSubSection>
</treatment>
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