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<document id="A9A8721BEB89A409F45046D6043EEEE2" ID-CLB-Dataset="298869" ID-DOI="10.5281/zenodo.12520585" ID-GBIF-Dataset="3a454625-d8aa-467e-a56b-4721cd06356c" ID-ISSN="1407-8619" ID-Zenodo-Dep="12520585" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719251034552" checkinUser="felipe" docAuthor="Anichtchenko, Alexander &amp; Medina, Milton Norman D." docDate="2021" docId="62064372A7357738F989FA9FFB72FBF5" docLanguage="en" docName="BalticJColeopterol.21.1.77-88.pdf" docOrigin="Baltic Journal of Coleopterology 21 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.3:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="3" docTitle="Dromoceryx obscurus Anichtchenko &amp; Medina 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="80" masterDocId="9E3F3B0AA734773BF96AFFDAF961FFE2" masterDocTitle="A new species of the genus Dromoceryx Schmidt-Goebel, 1846 (Coleoptera: Carabidae: Lebiini) from the Philippines" masterLastPageNumber="80" masterPageNumber="77" pageNumber="78" updateTime="1719257709814" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="44E4D7DDA0D2AC93D4CCFD2621357425">A new species of the genus Dromoceryx Schmidt-Goebel, 1846 (Coleoptera: Carabidae: Lebiini) from the Philippines</mods:title>
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<mods:namePart id="D8767274F0CC31F109EC796F39F07446">Anichtchenko, Alexander</mods:namePart>
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<mods:namePart id="362417E3D2A4759B10740CF39E720BE8">Medina, Milton Norman D.</mods:namePart>
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<mods:title id="124B46B13A894A4242F26049F402E82A">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="00FE951BAC158962BECC6D38A78A2464">2021</mods:date>
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<treatment id="62064372A7357738F989FA9FFB72FBF5" ID-DOI="http://doi.org/10.5281/zenodo.12520587" ID-GBIF-Taxon="233760276" ID-Zenodo-Dep="12520587" LSID="urn:lsid:plazi:treatment:62064372A7357738F989FA9FFB72FBF5" httpUri="http://treatment.plazi.org/id/62064372A7357738F989FA9FFB72FBF5" lastPageId="3" lastPageNumber="80" pageId="1" pageNumber="78">
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<heading id="B1584508A735773AF989FA9FFBACFABF" bold="true" box="[227,717,1349,1373]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="78" reason="4">
<emphasis id="D8DB2E76A735773AF989FA9FFBACFABF" bold="true" box="[227,717,1349,1373]" pageId="1" pageNumber="78">
<taxonomicName id="2DAF89E7A735773AF989FA9FFB13FABF" authority="Anichtchenko" authorityName="Anichtchenko &amp; Medina" authorityYear="2021" box="[227,626,1349,1373]" class="Insecta" family="Carabidae" genus="Dromoceryx" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="obscurus" status="sp. nov.">
<emphasis id="D8DB2E76A735773AF989FA9FF8B0FABF" bold="true" box="[227,465,1349,1373]" italics="true" pageId="1" pageNumber="78">Dromoceryx obscurus</emphasis>
Anichtchenko
</taxonomicName>
,
<taxonomicNameLabel id="C3E8930DA735773AFB16FA9FFBACFABF" box="[636,717,1349,1373]" pageId="1" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<heading id="B1584508A735773AF989FAB2F841FA62" bold="true" box="[227,288,1384,1408]" fontSize="9" level="2" pageId="1" pageNumber="78" reason="6">
<figureCitation id="7294EEE1A735773AF989FAB2F841FA62" box="[227,288,1384,1408]" captionStart="Fig" captionStartId="2.[303,343,904,928]" captionText="Fig. 1. Habitus of Dromoceryx obscurus sp. n. Holotype. Scale bar = 1mm." pageId="1" pageNumber="78">
<emphasis id="D8DB2E76A735773AF989FAB2F841FA62" bold="true" box="[227,288,1384,1408]" pageId="1" pageNumber="78">Fig. 1</emphasis>
</figureCitation>
</heading>
</paragraph>
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<subSubSection id="A2B5A1EFA735773AF989FA77F85FF9A6" pageId="1" pageNumber="78" type="materials_examined">
<paragraph id="EA10F264A735773AF989FA77F85FF9A6" blockId="1.[227,737,1452,1604]" pageId="1" pageNumber="78">
<emphasis id="D8DB2E76A735773AF989FA77F8F3FA27" bold="true" box="[227,402,1453,1477]" pageId="1" pageNumber="78">Type material.</emphasis>
<materialsCitation id="5AC7F839A735773AF8F5FA76F82EF9E6" ID-GBIF-Occurrence="4899242301" collectingDate="2020-12" collectionCode="DUBC" location="Mindanao" municipality="Sandayong" pageId="1" pageNumber="78" specimenCount="1" stateProvince="Zamboanga del Norte" typeStatus="holotype">
<typeStatus id="35144CC6A735773AF8F5FA76FB66FA26" box="[415,519,1452,1476]" pageId="1" pageNumber="78" type="holotype">Holotype</typeStatus>
, male:
<collectingMunicipality id="0A74681EA735773AFB35FA76FBBDFA26" box="[607,732,1452,1476]" pageId="1" pageNumber="78">Sandayong</collectingMunicipality>
,
<collectingRegion id="286B3C86A735773AF989FA16F8A3FA06" box="[227,450,1484,1508]" country="Philippines" name="Zamboanga del Norte" pageId="1" pageNumber="78">Zamboanga del Norte</collectingRegion>
,
<location id="EF70A4BFA735773AF8A0FA16FB56FA06" LSID="urn:lsid:plazi:treatment:62064372A7357738F989FA9FFB72FBF5:EF70A4BFA735773AF8A0FA16FB56FA06" box="[458,567,1484,1508]" municipality="Sandayong" name="Mindanao" pageId="1" pageNumber="78" stateProvince="Zamboanga del Norte">Mindanao</location>
,
<date id="9E11D4A4A735773AFB2AFA16FB80FA06" box="[576,737,1484,1508]" pageId="1" pageNumber="78" value="2020-12">
<collectingDate id="8E552D4CA735773AFB2AFA16FB80FA06" box="[576,737,1484,1508]" pageId="1" pageNumber="78" value="2020-12">December 2020</collectingDate>
</date>
(
<collectionCode id="8CBE6AA1A735773AF981FA36F824F9E6" box="[235,325,1516,1540]" pageId="1" pageNumber="78">DUBC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="5AC7F839A735773AF835FA37F85BF9A6" ID-GBIF-Occurrence="4899242302" collectingDate="2020-08" collectionCode="DUBC" location="Mindanao" municipality="Sandayong" pageId="1" pageNumber="78" specimenCount="1" specimenCount-male="1" stateProvince="Zamboanga del Norte" typeStatus="paratype">
<typeStatus id="35144CC6A735773AF835FA37F8AEF9E7" box="[351,463,1517,1541]" pageId="1" pageNumber="78" type="paratype">Paratype</typeStatus>
,
<specimenCount id="FCA939EDA735773AF888FA36FB21F9E6" box="[482,576,1516,1540]" count="1" pageId="1" pageNumber="78" type="male">1 male</specimenCount>
:
<collectingMunicipality id="0A74681EA735773AFB38FA36FBBDF9E6" box="[594,732,1516,1540]" pageId="1" pageNumber="78">Sandayong</collectingMunicipality>
,
<collectingRegion id="286B3C86A735773AF989F9D6F8B1F9C6" box="[227,464,1548,1572]" country="Philippines" name="Zamboanga del Norte" pageId="1" pageNumber="78">Zamboanga del Norte</collectingRegion>
,
<location id="EF70A4BFA735773AF8B1F9D6FB2CF9C6" LSID="urn:lsid:plazi:treatment:62064372A7357738F989FA9FFB72FBF5:EF70A4BFA735773AF8B1F9D6FB2CF9C6" box="[475,589,1548,1572]" municipality="Sandayong" name="Mindanao" pageId="1" pageNumber="78" stateProvince="Zamboanga del Norte">Mindanao</location>
,
<date id="9E11D4A4A735773AFB3CF9D7FB81F9C6" box="[598,736,1548,1573]" pageId="1" pageNumber="78" value="2020-08">
<collectingDate id="8E552D4CA735773AFB3CF9D7FB81F9C6" box="[598,736,1548,1573]" pageId="1" pageNumber="78" value="2020-08">August 2020</collectingDate>
</date>
(
<collectionCode id="8CBE6AA1A735773AF980F9F6F852F9A6" box="[234,307,1580,1604]" pageId="1" pageNumber="78">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B5A1EFA735773AF989F9B5FACCFA96" pageId="1" pageNumber="78" type="diagnosis">
<paragraph id="EA10F264A735773AF989F9B5FACCFA96" blockId="1.[227,737,1647,1735]" lastBlockId="1.[774,1285,1276,1396]" pageId="1" pageNumber="78">
<emphasis id="D8DB2E76A735773AF989F9B5F838F965" bold="true" box="[227,345,1647,1671]" pageId="1" pageNumber="78">Diagnosis.</emphasis>
This species is readily distinguished from other
<taxonomicName id="2DAF89E7A735773AF837F94AFB58F945" box="[349,569,1679,1703]" pageId="1" pageNumber="78">
<emphasis id="D8DB2E76A735773AF837F94AF884F945" box="[349,485,1680,1703]" italics="true" pageId="1" pageNumber="78">Dromoceryx</emphasis>
species
</taxonomicName>
by the dark piceous to black main body coloration, and by small yellow spots on elytra. New species is most similar to
<taxonomicName id="2DAF89E7A735773AFA22FAC7FAA0FAD6" authorityName="Schmidt-Goebel" authorityYear="1846" box="[840,961,1308,1332]" class="Insecta" family="Carabidae" genus="Dromoceryx" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="D8DB2E76A735773AFA22FAC7FAA0FAD6" box="[840,961,1308,1332]" italics="true" pageId="1" pageNumber="78">D. dorsalis</emphasis>
</taxonomicName>
, the two species can be diagnosed easily by elytral pattern and microsculpture.
</paragraph>
</subSubSection>
<subSubSection id="A2B5A1EFA7357738FA6CFA46FB72FBF5" lastPageId="3" lastPageNumber="80" pageId="1" pageNumber="78" type="description">
<paragraph id="EA10F264A7357739FA6CFA46FA08FB55" blockId="1.[774,1285,1436,1748]" lastBlockId="2.[851,1361,1151,1207]" lastPageId="2" lastPageNumber="79" pageId="1" pageNumber="78">
<emphasis id="D8DB2E76A735773AFA6CFA46FAE8FA56" bold="true" box="[774,905,1436,1460]" pageId="1" pageNumber="78">Description.</emphasis>
Length
<quantity id="2D575F81A735773AFAB6FA46FD39FA56" box="[988,1112,1436,1460]" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.4" metricValueMin="3.3" pageId="1" pageNumber="78" unit="mm" value="3.3499999999999996" valueMax="3.4" valueMin="3.3">3.33.4 mm</quantity>
. Dorsum of head, clypeus and labrum black; mandibles dark brown; palpi and anten-nae testaceous.
<collectingRegion id="286B3C86A735773AFD32FA07FDD5FA16" box="[1112,1204,1501,1524]" country="Myanmar" name="Chin" pageId="1" pageNumber="78">Mentum</collectingRegion>
brown. Pronotum black, with brownish lateral margins. Elytral disc black, epipleura dark brown. Elytral pattern consist of two brown, semilunar humeral maculae, extended from lateral margin to third or second striae, and two apical macula on intervals 5-7, very narrowly joined along apical margin. Ventral surface black, except for testaceous meso-sternum, coxa and base of sternites I and II.
</paragraph>
<caption id="BED0A2ECA7367739FA39FDF9FD7CFDB9" pageId="2" pageNumber="79" startId="2.[851,891,547,571]" targetType="figure">
<paragraph id="EA10F264A7367739FA39FDF9FD7CFDB9" blockId="2.[851,1360,547,603]" pageId="2" pageNumber="79">
Fig. 3. Aedeagus of
<taxonomicName id="2DAF89E7A7367739FD4CFDFFFC72FDD9" authorityName="Anichtchenko &amp; Medina" authorityYear="2021" box="[1062,1299,547,572]" class="Insecta" family="Carabidae" genus="Dromoceryx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="obscurus" status="sp. nov.">
<emphasis id="D8DB2E76A7367739FD4CFDFFFC72FDD9" box="[1062,1299,547,572]" italics="true" pageId="2" pageNumber="79">Dromoceryx obscurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E8930DA7367739FC72FDFEFC31FDD9" box="[1304,1360,548,571]" pageId="2" pageNumber="79" rank="species">sp. n.</taxonomicNameLabel>
Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="BED0A2ECA7367739F845FC52FB32FC22" pageId="2" pageNumber="79" startId="2.[303,343,904,928]" targetType="figure">
<paragraph id="EA10F264A7367739F845FC52FB32FC22" blockId="2.[303,812,904,960]" pageId="2" pageNumber="79">
Fig. 1. Habitus of
<taxonomicName id="2DAF89E7A7367739F89DFC50FB89FC42" authorityName="Anichtchenko &amp; Medina" authorityYear="2021" box="[503,744,904,929]" class="Insecta" family="Carabidae" genus="Dromoceryx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="obscurus" status="sp. nov.">
<emphasis id="D8DB2E76A7367739F89DFC50FB89FC42" box="[503,744,904,929]" italics="true" pageId="2" pageNumber="79">Dromoceryx obscurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E8930DA7367739FB9AFC53FA4DFC42" box="[752,812,905,928]" pageId="2" pageNumber="79" rank="species">sp. n.</taxonomicNameLabel>
Holotype. Scale bar = 1mm.
</paragraph>
</caption>
<caption id="BED0A2ECA7367739FA39FC2DFA01FBAD" pageId="2" pageNumber="79" startId="2.[851,897,1015,1039]" targetType="figure">
<paragraph id="EA10F264A7367739FA39FC2DFA01FBAD" blockId="2.[851,1361,1015,1103]" pageId="2" pageNumber="79">
Figs 4-5. Microreticulation of elytra. 4
<taxonomicName id="2DAF89E7A7367739FC69FC22FD1CFBCD" authority="Schmidt-Goebel, 1846" authorityName="Schmidt-Goebel" authorityYear="1846" class="Insecta" family="Carabidae" genus="Dromoceryx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="D8DB2E76A7367739FC69FC22FAE4FBCD" italics="true" pageId="2" pageNumber="79">D. dorsalis</emphasis>
Schmidt-Goebel, 1846
</taxonomicName>
; 5
<taxonomicName id="2DAF89E7A7367739FDC7FBC2FC4CFBCD" authorityName="Anichtchenko &amp; Medina" authorityYear="2021" box="[1197,1325,1047,1071]" class="Insecta" family="Carabidae" genus="Dromoceryx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="obscurus" status="sp. n">
<emphasis id="D8DB2E76A7367739FDC7FBC2FC4CFBCD" box="[1197,1325,1047,1071]" italics="true" pageId="2" pageNumber="79">D. obscurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E8930DA7367739FC58FBC2FA01FBAD" pageId="2" pageNumber="79" rank="species">sp. n</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="EA10F264A7367739FA39FB05FD74FA55" blockId="2.[851,1361,1247,1463]" pageId="2" pageNumber="79">
<emphasis id="D8DB2E76A7367739FA39FB05FAF6FB15" bold="true" box="[851,919,1247,1271]" pageId="2" pageNumber="79">Head.</emphasis>
Mandibles short, with wide base; labrum wider than long, rectangular, with six setae along apical margin; mentum with rectangular tooth; clypeus with two lateral setae; eyes convex, with two pairs of supraorbital setae. Disc of head with isodiametric to slightly polygonal microrreticulation.
</paragraph>
<paragraph id="EA10F264A7367738FA39FA05F8E1FEB5" blockId="2.[851,1362,1503,1719]" lastBlockId="3.[227,737,191,343]" lastPageId="3" lastPageNumber="80" pageId="2" pageNumber="79">
<emphasis id="D8DB2E76A7367739FA39FA05FAAAFA15" bold="true" box="[851,971,1503,1527]" pageId="2" pageNumber="79">Pronotum.</emphasis>
Anterior margin widely interrupted, lateral margins narrow, basal rebording complete; anterior transverse impression absent; posterior transverse im-pression very shallow; median longitudinal impression moderately shallow; disc mod-erately flat, basal angles obtuse. Sides broadly rounded. Disc of pronotum irregularly reticulated, with slightly transverse meshes medially to strongly transverse laterally. Pronotum with two setae along each lateral sides, one in hind angle and one in anterior half just before maximal width.
</paragraph>
<caption id="BED0A2ECA7367739F845F94AF880F90A" pageId="2" pageNumber="79" startId="2.[303,342,1680,1704]" targetType="figure">
<paragraph id="EA10F264A7367739F845F94AF880F90A" blockId="2.[303,814,1680,1768]" pageId="2" pageNumber="79">
Fig. 2. Habitus of
<taxonomicName id="2DAF89E7A7367739F882F94BF8DBF92A" authority="SchmidtGoebel, 1846" authorityName="Schmidt-Goebel" authorityYear="1846" class="Insecta" family="Carabidae" genus="Dromoceryx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="D8DB2E76A7367739F882F94BFBA6F94A" box="[488,711,1680,1704]" italics="true" pageId="2" pageNumber="79">Dromoceryx dorsalis</emphasis>
SchmidtGoebel, 1846
</taxonomicName>
from Borikhan province, Malaysia. Scale bar = 1mm.
</paragraph>
</caption>
<paragraph id="EA10F264A7377738F989FEA5F83CFDD5" blockId="3.[227,737,383,567]" pageId="3" pageNumber="80">
<emphasis id="D8DB2E76A7377738F989FEA5F855FE75" bold="true" box="[227,308,383,407]" pageId="3" pageNumber="80">Elytra.</emphasis>
Intervals moderately flat, striae deep, minutely punctate; elytral apices truncate. Microreticulation faint, consist of strongly transverse sculpticells. Elytra with two setae in interval 3, one seta just before mid-length, one seta in apical 1/3
<superScript id="1DDA5F2CA7377738F820FDC4F837FDCE" attach="none" box="[330,342,542,556]" fontSize="6" pageId="3" pageNumber="80">rd</superScript>
.
</paragraph>
<paragraph id="EA10F264A7377738F989FD85FB6DFD95" blockId="3.[227,524,607,631]" box="[227,524,607,631]" pageId="3" pageNumber="80">
<emphasis id="D8DB2E76A7377738F989FD85F80BFD95" bold="true" box="[227,362,607,631]" pageId="3" pageNumber="80">Hind wings.</emphasis>
Macropterous.
</paragraph>
<paragraph id="EA10F264A7377738F989FD45F826FCF5" blockId="3.[227,737,671,791]" pageId="3" pageNumber="80">
<emphasis id="D8DB2E76A7377738F989FD45F841FD55" bold="true" box="[227,288,671,695]" pageId="3" pageNumber="80">Legs.</emphasis>
Claws dentate with 5 teeth of about equal length at either side; 1st-3st tarsomeres of male protarsus dilated and with adhesive pads ventrally.
</paragraph>
<paragraph id="EA10F264A7377738F989FCE5FB4DFC35" blockId="3.[227,738,831,984]" pageId="3" pageNumber="80">
<emphasis id="D8DB2E76A7377738F989FCE5F8E9FCB5" bold="true" box="[227,392,831,855]" pageId="3" pageNumber="80">Male genitalia.</emphasis>
Length 0.95-1.00 mm. Aedeagus elongate, ventral side straight, apex relatively wide and truncated; endophallus with several microtrichial patches and ventrally with two groups of small spines (
<figureCitation id="7294EEE1A7377738F8B4FC65FB7FFC35" box="[478,542,959,983]" captionStart="Fig" captionStartId="2.[851,891,547,571]" captionText="Fig. 3. Aedeagus of Dromoceryx obscurus sp. n. Scale bar = 0.5 mm." pageId="3" pageNumber="80">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="EA10F264A7377738F989FC25FB72FBF5" blockId="3.[227,531,1023,1047]" box="[227,531,1023,1047]" pageId="3" pageNumber="80">
<emphasis id="D8DB2E76A7377738F989FC25F8C1FBF5" bold="true" box="[227,416,1023,1047]" pageId="3" pageNumber="80">Female genitalia.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>