266 lines
31 KiB
XML
266 lines
31 KiB
XML
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<mods:title id="E7AD3A744FD67759B17EA33386E875CC">Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data</mods:title>
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<mods:namePart id="712CE03B3ABC72279A2C9B3541C5E75A">Maddison, David R.</mods:namePart>
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<mods:affiliation id="ACC70CFB81844E74F6E442DB7957098F">Center for Comparative Genomics, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:affiliation id="96669AC63934E4C732DF8F5707DBD968">Department of Entomology, University of Wisconsin, Madison, WI 53706, USA</mods:affiliation>
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<mods:affiliation id="7D337172BAE99E4E86A29572A2C5FF45">Institute of Biosciences, University of Rostock, Universitaetsplatz 2, D- 18055 Rostock, Germany</mods:affiliation>
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<mods:affiliation id="07DC341A6B0ED8336B091627E499DBD0">Department of Entomology, University of Arizona, Tucson, AZ 85721 - 0036, USA</mods:affiliation>
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<mods:affiliation id="4A8C218DFA1C084CD39A85F2F87FEC0D">Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
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<mods:namePart id="46D19D19DEEB9185B31D0371A4C117BB">Archambeault, Sophie L.</mods:namePart>
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<mods:affiliation id="25AF8ED205252C3CAFCA54848A668ECA">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA & University of California, Berkeley, 142 Weill Hall # 3200, Berkeley, CA 94720, USA</mods:affiliation>
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<mods:affiliation id="EF0EBB1413823E6260DD9CD17D8F0223">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA & University of California, Santa Cruz, Long Marine Lab, 117 McAllister Way, Santa Cruz, CA 95060, USA</mods:affiliation>
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<treatment id="6226343FCCF3514DB574B8B0341021C8" ID-GBIF-Taxon="182754136" LSID="urn:lsid:zoobank.org:act:12F2832A-C3A8-44B3-9909-32482CEBFE6B" httpUri="http://treatment.plazi.org/id/6226343FCCF3514DB574B8B0341021C8" lastPageNumber="41" pageId="0" pageNumber="41" scope_family="Carabidae" scope_order="Coleoptera">
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<subSubSection id="DA146D1478D7E58E223240FDF7542F9B" pageId="0" pageNumber="41" type="nomenclature">
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<paragraph id="CF94416A6F3B9485241AFB51EE13370E" pageId="0" pageNumber="41">
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Subgenus
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<taxonomicName id="255B8351AFEA0229B964AEE98834D203" ID-CoL="8M55R" LSID="http://zoobank.org/12F2832A-C3A8-44B3-9909-32482CEBFE6B" authority="Kavanaugh," authorityName="Kavanaugh" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria (Erwinebria)" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Erwinebria">Nebria Erwinebria Kavanaugh,</taxonomicName>
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<taxonomicNameLabel id="F8080130D60D5E22737989CB3A11BFA1" pageId="0" pageNumber="41">subgen. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="3B7AA90FB0EDCA1F3E8A6DBEF3382667" pageId="0" pageNumber="41" type="reference_group">
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<paragraph id="F2F29D85A202D8A4532E5FBF9D0B1334" pageId="0" pageNumber="41">
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<taxonomicName id="59C8A1EE3CBF277D31332E33931AF60F" authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria gregaria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gregaria" status="subgen. nov.">Nebria gregaria</taxonomicName>
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= "
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<taxonomicName id="9B13F3F432542C908D63C2E5C88735E7" authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria gregaria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gregaria" status="subgen. nov.">Nebria gregaria</taxonomicName>
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group" sensu
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<bibRefCitation id="E76302540A8CA91FD3F64101D5AA673A" author="Lindroth, CH" journalOrPublisher="Proceedings of a symposium (28 August, 1996, Florence, Italy). XX International Congress of Entomology. Museo Regionale di Scienze Naturali, Torino" pageId="0" pageNumber="41" refId="B127" refString="Lindroth, CH, 1961. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 2. Opuscula Entomologica Supplementum No.20: 1-200." title="The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 2. Opuscula Entomologica Supplementum No. 20: 1 - 200." year="1961">Lindroth 1961</bibRefCitation>
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</paragraph>
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<paragraph id="59445CB48EA15B554E6B47CD72B9992A" pageId="0" pageNumber="41">
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<taxonomicName id="DCFD41060A18CE203F89F8286B9F1BDB" authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus" status="subgen. nov.">Reductonebria</taxonomicName>
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Shilenkov, 1975 (in part); = "
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<taxonomicName id="40BFA016EE1E3C380330C1E48990177C" authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria sahlbergi" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="sahlbergi" status="subgen. nov.">
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<emphasis id="77AF170397B8E8E1A33694E474E215FB" italics="true" pageId="0" pageNumber="41">Nebria sahlbergi</emphasis>
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</taxonomicName>
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" + "
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<taxonomicName id="8C01B5877DE7B0668D5B97C9CD666D7A" authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria gregaria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gregaria" status="subgen. nov.">Nebria gregaria</taxonomicName>
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" groups sensu
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<bibRefCitation id="D542F7A9FDB497BE75C25E3F4F15B477" author="Ledoux, G" journalOrPublisher="Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon" pageId="0" pageNumber="41" refId="B121" refString="Ledoux, G, Roux, P, 2005. Nebria (Coleoptera, Nebriidae). Faune mondiale. Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon" title="Nebria (Coleoptera, Nebriidae). Faune mondiale." year="2005">Ledoux and Roux 2005</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4384A1A1C4E27376EFBB7B125FFD8BA4" pageId="0" pageNumber="41" type="type species">
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<paragraph id="D3C79DC876A281A4DA1E0B99C267B0BA" pageId="0" pageNumber="41">Type species.</paragraph>
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<paragraph id="172F0D3390559CCA82B684FAED207329" pageId="0" pageNumber="41">
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<taxonomicName id="E327E25218B53BE78B4F1ECE36B178C2" authorityName="Fischer von Waldheim" authorityYear="1828" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria sahlbergii" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="sahlbergii">
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<emphasis id="E2CC4B305D7924254F207BBDE7B34B4E" italics="true" pageId="0" pageNumber="41">Nebria sahlbergii</emphasis>
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</taxonomicName>
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Fischer von Waldheim, 1828:254, by present designation.
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</paragraph>
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<subSubSection id="06316EF30AF65168A877C3A01669ABFC" pageId="0" pageNumber="41" type="diagnosis">
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<paragraph id="195EE778283B19CC028DB6F45A823049" pageId="0" pageNumber="41">Diagnosis.</paragraph>
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<paragraph id="2A1E1D859C89569C9872B6394EB75AD0" pageId="0" pageNumber="41">
|
||
Body size small to medium, SBL = 7.1 to 11.7 mm. Head width medium to slightly broadened, not constricted behind eyes; vertex with a pair of paramedial pale spots and a single pair of supraorbital setae. Eyes slightly reduced to moderate in size, moderately to markedly convex. Antennal scape with one (two in very few specimens) subapicodorsal seta; antennomeres 3 and 4 not laterally compressed, without extra setae. Labrum with three pairs of apical setae. Maxillary stipes typical for genus, with setae inserted flush on smooth surface. Penultimate labial palpomere with three setae. Pronotum with midlateral setae absent, a single pair of basolateral setae present. Elytral intervals smooth, without macrosculpture, interval 3 with three to eight or more setiferous pores, intervals 5 and 7 without setiferous pores, interval 3 not or only faintly catenate, intervals 5 and 7 not catenate. Hindwings full-sized or slightly to markedly reduced in size. Metepisterna smooth or faintly punctulate. Protarsomeres 1-3 expanded in males; mesotarsomeres 2-4 longer than their apical width; tarsi dorsally glabrous or with a few fine setae. Abdominal sternites IV-VI with two to four pairs of posterior paramedial setae, without paralateral setae. Median lobe of male aedeagus sclerotized dorsally at least to midlength on shaft, moderately to markedly deflected right in dorsal aspect; basal bulb expanded, quadrate, broadly open basally and closed dorsally, without a sagittal aileron present at base or with only a lightly sclerotized collar; mid-shaft parallel-sided or slightly to moderately narrowed basally in lateral aspect, slightly compressed in cross-section, with right lateral face unmodified; apical orifice markedly to extremely deflected right. Right paramere slender and moderately long. Female valvifers with vestiture (a few specimens of some species without vestiture); gonopods VIII fused to dorsomedial bases of gonocoxae; gonocoxae with ventral diagonal row of setiform setae and mediodorsal row of setae present. Bursa copulatrix without dorsal sclerites in vestibular chamber and with longitudinal axis of the bursa moderately to markedly sigmoid dorsally in lateral aspect; spermathecal chamber broadly cordate in dorsal aspect, without (in
|
||
<taxonomicName id="0F1CA0B18D611BC06D3582ED917A401D" authorityName="Van Dyke" authorityYear="1925" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria lyelli" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="lyelli">
|
||
<emphasis id="1DB59E5498EF4408FB68F35058CE9106" italics="true" pageId="0" pageNumber="41">Nebria lyelli</emphasis>
|
||
</taxonomicName>
|
||
Van Dyke, 1925 and
|
||
<taxonomicName id="6FF85A573B278AB64BBA1DBD097B391D" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria quileute" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="quileute">
|
||
<emphasis id="ECB67DC37BCDE301CEE2743422E23562" italics="true" pageId="0" pageNumber="41">Nebria quileute</emphasis>
|
||
</taxonomicName>
|
||
Kavanaugh, 1979) or with a small to large midline sclerotized dorsal plate, insemination duct not sclerotized; spermathecal duct medium-length to slightly long, uniform in diameter throughout or nearly so, inserted basodorsally on spermathecal chamber; spermathecal reservoir of medium length.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0E79B54704233499DBF0BB2A607510DB" pageId="0" pageNumber="41" type="etymology">
|
||
<paragraph id="4E75255E1344E5050BFC7E3905CDD8C6" pageId="0" pageNumber="41">Etymology.</paragraph>
|
||
<paragraph id="CDCE76A1D9F091B52CEFC5D9A2C1CBE1" pageId="0" pageNumber="41">
|
||
The subgeneric epithet is a noun of feminine gender and a combination of the surname of Terry
|
||
<taxonomicName id="94CBA2A39AF9FC0AE5EDF34B4AFEBCDB" genus="L." lsidName="L." pageId="0" pageNumber="41" rank="genus">L.</taxonomicName>
|
||
Erwin, in whose honor we name this subgenus, and the genus name,
|
||
<taxonomicName id="A8E0D9468B4C905DB760744D8FB08A73" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FD072321A7902187DF298890E209C6C8" italics="true" pageId="0" pageNumber="41">Nebria</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9552670B57AAC0DD840FC87ED9B57E6B" pageId="0" pageNumber="41" type="remarks">
|
||
<paragraph id="5D6ACA3898131524284A13F76C75F346" pageId="0" pageNumber="41">Remarks.</paragraph>
|
||
<paragraph id="6B2CB680E0F615F78C34BA79C48EBDC0" pageId="0" pageNumber="41">
|
||
Members of this group are easily identified as members of the
|
||
<taxonomicName id="79BA2E7BA70DAA67EC83DF6DBAD1D868" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BE1FC100C46BC75AED253D5AD0AE672C" italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
|
||
</taxonomicName>
|
||
Complex using the
|
||
<bibRefCitation id="C333A0E914B5E362626D698323BDA4C8" author="Ledoux, G" journalOrPublisher="Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon" pageId="0" pageNumber="41" refId="B121" refString="Ledoux, G, Roux, P, 2005. Nebria (Coleoptera, Nebriidae). Faune mondiale. Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon" title="Nebria (Coleoptera, Nebriidae). Faune mondiale." year="2005">
|
||
Ledoux and
|
||
<normalizedToken id="F3EDF9DBD616D2B63ABB9166AB4368DB" originalValue="Roux’s">Roux's</normalizedToken>
|
||
(2005)
|
||
</bibRefCitation>
|
||
key to subgenera. Features that distinguish them from members of subgenus
|
||
<taxonomicName id="A671494167BB0C00D298CB10FD20E72A" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reductonebria (Insulanebria)" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Insulanebria">Reductonebria Insulanebria</taxonomicName>
|
||
have been discussed above for that taxon. We have not yet identified any external morphological features that consistently distinguish all members of all species of
|
||
<taxonomicName id="322D269ECA9829A5F1A8EE8A75341F5F" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9F4CE72C551EFCE0FFBB370CE2779B37" italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
|
||
</taxonomicName>
|
||
from all
|
||
<taxonomicName id="B963619EAC2B023B7E5D8A6D067A8A37" genus="Erwinebria" lsidName="Erwinebria" pageId="0" pageNumber="41" rank="genus">
|
||
<emphasis id="70A8AC72A1C5FE4F42B75A3D830890E9" italics="true" pageId="0" pageNumber="41">Erwinebria</emphasis>
|
||
</taxonomicName>
|
||
members, but there are several distinguishing internal features evident in both sexes. The mid-shaft is parallel-sided or slightly to moderately narrowed basally and the apical orifice is markedly to extremely deflected right in
|
||
<taxonomicName id="0AFFBEF3F36F17090D18704EABFB9259" genus="Erwinebria" lsidName="Erwinebria" pageId="0" pageNumber="41" rank="genus">
|
||
<emphasis id="458E84FDE5E51670BA28BA7D23301211" italics="true" pageId="0" pageNumber="41">Erwinebria</emphasis>
|
||
</taxonomicName>
|
||
males. In
|
||
<taxonomicName id="18BEE66AD1F2918B96046EC2D5B88050" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="616A02F80799249D4462268CAF9EF698" italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
|
||
</taxonomicName>
|
||
males, the mid-shaft is markedly narrowed apically and the apical orifice is only moderately deflected right. As in most other
|
||
<taxonomicName id="D68856F06F9E24824F403EA187B2E698" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8EAF294ECD6C756581BD2B10612719A3" italics="true" pageId="0" pageNumber="41">Nebria</emphasis>
|
||
</taxonomicName>
|
||
examined, the gonopods VIII in
|
||
<taxonomicName id="1F7CB7F3BDA05BF6B4BCFCA3AF33BCD8" genus="Erwinebria" lsidName="Erwinebria" pageId="0" pageNumber="41" rank="genus">
|
||
<emphasis id="92148913DBDDF33D9C48B9780B40C335" italics="true" pageId="0" pageNumber="41">Erwinebria</emphasis>
|
||
</taxonomicName>
|
||
females are fused dorsomedially with the bases of gonocoxae. In
|
||
<taxonomicName id="CFCCBC3E51D3CCC9423455F610AFF90B" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="102BB9D8867A742F8A0CACAE84397D70" italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
|
||
</taxonomicName>
|
||
females, these gonopods are distinctly separate from, although in the same position in relation to, the gonocoxae. The spermathecal chamber is broadly cordate in dorsal aspect in
|
||
<taxonomicName id="244705828E47E3336799EAD9C87B4CA7" genus="Erwinebria" lsidName="Erwinebria" pageId="0" pageNumber="41" rank="genus">
|
||
<emphasis id="C9790E68DDBE4016195B04E25A9EDBEB" italics="true" pageId="0" pageNumber="41">Erwinebria</emphasis>
|
||
</taxonomicName>
|
||
females, but somewhat broadly ovoid in
|
||
<taxonomicName id="06A9F0D979104AB5FA3937713CB25B44" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F5BF872BAD4FE34C8E118EDA76FC2C5" italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
|
||
</taxonomicName>
|
||
females. Although most members of both of the subgenera have sclerotized plates in the dorsal or dorsoapical wall of the spermathecal chamber of bursa copulatrix, they are of a different form in the two groups. In
|
||
<taxonomicName id="1BFC65EC208A9D4E3780585F1AF0676C" genus="Erwinebria" lsidName="Erwinebria" pageId="0" pageNumber="41" rank="genus">
|
||
<emphasis id="7A52D7083AE6C438376B98FC4107BA6B" italics="true" pageId="0" pageNumber="41">Erwinebria</emphasis>
|
||
</taxonomicName>
|
||
females (except in
|
||
<taxonomicName id="30ACAB9238AA330558323DC790CD1181" lsidName="N. lyelli" pageId="0" pageNumber="41" rank="species" species="lyelli">
|
||
<emphasis id="688BED1623A003F2D99A1395DEDFD437" italics="true" pageId="0" pageNumber="41">N. lyelli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DA8E83705A008DB5D05BFDE2088B90D2" lsidName="N. quileute" pageId="0" pageNumber="41" rank="species" species="quileute">
|
||
<emphasis id="FC1DE60DC6158E0CB55854DE5A3B5BE6" italics="true" pageId="0" pageNumber="41">N. quileute</emphasis>
|
||
</taxonomicName>
|
||
, which have no dorsal sclerites), the dorsal sclerite is variously formed as a small to large domed, horseshoe- or saddle-shaped plate associated with the point insertion of the spermathecal duct on the chamber. In
|
||
<taxonomicName id="E0A96188282C4AEC067A8CCE89558FD7" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="024001044AF818CA7B6CA37400910151" italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
|
||
</taxonomicName>
|
||
females, the sclerotized area is predominately an apicodorsal cap on the spermathecal chamber. This cap sclerite may be relatively small, round, and smooth, as in
|
||
<taxonomicName id="B90C6A72E74B4A378D87B39ACFE0A16B" lsidName="N. ochotica" pageId="0" pageNumber="41" rank="species" species="ochotica">
|
||
<emphasis id="2290F037A3655E1479CF99D0A15E223D" italics="true" pageId="0" pageNumber="41">N. ochotica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="BD04BB2536D048806D0EA80D5AD89BCB" lsidName="N. mannerheimii" pageId="0" pageNumber="41" rank="species" species="mannerheimii">
|
||
<emphasis id="6B7C3CA9766C4F88547BB1289016A109" italics="true" pageId="0" pageNumber="41">N. mannerheimii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AC56C764C61BAFA1108078BCA20F38F6" lsidName="N. darlingtoni" pageId="0" pageNumber="41" rank="species" species="darlingtoni">
|
||
<emphasis id="DF4566D78308FC47B6E7DA0EEAB448EB" italics="true" pageId="0" pageNumber="41">N. darlingtoni</emphasis>
|
||
</taxonomicName>
|
||
females, or larger, crenulated rather than smooth, and/or expanded laterally or basally to or beyond the insertion point of the spermathecal duct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="379B719CB2B07740AE24F8F0D90D15BA" pageId="0" pageNumber="41" type="distribution">
|
||
<paragraph id="932003030DA3A40D91D13E806E35FDA4" pageId="0" pageNumber="41">Known distribution and diversity.</paragraph>
|
||
<paragraph id="4F8BC318959BBE74DA7422FE24278473" pageId="0" pageNumber="41">The geographical range of this clade covers much of western North America, from the outer Aleutian Islands in the northwest, south to eastcentral California, northern Arizona, and New Mexico and east to the eastern edge of the Rocky Mountain system from Yukon Territory in Canada south to northcentral New Mexico.</paragraph>
|
||
<paragraph id="BE77F3F922BAF725F4CC47AB5F2BC0C4" pageId="0" pageNumber="41">Twenty-one species in this clade have been described.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |