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<mods:title id="16D0499B03678094CAA4994567EBDD9A">Tortonian teleost otoliths from northern Italy: taxonomic synthesis and stratigraphic significance</mods:title>
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<treatment id="615996129474CE257A29A205FE4092DE" ID-DOI="http://doi.org/10.5281/zenodo.3848343" ID-GBIF-Taxon="164159965" ID-Zenodo-Dep="3848343" LSID="urn:lsid:plazi:treatment:615996129474CE257A29A205FE4092DE" httpUri="http://treatment.plazi.org/id/615996129474CE257A29A205FE4092DE" lastPageId="19" lastPageNumber="19" pageId="18" pageNumber="19">
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Genus
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<emphasis id="DB84FB169474CE247AF7A206FCFF953D" bold="true" box="[696,835,1617,1644]" italics="true" pageId="18" pageNumber="19">Lobianchia</emphasis>
Gatti, 1904
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The otoliths of
<taxonomicName id="2EF05C879474CE24793AA2C4FC9095FC" authority="(Cocco, 1838)" baseAuthorityName="Cocco" baseAuthorityYear="1838" box="[373,812,1683,1709]" class="Actinopterygii" family="Myctophidae" genus="Lobianchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="gemellarii">
<emphasis id="DB84FB169474CE24793AA2C4FDC095FC" box="[373,636,1683,1709]" italics="true" pageId="18" pageNumber="19">Lobianchia gemellarii</emphasis>
(Cocco, 1838)
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show an apparent ontogenetic variability (
<figureCitation id="71CB3B819474CE247D67A2C4FACD95FC" box="[1320,1393,1683,1709]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[305,1303,685,1840]" captionTargetId="figure@19.[285,1303,680,1845]" captionTargetPageId="19" captionText="Fig. 6. Recent otoliths of Lobianchia gemellarii (Cocco, 1838). A, C, E–F, I. Strait of Messina, fish total length (TL) 9.0 cm, 9.0 cm, 8.1 cm, 7.0 cm and 7.0 cm, respectively. B, D, G–H, J. Off Canaries, TL 9.0 cm, 9.0 cm, 4.8 cm, 9.0 cm and 4.8 cm, respectively. 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829317" httpUri="https://zenodo.org/record/3829317/files/figure.png" pageId="18" pageNumber="19">Fig. 6</figureCitation>
, Recent specimens). The small specimens of
<taxonomicName id="2EF05C879474CE247A83A2E0FCD99580" baseAuthorityName="Cocco" baseAuthorityYear="1838" box="[716,869,1719,1745]" class="Actinopterygii" family="Myctophidae" genus="Lobianchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="gemellarii">
<emphasis id="DB84FB169474CE247A83A2E0FD609581" box="[716,732,1719,1744]" italics="true" pageId="18" pageNumber="19">L</emphasis>
.
<emphasis id="DB84FB169474CE247AA3A2E0FCD99580" box="[748,869,1719,1745]" italics="true" pageId="18" pageNumber="19">gemellarii</emphasis>
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are rather similar to those of the same-sized
<taxonomicName id="2EF05C879474CE2478F2A28DFD3195A5" authority="(Zugmayer, 1911)" baseAuthorityName="Zugmayer" baseAuthorityYear="1911" box="[189,653,1754,1780]" class="Actinopterygii" family="Myctophidae" genus="Lobianchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="dofleini">
<emphasis id="DB84FB169474CE2478F2A28DFE1B95A5" box="[189,423,1754,1780]" italics="true" pageId="18" pageNumber="19">Lobianchia dofleini</emphasis>
(Zugmayer, 1911)
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, from which they differ in having a narrower sulcus and an obliquely inclining antero-dorsal rim that runs downwards to the antirostrum (
<figureCitation id="71CB3B819474CE247C1EA2AAFB0F9449" box="[1105,1203,1789,1816]" captionStart="Fig" captionStartId="17.[189,231,1514,1540]" captionTargetBox="[298,1290,265,1505]" captionTargetId="figure@17.[298,1290,265,1505]" captionTargetPageId="17" captionText="Fig. 5. Tortonian fish otoliths from northern Italy. A. Diaphus holti Tåning, 1918, Mondovi, Madonna della Neve (IRSNB P 9732). B. Diaphus metopoclampoides Steurbaut, 1983, Sant’Alosio (IRSNB P 9733). C–E. Diaphus regani Tåning, 1932, Montegibbio (IRSNB P 9734–P 9736). F–G. Diaphus pedemontanus (Robba, 1970), Sant’Alosio (IRSNB P 9737–P 9738). H. Diaphus aff. rafinesquii (Cocco, 1838), Mondovi, Madonna della Neve (IRSNB P 9739). I. Diaphus aff. splendidus (Brauer, 1904), Costa Vescovato (IRSNB P 9740). J. Diaphus cahuzaci Steurbaut, 1979, Mondovi, Madonna della Neve (IRSNB P 9741). K. Lampadena aff. dea Fraser-Brunner, 1949, Torrente Stirone (IRSNB P 9742). L. Lampadena aff. speculigeroides Brzobohatý &amp; Nolf, 1996, Stazzano (IRSNB P 9743). M. Hygophum hygomii (Lütken, 1892), Mondovi, Madonna della Neve (IRSNB P 9744). N–O. Lampadena gracilis (Schubert, 1912); N. Alba, Tanaro (50 m), O. Mondovi, Madonna della Neve (IRSNB P 9745–P 9746). P–Q. Lobianchia gemellarii (Cocco, 1838), Sant’Alosio (IRSNB P 9747–P 9748). R. Hygophum derthonensis (Anfossi &amp; Mosna, 1969), Torrente Stirone (IRSNB P 9749). S–T. Lampanyctus latesulcatus Nolf &amp; Steurbaut, 1983, Sant’Agata Fossili (IRSNB P 9750–P 9751). U. Merluccius cf. merluccius (Linnaeus, 1758), Sant’Agata Fossili (IRSNB P 9752). V. Lobianchia dofleini (Zugmayer, 1911), Costa Vescovato (IRSNB P 9753). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829315" httpUri="https://zenodo.org/record/3829315/files/figure.png" pageId="18" pageNumber="19">Figs 5Q</figureCitation>
,
<figureCitation id="71CB3B819474CE247CF0A2AAFB729446" box="[1215,1230,1789,1815]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[305,1303,685,1840]" captionTargetId="figure@19.[285,1303,680,1845]" captionTargetPageId="19" captionText="Fig. 6. Recent otoliths of Lobianchia gemellarii (Cocco, 1838). A, C, E–F, I. Strait of Messina, fish total length (TL) 9.0 cm, 9.0 cm, 8.1 cm, 7.0 cm and 7.0 cm, respectively. B, D, G–H, J. Off Canaries, TL 9.0 cm, 9.0 cm, 4.8 cm, 9.0 cm and 4.8 cm, respectively. 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829317" httpUri="https://zenodo.org/record/3829317/files/figure.png" pageId="18" pageNumber="19">6</figureCitation>
vs
<bibRefCitation id="8D615AF59474CE247CB9A2AAFACF9446" author="Nolf D." box="[1270,1395,1789,1815]" pageId="18" pageNumber="19" refId="ref22613" refString="Nolf D. 2013. The Diversity of Fish Otoliths, Past and Present. Royal Belgian Institute of Natural Sciences, Brussels." type="book" year="2013">
Nolf
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: pl. 81). In large otoliths of
<taxonomicName id="2EF05C879474CE247A47A376FD1E946B" baseAuthorityName="Cocco" baseAuthorityYear="1838" box="[520,674,1824,1850]" class="Actinopterygii" family="Myctophidae" genus="Lobianchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="gemellarii">
<emphasis id="DB84FB169474CE247A47A376FD1E946B" box="[520,674,1824,1850]" italics="true" pageId="18" pageNumber="19">L. gemellarii</emphasis>
</taxonomicName>
, the posterior rim becomes straight and the depression after the postero-dorsal angle is more pronounced (
<figureCitation id="71CB3B819474CE247AACA313FC83940F" box="[739,831,1859,1886]" captionStart="Fig" captionStartId="17.[189,231,1514,1540]" captionTargetBox="[298,1290,265,1505]" captionTargetId="figure@17.[298,1290,265,1505]" captionTargetPageId="17" captionText="Fig. 5. Tortonian fish otoliths from northern Italy. A. Diaphus holti Tåning, 1918, Mondovi, Madonna della Neve (IRSNB P 9732). B. Diaphus metopoclampoides Steurbaut, 1983, Sant’Alosio (IRSNB P 9733). C–E. Diaphus regani Tåning, 1932, Montegibbio (IRSNB P 9734–P 9736). F–G. Diaphus pedemontanus (Robba, 1970), Sant’Alosio (IRSNB P 9737–P 9738). H. Diaphus aff. rafinesquii (Cocco, 1838), Mondovi, Madonna della Neve (IRSNB P 9739). I. Diaphus aff. splendidus (Brauer, 1904), Costa Vescovato (IRSNB P 9740). J. Diaphus cahuzaci Steurbaut, 1979, Mondovi, Madonna della Neve (IRSNB P 9741). K. Lampadena aff. dea Fraser-Brunner, 1949, Torrente Stirone (IRSNB P 9742). L. Lampadena aff. speculigeroides Brzobohatý &amp; Nolf, 1996, Stazzano (IRSNB P 9743). M. Hygophum hygomii (Lütken, 1892), Mondovi, Madonna della Neve (IRSNB P 9744). N–O. Lampadena gracilis (Schubert, 1912); N. Alba, Tanaro (50 m), O. Mondovi, Madonna della Neve (IRSNB P 9745–P 9746). P–Q. Lobianchia gemellarii (Cocco, 1838), Sant’Alosio (IRSNB P 9747–P 9748). R. Hygophum derthonensis (Anfossi &amp; Mosna, 1969), Torrente Stirone (IRSNB P 9749). S–T. Lampanyctus latesulcatus Nolf &amp; Steurbaut, 1983, Sant’Agata Fossili (IRSNB P 9750–P 9751). U. Merluccius cf. merluccius (Linnaeus, 1758), Sant’Agata Fossili (IRSNB P 9752). V. Lobianchia dofleini (Zugmayer, 1911), Costa Vescovato (IRSNB P 9753). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829315" httpUri="https://zenodo.org/record/3829315/files/figure.png" pageId="18" pageNumber="19">Figs 5P</figureCitation>
,
<figureCitation id="71CB3B819474CE247B02A314FCE2940F" box="[845,862,1859,1886]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[305,1303,685,1840]" captionTargetId="figure@19.[285,1303,680,1845]" captionTargetPageId="19" captionText="Fig. 6. Recent otoliths of Lobianchia gemellarii (Cocco, 1838). A, C, E–F, I. Strait of Messina, fish total length (TL) 9.0 cm, 9.0 cm, 8.1 cm, 7.0 cm and 7.0 cm, respectively. B, D, G–H, J. Off Canaries, TL 9.0 cm, 9.0 cm, 4.8 cm, 9.0 cm and 4.8 cm, respectively. 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829317" httpUri="https://zenodo.org/record/3829317/files/figure.png" pageId="18" pageNumber="19">6</figureCitation>
). Otoliths of such a dimension (length about
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) are derived from fish attaining a length of about
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(IRSNB collection), which is very huge compared to the average individual’s maximum length of
<quantity id="2E088AE19474CE247B12A3DDFC2A94F4" box="[861,918,1930,1957]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="cm" value="6.0">6 cm</quantity>
(
<bibRefCitation id="8D615AF59474CE247BEBA3DDFB1194F5" author="Nafpaktitis B. G. &amp; Backus R. H. &amp; Craddock J. E. &amp; Haedrich R. L. &amp; Robinson B. H. &amp; Karnella C." box="[932,1197,1930,1957]" pageId="18" pageNumber="19" pagination="1 - 299" refId="ref22438" refString="Nafpaktitis B. G., Backus R. H., Craddock J. E., Haedrich R. L., Robinson B. H. &amp; Karnella C. 1977. Family Myctophidae. In: Gibbs R. H. Jr. (ed.) Fishes of the Western North Atlantic: 1 - 299. Memoir of the Sears Foundation for Marine Research, New Haven 7." type="journal article" year="1977">
Nafpaktitis
<emphasis id="DB84FB169474CE247C60A3DCFBDD94F5" box="[1071,1121,1930,1956]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
.
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;
<bibRefCitation id="8D615AF59474CE247CF8A3DDFF469499" author="Whitehead P. J. P. &amp; Bauchot M. L. &amp; Hureau J. C. &amp; Nielsen J. &amp; Tortonese E." pageId="18" pageNumber="19" refId="ref24088" refString="Whitehead P. J. P., Bauchot M. L., Hureau J. C., Nielsen J. &amp; Tortonese E. (eds). 1986 - 1989. Fishes of the North-Eastern Atlantic and the Mediterranean, 3 volumes. UNESCO, Paris." type="book" year="1986">
Whitehead
<emphasis id="DB84FB169474CE247D72A3DCFAD394F5" box="[1341,1391,1930,1956]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
.
<date id="9D4E01C49474CE2478F2A3FAFF469499" box="[189,250,1965,1992]" pageId="18" pageNumber="19" value="1986" valueMax="1989">1986</date>
</bibRefCitation>
–1989). In the slope water off New
<collectingCountry id="91E767949474CE247ADBA3FAFD459496" box="[660,761,1965,1991]" name="United Kingdom" pageId="18" pageNumber="19">England</collectingCountry>
,
<bibRefCitation id="8D615AF59474CE247B4BA3FAFBA09499" author="Nafpaktitis B. G. &amp; Backus R. H. &amp; Craddock J. E. &amp; Haedrich R. L. &amp; Robinson B. H. &amp; Karnella C." box="[772,1052,1965,1992]" pageId="18" pageNumber="19" pagination="1 - 299" refId="ref22438" refString="Nafpaktitis B. G., Backus R. H., Craddock J. E., Haedrich R. L., Robinson B. H. &amp; Karnella C. 1977. Family Myctophidae. In: Gibbs R. H. Jr. (ed.) Fishes of the Western North Atlantic: 1 - 299. Memoir of the Sears Foundation for Marine Research, New Haven 7." type="journal article" year="1977">
Nafpaktitis
<emphasis id="DB84FB169474CE247BC2A3F9FC039496" box="[909,959,1965,1991]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (
<date id="9D4E01C49474CE247B99A3F9FBAF9499" box="[982,1043,1966,1992]" pageId="18" pageNumber="19" value="1977">1977</date>
)
</bibRefCitation>
noted a number of specimens of
<taxonomicName id="2EF05C879474CE247894A385FECD94BA" baseAuthorityName="Cocco" baseAuthorityYear="1838" box="[219,369,2001,2027]" class="Actinopterygii" family="Myctophidae" genus="Lobianchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="gemellarii">
<emphasis id="DB84FB169474CE247894A385FF5794BA" box="[219,235,2002,2027]" italics="true" pageId="18" pageNumber="19">L</emphasis>
.
<emphasis id="DB84FB169474CE2478B7A386FECD94BA" box="[248,369,2001,2027]" italics="true" pageId="18" pageNumber="19">gemellarii</emphasis>
</taxonomicName>
ranging in length between 85 and
<quantity id="2E088AE19474CE247AB3A386FCE294BA" box="[764,862,2001,2027]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="mm" value="100.0">100 mm</quantity>
. Such large individuals are occasionally found outside the spawning area. Studies on comparative morphology of Recent
<taxonomicName id="2EF05C879475CE257C04A55CFB6D9274" authorityName="Gatti" authorityYear="1904" box="[1099,1233,267,293]" class="Actinopterygii" family="Myctophidae" genus="Lobianchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="DB84FB169475CE257C04A55CFB6D9274" box="[1099,1233,267,293]" italics="true" pageId="19" pageNumber="20">Lobianchia</emphasis>
</taxonomicName>
otoliths have obtained similar results (
<bibRefCitation id="8D615AF59475CE257992A579FD719218" author="Schwarzhans W." box="[477,717,302,329]" pageId="19" pageNumber="20" pagination="41 - 82" refId="ref23659" refString="Schwarzhans W. 2013 a. A comparative morphological study of the Recent otoliths of the genera Diaphus, Idiolychnus and Lobianchia (Myctophidae). Palaeo Ichthyologica 13: 41 - 82. https: // doi. org / 10.13140 / 2.1.2872.3843" type="journal article" year="2013">
Schwarzhans
<date id="9D4E01C49475CE257ACEA578FD7C9218" box="[641,704,303,329]" pageId="19" pageNumber="20" value="2013">2013</date>
a
</bibRefCitation>
: pl. 15, figs 6, 7). Based on these observations, the studied material testifies to the occurrence of both species,
<taxonomicName id="2EF05C879475CE257B53A504FC2F923D" baseAuthorityName="Zugmayer" baseAuthorityYear="1911" box="[796,915,338,364]" class="Actinopterygii" family="Myctophidae" genus="Lobianchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="dofleini">
<emphasis id="DB84FB169475CE257B53A504FC90923D" box="[796,812,339,364]" italics="true" pageId="19" pageNumber="20">L</emphasis>
.
<emphasis id="DB84FB169475CE257B74A505FC2F923D" box="[827,915,338,364]" italics="true" pageId="19" pageNumber="20">dofleini</emphasis>
</taxonomicName>
(
<figureCitation id="71CB3B819475CE257BEBA505FBBD923C" box="[932,1025,338,365]" captionStart="Fig" captionStartId="17.[189,231,1514,1540]" captionTargetBox="[298,1290,265,1505]" captionTargetId="figure@17.[298,1290,265,1505]" captionTargetPageId="17" captionText="Fig. 5. Tortonian fish otoliths from northern Italy. A. Diaphus holti Tåning, 1918, Mondovi, Madonna della Neve (IRSNB P 9732). B. Diaphus metopoclampoides Steurbaut, 1983, Sant’Alosio (IRSNB P 9733). C–E. Diaphus regani Tåning, 1932, Montegibbio (IRSNB P 9734–P 9736). F–G. Diaphus pedemontanus (Robba, 1970), Sant’Alosio (IRSNB P 9737–P 9738). H. Diaphus aff. rafinesquii (Cocco, 1838), Mondovi, Madonna della Neve (IRSNB P 9739). I. Diaphus aff. splendidus (Brauer, 1904), Costa Vescovato (IRSNB P 9740). J. Diaphus cahuzaci Steurbaut, 1979, Mondovi, Madonna della Neve (IRSNB P 9741). K. Lampadena aff. dea Fraser-Brunner, 1949, Torrente Stirone (IRSNB P 9742). L. Lampadena aff. speculigeroides Brzobohatý &amp; Nolf, 1996, Stazzano (IRSNB P 9743). M. Hygophum hygomii (Lütken, 1892), Mondovi, Madonna della Neve (IRSNB P 9744). N–O. Lampadena gracilis (Schubert, 1912); N. Alba, Tanaro (50 m), O. Mondovi, Madonna della Neve (IRSNB P 9745–P 9746). P–Q. Lobianchia gemellarii (Cocco, 1838), Sant’Alosio (IRSNB P 9747–P 9748). R. Hygophum derthonensis (Anfossi &amp; Mosna, 1969), Torrente Stirone (IRSNB P 9749). S–T. Lampanyctus latesulcatus Nolf &amp; Steurbaut, 1983, Sant’Agata Fossili (IRSNB P 9750–P 9751). U. Merluccius cf. merluccius (Linnaeus, 1758), Sant’Agata Fossili (IRSNB P 9752). V. Lobianchia dofleini (Zugmayer, 1911), Costa Vescovato (IRSNB P 9753). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829315" httpUri="https://zenodo.org/record/3829315/files/figure.png" pageId="19" pageNumber="20">Fig. 5V</figureCitation>
) and
<taxonomicName id="2EF05C879475CE257C0BA504FB61923D" baseAuthorityName="Cocco" baseAuthorityYear="1838" box="[1092,1245,338,364]" class="Actinopterygii" family="Myctophidae" genus="Lobianchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="gemellarii">
<emphasis id="DB84FB169475CE257C0BA504FB61923D" box="[1092,1245,338,364]" italics="true" pageId="19" pageNumber="20">L. gemellarii</emphasis>
</taxonomicName>
(
<figureCitation id="71CB3B819475CE257CA2A505FADA923D" box="[1261,1382,338,365]" captionStart="Fig" captionStartId="17.[189,231,1514,1540]" captionTargetBox="[298,1290,265,1505]" captionTargetId="figure@17.[298,1290,265,1505]" captionTargetPageId="17" captionText="Fig. 5. Tortonian fish otoliths from northern Italy. A. Diaphus holti Tåning, 1918, Mondovi, Madonna della Neve (IRSNB P 9732). B. Diaphus metopoclampoides Steurbaut, 1983, Sant’Alosio (IRSNB P 9733). C–E. Diaphus regani Tåning, 1932, Montegibbio (IRSNB P 9734–P 9736). F–G. Diaphus pedemontanus (Robba, 1970), Sant’Alosio (IRSNB P 9737–P 9738). H. Diaphus aff. rafinesquii (Cocco, 1838), Mondovi, Madonna della Neve (IRSNB P 9739). I. Diaphus aff. splendidus (Brauer, 1904), Costa Vescovato (IRSNB P 9740). J. Diaphus cahuzaci Steurbaut, 1979, Mondovi, Madonna della Neve (IRSNB P 9741). K. Lampadena aff. dea Fraser-Brunner, 1949, Torrente Stirone (IRSNB P 9742). L. Lampadena aff. speculigeroides Brzobohatý &amp; Nolf, 1996, Stazzano (IRSNB P 9743). M. Hygophum hygomii (Lütken, 1892), Mondovi, Madonna della Neve (IRSNB P 9744). N–O. Lampadena gracilis (Schubert, 1912); N. Alba, Tanaro (50 m), O. Mondovi, Madonna della Neve (IRSNB P 9745–P 9746). P–Q. Lobianchia gemellarii (Cocco, 1838), Sant’Alosio (IRSNB P 9747–P 9748). R. Hygophum derthonensis (Anfossi &amp; Mosna, 1969), Torrente Stirone (IRSNB P 9749). S–T. Lampanyctus latesulcatus Nolf &amp; Steurbaut, 1983, Sant’Agata Fossili (IRSNB P 9750–P 9751). U. Merluccius cf. merluccius (Linnaeus, 1758), Sant’Agata Fossili (IRSNB P 9752). V. Lobianchia dofleini (Zugmayer, 1911), Costa Vescovato (IRSNB P 9753). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829315" httpUri="https://zenodo.org/record/3829315/files/figure.png" pageId="19" pageNumber="20">Fig. 5P, Q</figureCitation>
), in the Tortonian sediments.
</paragraph>
</subSubSection>
</treatment>
</document>
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