187 lines
20 KiB
XML
187 lines
20 KiB
XML
<document id="D8AFC1E2A2307F3CDF82E11C4E9F1B93" ID-DOI="10.11646/zootaxa.5551.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14389841" ID-ZooBank="BEBB4D36-855E-4AD3-829D-A3B2B17B81B7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1733924247133" checkinUser="plazi" docAuthor="Brown, John W." docDate="2024" docId="626F651B1871FF9CFF3811A3FB70FB20" docLanguage="en" docName="zootaxa.5551.1.2.pdf" docOrigin="Zootaxa 5551 (1)" docSource="https://doi.org/10.11646/zootaxa.5551.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Varifula geomops Brown 2024, new species" docType="treatment" docVersion="2" lastPageNumber="62" masterDocId="9E561D63187AFF97FFAF1134FFAFFFE6" masterDocTitle="New genera and species of tortricid moths from Chile and Argentina (Lepidoptera: Tortricidae)" masterLastPageNumber="90" masterPageNumber="51" pageNumber="62" updateTime="1733925089986" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AC70BD228EBA1C59766DA9CFF69B71FA">New genera and species of tortricid moths from Chile and Argentina (Lepidoptera: Tortricidae)</mods:title>
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<mods:name id="BA4DFAC8333F063B3662233C39299117" type="personal">
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<mods:namePart id="4D511DFA2F73368024F52A23A78FBDF8">Brown, John W.</mods:namePart>
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<mods:title id="49B01190F86CB28110DD7102AB208FA5">Zootaxa</mods:title>
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<mods:part id="C383F27D44215F0E26E8A548106E3552">
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<mods:date id="2425A5CB7475B5543EA8219835A487CC">2024</mods:date>
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<mods:number id="30CA8A1E8C7B80B8E9C735E4C78C4406">2024-12-11</mods:number>
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<mods:number id="7F651A5582FDE68BA0DA36B80DA74950">5551</mods:number>
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<mods:url id="17C5347D7FF299EE643200ADDEF0A9D8">https://doi.org/10.11646/zootaxa.5551.1.2</mods:url>
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<mods:classification id="45465EDC1E411911BDDA66D325EE27B6">journal article</mods:classification>
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<mods:identifier id="BEA804F095B916EC38F5A4523F027D3D" type="DOI">10.11646/zootaxa.5551.1.2</mods:identifier>
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<mods:identifier id="8832796DBDB28C792FB4CCC2CB52D797" type="ISSN">1175-5326</mods:identifier>
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<treatment id="626F651B1871FF9CFF3811A3FB70FB20" LSID="urn:lsid:plazi:treatment:626F651B1871FF9CFF3811A3FB70FB20" httpUri="http://treatment.plazi.org/id/626F651B1871FF9CFF3811A3FB70FB20" lastPageNumber="62" pageId="11" pageNumber="62">
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<subSubSection id="A2DC87861871FF9CFF3811A3FBACFF30" pageId="11" pageNumber="62" type="nomenclature">
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<paragraph id="EA79D40D1871FF9CFF3811A3FDC9FF57" blockId="11.[151,1027,151,249]" box="[151,614,151,178]" pageId="11" pageNumber="62">
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<heading id="B13163611871FF9CFF3811A3FDC9FF57" bold="true" box="[151,614,151,178]" fontSize="11" level="1" pageId="11" pageNumber="62" reason="1">
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<emphasis id="D8B2081F1871FF9CFF3811A3FDC9FF57" bold="true" box="[151,614,151,178]" pageId="11" pageNumber="62">
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<taxonomicName id="2DC6AF8E1871FF9CFF3811A3FE66FF57" authority="Brown" authorityName="Brown" authorityYear="2024" box="[151,457,151,178]" class="Insecta" family="Tortricidae" genus="Varifula" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="geomops" status="sp. nov.">
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<emphasis id="D8B2081F1871FF9CFF3811A3FEC3FF57" bold="true" box="[151,364,151,178]" italics="true" pageId="11" pageNumber="62">Varifula geomops</emphasis>
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Brown
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</taxonomicName>
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,
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<taxonomicNameLabel id="C381B5641871FF9CFE7911A3FDC9FF57" box="[470,614,151,177]" pageId="11" pageNumber="62" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EA79D40D1871FF9CFF38118FFBACFF30" blockId="11.[151,1027,151,249]" box="[151,1027,187,214]" pageId="11" pageNumber="62">
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LSIDurn:lsid:zoobank.org:act:
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<uuid id="9E60EED81871FF9CFE41118FFBACFF30" box="[494,1027,187,214]" pageId="11" pageNumber="62">E6BD94A1-FBB5-4335-8361-4DF7E50761A5</uuid>
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</paragraph>
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</subSubSection>
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<subSubSection id="A2DC87861871FF9CFF3811EBFE8FFF1F" box="[151,288,223,249]" pageId="11" pageNumber="62" type="description">
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<paragraph id="EA79D40D1871FF9CFF3811EBFE8FFF1F" blockId="11.[151,1027,151,249]" box="[151,288,223,249]" pageId="11" pageNumber="62">
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(
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<figureCitation id="72FDC8881871FF9CFF3011EBFF41FF1F" box="[159,238,223,249]" captionStart="FIGURES 1‒8" captionStartId="3.[151,264,1592,1616]" captionTargetBox="[151,1436,171,1572]" captionTargetId="figure-191@3.[151,1436,171,1572]" captionTargetPageId="3" captionText="FIGURES 1‒8.Adults of tortricid species from Chile andArgentina.1. Argentiliana basipallidana, paratype female. 2. Terifrona fronteria, holotype male. 3. Transtilliana biloba, paratype male. 4. Transtilliana biloba, paratype male. 5. Vargasia karsholti, holotype male. 6. Hamiamia trancasiana, Chile, Ñuble Province. 7. Varifula fulvaria, Chile, Aconcagua Province. 8. Varifula geomops, holotype male." figureDoi="http://doi.org/10.5281/zenodo.14389843" httpUri="https://zenodo.org/record/14389843/files/figure.png" pageId="11" pageNumber="62">Figs. 8</figureCitation>
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,
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<figureCitation id="72FDC8881871FF9CFF5411EBFEB7FF1F" box="[251,280,223,249]" captionStart="FIGURES 21‒28" captionStartId="23.[151,264,1890,1914]" captionTargetBox="[151,1436,481,1867]" captionTargetId="figure-156@23.[151,1436,478,1867]" captionTargetPageId="23" captionText="FIGURES 21‒28. Male genitalia of tortricid species from Chile and Argentina. 21. Terifrona fronteria, USNM slide 145,517. 22. Transtilliana biloba, USNM slide 154,846. 23. Vargasia karsholti, Horak slide GP 733. 24. Hamiamia trancasiana, USNM slide 92,490. 25. Varifula fulvaria, USNM slide 154,848. 26. Varifula geomops, USNM slide 68,492. 27. Proeulia hasta, furca attached to phallus, USNM slide 154,802. 28. Proeulia hasta, furca in situ, USNM slide 81,233." figureDoi="http://doi.org/10.5281/zenodo.14389851" httpUri="https://zenodo.org/record/14389851/files/figure.png" pageId="11" pageNumber="62">26</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="A2DC87861871FF9CFF381013FACEFE10" pageId="11" pageNumber="62" type="discussion">
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<paragraph id="EA79D40D1871FF9CFF381013FACEFE10" blockId="11.[151,1437,295,1222]" pageId="11" pageNumber="62">
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<emphasis id="D8B2081F1871FF9CFF381013FE1BFEA7" bold="true" box="[151,436,295,321]" pageId="11" pageNumber="62">Remarks and diagnosis</emphasis>
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.
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<taxonomicName id="2DC6AF8E1871FF9CFE691013FD21FEA7" authorityName="Brown" authorityYear="2024" box="[454,654,295,321]" class="Insecta" family="Tortricidae" genus="Varifula" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="geomops">
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<emphasis id="D8B2081F1871FF9CFE691013FD21FEA7" box="[454,654,295,321]" italics="true" pageId="11" pageNumber="62">Varifula geomops</emphasis>
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</taxonomicName>
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was originally recognized by Nicolas Obraztsov as an undescribed species, and he placed it in his manuscript genus “Apoeulia” (in 1964 based on specimen labels), long before Razowski’s description of
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<taxonomicName id="2DC6AF8E1871FF9CFE11105BFDB7FE6F" authorityName="Razowski" authorityYear="1995" box="[446,536,367,393]" class="Insecta" family="Tortricidae" genus="Varifula" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="genus">
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<emphasis id="D8B2081F1871FF9CFE11105BFDB7FE6F" box="[446,536,367,393]" italics="true" pageId="11" pageNumber="62">Varifula</emphasis>
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</taxonomicName>
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. In comparison to
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<taxonomicName id="2DC6AF8E1871FF9CFD481044FCF5FE6F" authorityName="Blanchard" authorityYear="1852" box="[743,858,367,393]" class="Insecta" family="Tortricidae" genus="Tortrix" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="fulvaria">
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<emphasis id="D8B2081F1871FF9CFD481044FCF5FE6F" box="[743,858,367,393]" italics="true" pageId="11" pageNumber="62">V. fulvaria</emphasis>
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</taxonomicName>
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, the forewing is conspicuously shorter in length and has a more reticulated pattern. In the male genitalia of
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<taxonomicName id="2DC6AF8E1871FF9CFCA910A7FC63FE4B" authorityName="Brown" authorityYear="2024" box="[774,972,403,429]" class="Insecta" family="Tortricidae" genus="Varifula" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="geomops">
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<emphasis id="D8B2081F1871FF9CFCA910A7FC63FE4B" box="[774,972,403,429]" italics="true" pageId="11" pageNumber="62">Varifula geomops</emphasis>
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</taxonomicName>
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, the transtilla has a larger folded median region with a broadly U-shaped ventral margin; the valvae are more parallel-sided throughout; and the termination of the sacculus is indicated by a small angle at the ventral margin of the valva, which is lacking in
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<taxonomicName id="2DC6AF8E1871FF9CFB4910E8FAF4FE13" authorityName="Blanchard" authorityYear="1852" box="[1254,1371,475,501]" class="Insecta" family="Tortricidae" genus="Tortrix" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="fulvaria">
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<emphasis id="D8B2081F1871FF9CFB4910E8FAF4FE13" box="[1254,1371,475,501]" italics="true" pageId="11" pageNumber="62">V. fulvaria</emphasis>
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</taxonomicName>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A2DC87861871FF9CFF6810CBFBEAFBD3" pageId="11" pageNumber="62" type="description">
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<paragraph id="EA79D40D1871FF9CFF6810CBFDB9FDD8" blockId="11.[151,1437,295,1222]" pageId="11" pageNumber="62">
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<emphasis id="D8B2081F1871FF9CFF6810CBFEFEFDFF" bold="true" box="[199,337,511,537]" pageId="11" pageNumber="62">Description</emphasis>
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. Head: Scales of vertex and frons cream; scales of labial palpus mostly cream on outer and inner surface; scaling of antenna cream.
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</paragraph>
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<paragraph id="EA79D40D1871FF9CFF681373FE0AFD17" blockId="11.[151,1437,295,1222]" pageId="11" pageNumber="62">
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Thorax: Dorsum scaling cream. Forewing length
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<quantity id="2D3E79E81871FF9CFD591373FCE2FD84" box="[758,845,583,610]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="11" pageNumber="62" unit="mm" value="9.5">9.5 mm</quantity>
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(n = 2); ground color pale cream, faintly reticulated throughout with pale gold-brown scales, densest in distal 0.5 of wing, creating faint, narrow, incomplete, median and terminal fasciae; fringe pale cream. Underside brownish cream, darker than upperside, reticulated throughout. Hindwing concolorous with forewing, with faint, pale gold-brown mottling; fringe cream. Underside dingy white with pale gray mottling.
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</paragraph>
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<paragraph id="EA79D40D1871FF9CFF6813CFFD7EFC0B" blockId="11.[151,1437,295,1222]" pageId="11" pageNumber="62">
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Abdomen: Male genitalia (
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<figureCitation id="72FDC8881871FF9CFE5413CFFDE1FCF3" box="[507,590,763,789]" captionStart="FIGURES 21‒28" captionStartId="23.[151,264,1890,1914]" captionTargetBox="[151,1436,481,1867]" captionTargetId="figure-156@23.[151,1436,478,1867]" captionTargetPageId="23" captionText="FIGURES 21‒28. Male genitalia of tortricid species from Chile and Argentina. 21. Terifrona fronteria, USNM slide 145,517. 22. Transtilliana biloba, USNM slide 154,846. 23. Vargasia karsholti, Horak slide GP 733. 24. Hamiamia trancasiana, USNM slide 92,490. 25. Varifula fulvaria, USNM slide 154,848. 26. Varifula geomops, USNM slide 68,492. 27. Proeulia hasta, furca attached to phallus, USNM slide 154,802. 28. Proeulia hasta, furca in situ, USNM slide 81,233." figureDoi="http://doi.org/10.5281/zenodo.14389851" httpUri="https://zenodo.org/record/14389851/files/figure.png" pageId="11" pageNumber="62">Fig. 26</figureCitation>
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) with rod-like uncus; socius moderately long, pendant, densely clothes in fine hairs; gnathos well defined with pointed terminal plate representing the fusion of the lateral arms; transtilla with conspicuous, subrectangular folded region medially with a broadly U-shaped concavity, and with irregular rows of tiny spines; valva mostly parallel-sided from base to ca. distal 0.1; weakly upcurved, sacculus a simple, linear, sclerotized region along venter of basal 0.4 of valva, with a small angle at its distal termination, lacking a free distal process; phallus comparative large, robust, rather short; vesica with tiny patch of 10–12 extremely small non-deciduous cornuti. Female genitalia unknown.
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</paragraph>
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<paragraph id="EA79D40D1871FF9CFF6812C3FBD2FBF7" blockId="11.[151,1437,295,1222]" box="[199,1149,1015,1042]" pageId="11" pageNumber="62">
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<emphasis id="D8B2081F1871FF9CFF6812C3FEA3FBF7" bold="true" box="[199,268,1015,1041]" pageId="11" pageNumber="62">Types</emphasis>
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.
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<materialsCitation id="5AAEDE501871FF9CFEB412C3FBD2FBF7" box="[283,1149,1015,1042]" collectingDate="1953-12-05" collectorName="L. Pena" country="Chile" location="Chovellen" pageId="11" pageNumber="62" specimenCount="1" specimenCount-male="1" stateProvince="Maule" typeStatus="holotype">
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<typeStatus id="357D6AAF1871FF9CFEB412C3FE2CFBF7" box="[283,387,1015,1041]" pageId="11" pageNumber="62" type="holotype">Holotype</typeStatus>
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<specimenCount id="FCC01F841871FF9CFE2512CCFE0EFBF4" box="[394,417,1016,1042]" count="1" pageId="11" pageNumber="62" type="male">♂</specimenCount>
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,
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||
<collectingCountry id="92D1949D1871FF9CFE0212C3FE43FBF7" box="[429,492,1015,1041]" name="Chile" pageId="11" pageNumber="62">Chile</collectingCountry>
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,
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<collectingRegion id="28021AEF1871FF9CFE5712C3FD35FBF7" box="[504,666,1015,1041]" country="Chile" name="Maule" pageId="11" pageNumber="62">Maule Region</collectingRegion>
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,
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||
<location id="EF1982D61871FF9CFD0A12C3FCB4FBF7" LSID="urn:lsid:plazi:treatment:626F651B1871FF9CFF3811A3FB70FB20:EF1982D61871FF9CFD0A12C3FCB4FBF7" box="[677,795,1015,1041]" country="Chile" name="Chovellen" pageId="11" pageNumber="62" stateProvince="Maule">Chovellén</location>
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,
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||
<date id="9E78F2CD1871FF9CFC8912C3FC05FBF4" box="[806,938,1015,1042]" pageId="11" pageNumber="62" value="1953-12-05">
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||
<collectingDate id="8E3C0B251871FF9CFC8912C3FC05FBF4" box="[806,938,1015,1042]" pageId="11" pageNumber="62" value="1953-12-05">5 Dec 1953</collectingDate>
|
||
</date>
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||
,
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||
<collectorName id="4733B1DB1871FF9CFC1A12CCFBA5FBF4" box="[949,1034,1016,1042]" pageId="11" pageNumber="62">L. Peña</collectorName>
|
||
(
|
||
<taxonomicName id="2DC6AF8E1871FF9CFBB512C3FBDEFBF7" box="[1050,1137,1015,1041]" kingdom="Usnm" pageId="11" pageNumber="52" rank="kingdom">USNM</taxonomicName>
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||
).
|
||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="EA79D40D1871FF9CFF681528FBEAFBD3" blockId="11.[151,1437,295,1222]" box="[199,1093,1051,1078]" pageId="11" pageNumber="62">
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||
<materialsCitation id="5AAEDE501871FF9CFF681528FBEEFBD3" box="[199,1089,1051,1078]" collectingDate="1953-12-05" collectorName="L. Pena" country="Chile" location="Chovellen" pageId="11" pageNumber="62" specimenCount="1" specimenCount-male="1" stateProvince="Maule" typeStatus="paratype">
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||
<typeStatus id="357D6AAF1871FF9CFF681528FE86FBD0" box="[199,297,1052,1078]" pageId="11" pageNumber="62" type="paratype">Paratype</typeStatus>
|
||
(
|
||
<specimenCount id="FCC01F841871FF9CFE95152FFEF0FBD3" box="[314,351,1051,1077]" count="1" pageId="11" pageNumber="62" type="male">1♂</specimenCount>
|
||
).
|
||
<collectingCountry id="92D1949D1871FF9CFEDB152FFE1AFBD3" box="[372,437,1051,1077]" name="Chile" pageId="11" pageNumber="62">Chile</collectingCountry>
|
||
:
|
||
<collectingRegion id="28021AEF1871FF9CFE10152FFDCCFBD3" box="[447,611,1051,1077]" country="Chile" name="Maule" pageId="11" pageNumber="62">Maule Region</collectingRegion>
|
||
:
|
||
<location id="EF1982D61871FF9CFDC2152FFD4CFBD3" LSID="urn:lsid:plazi:treatment:626F651B1871FF9CFF3811A3FB70FB20:EF1982D61871FF9CFDC2152FFD4CFBD3" box="[621,739,1051,1077]" country="Chile" name="Chovellen" pageId="11" pageNumber="62" stateProvince="Maule">Chovellén</location>
|
||
,
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||
<date id="9E78F2CD1871FF9CFD40152FFCDCFBD0" box="[751,883,1051,1078]" pageId="11" pageNumber="62" value="1953-12-05">
|
||
<collectingDate id="8E3C0B251871FF9CFD40152FFCDCFBD0" box="[751,883,1051,1078]" pageId="11" pageNumber="62" value="1953-12-05">5 Dec 1953</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="4733B1DB1871FF9CFCD11528FC7CFBD0" box="[894,979,1052,1078]" pageId="11" pageNumber="62">L. Peña</collectorName>
|
||
(
|
||
<taxonomicName id="2DC6AF8E1871FF9CFC4D152FFB96FBD3" box="[994,1081,1051,1077]" kingdom="Usnm" pageId="11" pageNumber="52" rank="kingdom">USNM</taxonomicName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A2DC87861871FF9CFF68150BFC2CFB9B" pageId="11" pageNumber="62" type="distribution">
|
||
<paragraph id="EA79D40D1871FF9CFF68150BFC2CFB9B" blockId="11.[151,1437,295,1222]" pageId="11" pageNumber="62">
|
||
<emphasis id="D8B2081F1871FF9CFF68150BFE46FBBF" bold="true" box="[199,489,1087,1113]" pageId="11" pageNumber="62">Distribution and biology</emphasis>
|
||
.
|
||
<taxonomicName id="2DC6AF8E1871FF9CFE59150BFD15FBBF" authorityName="Brown" authorityYear="2024" box="[502,698,1087,1113]" class="Insecta" family="Tortricidae" genus="Varifula" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="geomops">
|
||
<emphasis id="D8B2081F1871FF9CFE59150BFD15FBBF" box="[502,698,1087,1113]" italics="true" pageId="11" pageNumber="62">Varifula geomops</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="357D6AAF1871FF9CFC6A1574FC5AFBBC" box="[965,1013,1088,1114]" pageId="11" pageNumber="62">type</typeStatus>
|
||
locality in the
|
||
<collectingRegion id="28021AEF1871FF9CFB37150BFA99FBBF" box="[1176,1334,1087,1113]" country="Chile" name="Maule" pageId="11" pageNumber="62">Maule Region</collectingRegion>
|
||
of
|
||
<collectingCountry id="92D1949D1871FF9CFAF7150BFA38FBBF" box="[1368,1431,1087,1113]" name="Chile" pageId="11" pageNumber="62">Chile</collectingCountry>
|
||
, at an elevation of about
|
||
<specimenCount id="FCC01F841871FF9CFE0A1557FE4AFB98" box="[421,485,1123,1150]" count="75" pageId="11" pageNumber="62" type="generic">
|
||
<quantity id="2D3E79E81871FF9CFE0A1557FE4EFB98" box="[421,481,1123,1150]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="62" unit="m" value="75.0">75 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Nothing is known of the life history.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A2DC87861871FF9CFF6815B3FB70FB20" pageId="11" pageNumber="62" type="etymology">
|
||
<paragraph id="EA79D40D1871FF9CFF6815B3FB70FB20" blockId="11.[151,1437,295,1222]" pageId="11" pageNumber="62">
|
||
<emphasis id="D8B2081F1871FF9CFF6815B3FEE7FB47" bold="true" box="[199,328,1159,1185]" pageId="11" pageNumber="62">Etymology</emphasis>
|
||
. This species was originally recognized as distinct by Nicholas Obraztsov, who used the name “Apoeulia geomops” on its genitalia slide label. Hence, it seems fitting to adopt his species name.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |