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<mods:title id="4FD7F02FD9003DDCFDE49BAC448104D8">New nomenclatural and taxonomic adjustments in Dracaena (Asparagaceae)</mods:title>
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<mods:namePart id="80AA74E7D2097F9F576F3DABB2915E29">Takawira-Nyenya, Ratidzayi</mods:namePart>
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<mods:namePart id="3BD620E7ED7D2003BABDF2F61EC9E888">Thiede, Joachim</mods:namePart>
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<mods:affiliation id="E44A46820B19F00D01E57153C20FD51C">Mechelnbusch 2, 22559 Hamburg, Germany. &amp; joachim _ thiede @ gmx. de; https: // orcid. org / 0000 - 0002 - 0130 - 8424</mods:affiliation>
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<mods:namePart id="7386F2A22DE9CF61C976EDECEB4D80DB">Mucina, Ladislav</mods:namePart>
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<emphasis id="DBECA909FF89FFBDFF1BED42FEDAFCBF" bold="true" box="[136,321,842,869]" italics="true" pageId="4" pageNumber="297">Dracaena ballyi</emphasis>
(
<bibRefCitation id="8D0908EAFF89FFBDFEDCED43FD89FCBE" author="Newton, L. E." box="[335,530,842,868]" pageId="4" pageNumber="297" pagination="10 - 13" refId="ref6187" refString="Newton, L. E. (2004) Sansevieria ballyi, a name at last for a long-cultivated plant. British Cactus and Succulent Journal 22: 10 - 13. [https: // www. jstor. org / stable / 42793926]" type="journal article" year="2004">Newton 2004: 11</bibRefCitation>
)
<bibRefCitation id="8D0908EAFF89FFBDFDB1ED42FC0AFCBF" author="Byng, J. W. &amp; Christenhusz, J. M. M." box="[546,913,842,869]" pageId="4" pageNumber="297" pagination="64 - 67" refId="ref5266" refString="Byng, J. W. &amp; Christenhusz, J. M. M. (2018) Asparagaceae. In: Christenhusz, J. M. M., Fay, F. M. &amp; Byng, J. W. (Eds.) The Global Flora. A practical flora to vascular plant species of the world. Special Edition. GLOVAP Nomenclature 1, vol. 4. Plant Gateway Ltd., Bradford, pp. 64 - 67." type="book chapter" year="2018">Byng &amp; Christenhusz (2018: 65)</bibRefCitation>
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:—
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[Coast Province,
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]:
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, soil pockets on rock faces, amongst sedges, and in deeper soil in the shelter of taller vegetation, elev.
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,
<emphasis id="DBECA909FF89FFBDFD0AED9EFC8CFC71" box="[665,791,918,940]" italics="true" pageId="4" pageNumber="297">Newton 5594</emphasis>
(
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<specimenCode id="B93EDD60FF89FFBDFCEFED9FFC66FC76" box="[892,1021,918,940]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="297" type="Herbarium">K 000099514</specimenCode>
digital image!, available at: http://specimens. kew.org/herbarium/
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; isotype
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digital image!, available at: https://plants.jstor.org/stable/10.5555/al.ap. specimen.ea000002620)
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.
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<paragraph id="E927751BFF89FFBDFF1BEA08FDB5FBC2" blockId="4.[136,1453,881,1156]" box="[136,558,1025,1048]" pageId="4" pageNumber="297">
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<taxonomicName id="2E980E98FF89FFBDFF05EA0BFDB2FBC2" ID-CoL="79J52" authority="L. E. Newton (2004: 11)" authorityName="L. E. Newton" authorityPageNumber="11" authorityYear="2004" box="[150,553,1026,1048]" class="Liliopsida" family="Asparagaceae" genus="Sansevieria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="ballyi">
<emphasis id="DBECA909FF89FFBDFF05EA0BFED8FBCD" box="[150,323,1026,1047]" italics="true" pageId="4" pageNumber="297">Sansevieria ballyi</emphasis>
L.E.
<bibRefCitation id="8D0908EAFF89FFBDFEFEEA0BFDB2FBC2" author="Newton, L. E." box="[365,553,1026,1048]" pageId="4" pageNumber="297" pagination="10 - 13" refId="ref6187" refString="Newton, L. E. (2004) Sansevieria ballyi, a name at last for a long-cultivated plant. British Cactus and Succulent Journal 22: 10 - 13. [https: // www. jstor. org / stable / 42793926]" type="journal article" year="2004">Newton (2004: 11)</bibRefCitation>
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.
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<paragraph id="E927751BFF89FFBDFF1BEA2FFD78FB85" blockId="4.[136,1453,881,1156]" pageId="4" pageNumber="297">
=
<taxonomicName id="2E980E98FF89FFBDFF05EA2FFD55FBE6" ID-CoL="8XFZ3" authority="L. E. Newton (2015: 11)" authorityName="L. E. Newton" authorityPageNumber="11" authorityYear="2015" box="[150,718,1062,1084]" class="Liliopsida" family="Asparagaceae" genus="Sansevieria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="variety" species="ballyi" variety="robertsoniae">
<emphasis id="DBECA909FF89FFBDFF05EA2FFED9FBE1" box="[150,322,1062,1083]" italics="true" pageId="4" pageNumber="297">Sansevieria ballyi</emphasis>
<materialsCitation id="59F07F46FF89FFBDFED4EA2FFD55FBE6" ID-GBIF-Occurrence="3400173309" box="[327,718,1062,1084]" collectionCode="EA" collectorName="Stedje &amp; Takawira-Nyenya" country="Kenya" location="Coast Province" pageId="4" pageNumber="297" specimenCount="1" typeStatus="holotype">
var.
<emphasis id="DBECA909FF89FFBDFEFCEA2FFE72FBE1" box="[367,489,1062,1083]" italics="true" pageId="4" pageNumber="297">robertsoniae</emphasis>
L.E.Newton (2015: 11)
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. Type:—
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.
<location id="EC4723C0FF89FFBDFC11EA2FFB8DFBE6" LSID="urn:lsid:plazi:treatment:6131C40DFF89FFBDFF1BED42FCC1F8EB:EC4723C0FF89FFBDFC11EA2FFB8DFBE6" box="[898,1046,1062,1084]" country="Kenya" name="Coast Province" pageId="4" pageNumber="297">Coast Province</location>
, Chakama Road, s. d.,
<emphasis id="DBECA909FF89FFBDFB7DEA2EFE6CFB85" italics="true" pageId="4" pageNumber="297">
Newton, Robertson,
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&amp;
<collectorName id="446D10CDFF89FFBDFE84EA43FE5AFB85" box="[279,449,1098,1119]" pageId="4" pageNumber="297">Takawira-Nyenya</collectorName>
5981
</emphasis>
(
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EA)
<emphasis id="DBECA909FF89FFBDFD1FEA43FD78FB85" bold="true" box="[652,739,1098,1119]" italics="true" pageId="4" pageNumber="297">syn. nov.</emphasis>
</paragraph>
<paragraph id="E927751BFF89FFBDFF1BEA67FCF6FB59" blockId="4.[136,1453,881,1156]" box="[136,877,1134,1156]" pageId="4" pageNumber="297">
<treatmentCitationGroup id="C9885235FF89FFBDFF1BEA67FCF6FB59" box="[136,877,1134,1156]" pageId="4" pageNumber="297">
=
<taxonomicName id="2E980E98FF89FFBDFF05EA67FD4DFB5E" authority="Pfennig in Heitz &amp; Zeller" box="[150,726,1134,1156]" class="Liliopsida" family="Asparagaceae" genus="Sansevieria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="tsavoensis">
<emphasis id="DBECA909FF89FFBDFF05EA67FEF4FB59" box="[150,367,1134,1155]" italics="true" pageId="4" pageNumber="297">Sansevieria tsavoensis</emphasis>
Pfennig in
<treatmentCitation id="6839530AFF89FFBDFE4DEA67FD4DFB5E" author="Heitz, B. &amp; Zeller, J." box="[478,726,1134,1156]" page="21" pageId="4" pageNumber="297" year="1990">
<bibRefCitation id="8D0908EAFF89FFBDFE4DEA67FD4DFB5E" author="Heitz, B. &amp; Zeller, J." box="[478,726,1134,1156]" pageId="4" pageNumber="297" pagination="12 - 24" refId="ref5627" refString="Heitz, B. &amp; Zeller, J. (1990) Comment s'appelle cette sanseviere? Succulentes 13: 12 - 24." type="journal article" year="1990">Heitz &amp; Zeller (1990: 21)</bibRefCitation>
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</taxonomicName>
,
<emphasis id="DBECA909FF89FFBDFD73EA66FCF6FB59" box="[736,877,1134,1156]" italics="true" pageId="4" pageNumber="297">nomen nudum.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E927751BFF89FFBDFF1BEABBFDE6F91E" blockId="4.[136,1452,1202,1841]" pageId="4" pageNumber="297">
<emphasis id="DBECA909FF89FFBDFF1BEABBFF5BFB16" bold="true" box="[136,192,1202,1228]" pageId="4" pageNumber="297">Note</emphasis>
:—German botanist Horst Pfennig intended to publish this taxon as
<taxonomicName id="2E980E98FF89FFBDFC3FEABAFBA1FB16" authority="Pfennig in Heitz &amp; Zeller" box="[940,1082,1203,1229]" class="Liliopsida" family="Asparagaceae" genus="Sansevieria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="tsavoensis">
<emphasis id="DBECA909FF89FFBDFC3FEABAFBA1FB16" box="[940,1082,1203,1229]" italics="true" pageId="4" pageNumber="297">S. tsavoensis</emphasis>
</taxonomicName>
, but this name remained a
<emphasis id="DBECA909FF89FFBDFAF2EABAFF4FFB2A" italics="true" pageId="4" pageNumber="297">nomen nudum</emphasis>
according to ICN Article 38.1 (
<bibRefCitation id="8D0908EAFF89FFBDFDD4EADFFCB7FB2A" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[583,812,1238,1264]" pageId="4" pageNumber="297" refId="ref6937" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashutten. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="DBECA909FF89FFBDFD39EADEFD7EFB2A" box="[682,741,1238,1264]" italics="true" pageId="4" pageNumber="297">et al.</emphasis>
2018
</bibRefCitation>
). The taxon was later published as
<taxonomicName id="2E980E98FF89FFBDFB56EADEFABFFB2A" ID-CoL="79J52" authorityName="L. E. Newton" authorityYear="2004" box="[1221,1316,1238,1265]" class="Liliopsida" family="Asparagaceae" genus="Sansevieria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="ballyi">
<emphasis id="DBECA909FF89FFBDFB56EADEFABFFB2A" box="[1221,1316,1238,1265]" italics="true" pageId="4" pageNumber="297">S. ballyi</emphasis>
</taxonomicName>
by
<bibRefCitation id="8D0908EAFF89FFBDFAC0EADEFF48FACE" author="Newton, L. E." pageId="4" pageNumber="297" pagination="10 - 13" refId="ref6187" refString="Newton, L. E. (2004) Sansevieria ballyi, a name at last for a long-cultivated plant. British Cactus and Succulent Journal 22: 10 - 13. [https: // www. jstor. org / stable / 42793926]" type="journal article" year="2004">Newton (2004)</bibRefCitation>
without knowledge of Pfennigs intended name, to accomodate dwarf, stoloniferous rosette plants with very short leaves with a distinct rough surface and distinct colour banding and short inflorescences, from south-eastern
<collectingCountry id="918F358BFF89FFBDFF1BEB4AFF4EFA87" box="[136,213,1347,1373]" name="Kenya" pageId="4" pageNumber="297">Kenya</collectingCountry>
. Later, studies by
<bibRefCitation id="8D0908EAFF89FFBDFE0DEB4AFDCDFA87" author="Newton, L. E." box="[414,598,1346,1373]" pageId="4" pageNumber="297" pagination="10 - 12" refId="ref6278" refString="Newton, L. E. (2010 b) Notes on Sansevieria ballyi, with description of a new cultivar. Sansevieria 22: 10 - 12." type="journal article" year="2010">Newton (2010b)</bibRefCitation>
and
<bibRefCitation id="8D0908EAFF89FFBDFD1FEB4BFCDAFA86" author="Mansfeld, P. A." box="[652,833,1346,1372]" pageId="4" pageNumber="297" pagination="225 - 229" refId="ref6030" refString="Mansfeld, P. A. (2016) Sansevieria ballyi L. E. Newton (Asparagaceae) and newly found field notes. Bradleya 34: 225 - 229. https: // doi. org / 10.25223 / brad. n 34.2016. a 10" type="journal article" year="2016">Mansfeld (2016</bibRefCitation>
,
<bibRefCitation id="8D0908EAFF89FFBDFCDFEB4BFC1DFA86" author="Mansfeld, P. A." box="[844,902,1346,1372]" pageId="4" pageNumber="297" pagination="24 - 35" refId="ref6078" refString="Mansfeld, P. A. (2017) Die beliebte Sansevieria ballyi und die wahre Geschichte ihrer Entdeckung. Sansevieria Online 5: 24 - 35." type="journal article" year="2017">2017</bibRefCitation>
) showed
<taxonomicName id="2E980E98FF89FFBDFC63EB4AFBD7FA86" ID-CoL="79J52" authorityName="L. E. Newton" authorityYear="2004" box="[1008,1100,1346,1373]" class="Liliopsida" family="Asparagaceae" genus="Sansevieria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="ballyi">
<emphasis id="DBECA909FF89FFBDFC63EB4AFBD7FA86" box="[1008,1100,1346,1373]" italics="true" pageId="4" pageNumber="297">S. ballyi</emphasis>
</taxonomicName>
to be common in south-eastern
<collectingCountry id="918F358BFF89FFBDFF1BEB6EFF4AFA5B" box="[136,209,1383,1409]" name="Kenya" pageId="4" pageNumber="297">Kenya</collectingCountry>
and to extend also into adjacent northeast
<collectingCountry id="918F358BFF89FFBDFD2EEB6FFCBEFA5A" box="[701,805,1382,1408]" name="Tanzania" pageId="4" pageNumber="297">Tanzania</collectingCountry>
. It exhibits a considerable variability in rosette size, leaf length, stolon length, and leaf colour ranging from pale green to almost white with transverse bands varying in width. A population with paler leaf colour and very smooth leaf surface at the northern edge of the species distribution range was first published as a cultivar;
<emphasis id="DBECA909FF89FFBDFE6CEBDAFD19FA37" box="[511,642,1491,1517]" italics="true" pageId="4" pageNumber="297">Sansevieria</emphasis>
Ann Robertson (
<bibRefCitation id="8D0908EAFF89FFBDFCC6EBDBFBB1FA36" author="Newton, L. E." box="[853,1066,1490,1516]" pageId="4" pageNumber="297" pagination="10 - 12" refId="ref6278" refString="Newton, L. E. (2010 b) Notes on Sansevieria ballyi, with description of a new cultivar. Sansevieria 22: 10 - 12." type="journal article" year="2010">Newton 2010b: 12</bibRefCitation>
) and its pale leaf colour was later concluded to be “normal variation within a species” (
<bibRefCitation id="8D0908EAFF89FFBDFD40EBFFFC3CF9CA" author="Newton, L. E." box="[723,935,1526,1552]" pageId="4" pageNumber="297" pagination="10 - 12" refId="ref6278" refString="Newton, L. E. (2010 b) Notes on Sansevieria ballyi, with description of a new cultivar. Sansevieria 22: 10 - 12." type="journal article" year="2010">Newton 2010b: 12</bibRefCitation>
). Later, the population was elevated to varietal rank as
<taxonomicName id="2E980E98FF89FFBDFF77E812FCC4F9EE" ID-CoL="8XFZ3" authority="(Newton 2015: 11)" baseAuthorityName="Newton" baseAuthorityPageNumber="11" baseAuthorityYear="2015" box="[228,863,1562,1589]" class="Liliopsida" family="Asparagaceae" genus="Sansevieria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="variety" species="ballyi" variety="robertsoniae">
<emphasis id="DBECA909FF89FFBDFF77E812FE2BF9EE" box="[228,432,1562,1589]" italics="true" pageId="4" pageNumber="297">Sansevieria ballyi</emphasis>
var.
<emphasis id="DBECA909FF89FFBDFE7FE813FDE1F9EE" box="[492,634,1562,1588]" italics="true" pageId="4" pageNumber="297">robertsoniae</emphasis>
(Newton 2015: 11)
</taxonomicName>
. Plants with short, smooth leaves had already been collected back in 1960 by W. Rauh, Heidelberg [D =
<collectingCountry id="918F358BFF89FFBDFD48E837FCFCF982" box="[731,871,1598,1624]" name="Germany" pageId="4" pageNumber="297">Deutschland</collectingCountry>
(
<collectingCountry id="918F358BFF89FFBDFCE5E837FC7FF983" box="[886,996,1598,1625]" name="Germany" pageId="4" pageNumber="297">Germany</collectingCountry>
)] together with typical plants with rough surface at Mutomo Hills. The smooth-leaved plants were later seen as natural variation by
<bibRefCitation id="8D0908EAFF89FFBDFB02E86BFA33F9A6" author="Mansfeld, P. A." box="[1169,1448,1634,1660]" pageId="4" pageNumber="297" pagination="24 - 35" refId="ref6078" refString="Mansfeld, P. A. (2017) Die beliebte Sansevieria ballyi und die wahre Geschichte ihrer Entdeckung. Sansevieria Online 5: 24 - 35." type="journal article" year="2017">Mansfeld (2017: 3334)</bibRefCitation>
. Consequently,
<taxonomicName id="2E980E98FF89FFBDFEA3E88EFDCEF97A" ID-CoL="8XFZ3" baseAuthorityName="Newton" baseAuthorityYear="2015" box="[304,597,1670,1697]" class="Liliopsida" family="Asparagaceae" genus="Sansevieria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="variety" species="ballyi" variety="robertsoniae">
<emphasis id="DBECA909FF89FFBDFEA3E88EFE16F97A" box="[304,397,1670,1697]" italics="true" pageId="4" pageNumber="297">S. ballyi</emphasis>
var.
<emphasis id="DBECA909FF89FFBDFE54E88FFDCEF97A" box="[455,597,1670,1696]" italics="true" pageId="4" pageNumber="297">robertsoniae</emphasis>
</taxonomicName>
falls within the range of variation encountered in
<taxonomicName id="2E980E98FF89FFBDFB02E88EFAEAF97A" ID-CoL="79J52" authorityName="L. E. Newton" authorityYear="2004" box="[1169,1393,1670,1697]" class="Liliopsida" family="Asparagaceae" genus="Sansevieria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="ballyi">
<emphasis id="DBECA909FF89FFBDFB02E88EFB74F97A" box="[1169,1263,1670,1697]" italics="true" pageId="4" pageNumber="297">S. ballyi</emphasis>
(
<emphasis id="DBECA909FF89FFBDFA93E88EFAEAF97A" box="[1280,1393,1670,1696]" italics="true" pageId="4" pageNumber="297">var. ballyi</emphasis>
</taxonomicName>
) and therefore it is referred to the synonymy here.
</paragraph>
</subSubSection>
<subSubSection id="A1822690FF89FFBDFF2EE8C7FCC1F8EB" pageId="4" pageNumber="297" type="description">
<paragraph id="E927751BFF89FFBDFF2EE8C7FCC1F8EB" blockId="4.[136,1452,1202,1841]" pageId="4" pageNumber="297">
<emphasis id="DBECA909FF89FFBDFF2EE8C7FED6F932" bold="true" box="[189,333,1742,1768]" pageId="4" pageNumber="297">Typification</emphasis>
:—
<materialsCitation id="59F07F46FF89FFBDFEFCE8C7FCCEF8EB" ID-GBIF-Occurrence="3400173306" collectionCode="JSTOR" pageId="4" pageNumber="297" specimenCode="K000099514" specimenCount="1" typeStatus="holotype">
The
<typeStatus id="3623CBB9FF89FFBDFE32E8C7FD98F932" box="[417,515,1742,1768]" pageId="4" pageNumber="297" type="holotype">holotype</typeStatus>
of
<taxonomicName id="2E980E98FF89FFBDFDB7E8C6FD70F932" authorityName="L. E. Newton" authorityYear="2004" box="[548,747,1742,1769]" class="Liliopsida" family="Asparagaceae" genus="Sansevieria" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="ballyi">
<emphasis id="DBECA909FF89FFBDFDB7E8C6FD70F932" box="[548,747,1742,1769]" italics="true" pageId="4" pageNumber="297">Sansevieria ballyi</emphasis>
</taxonomicName>
at Kew (=
<specimenCode id="B93EDD60FF89FFBDFCCDE8C7FC68F933" box="[862,1011,1742,1769]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="297" type="Herbarium">K000099514</specimenCode>
, see above) designated in the protologue (Newton 2015: 11) is erroneously labelled as “ISOTYPE” on the sheet and in the
<bibRefCitation id="8D0908EAFF89FFBDFBBAE8FAFA30F8D6" author="Kew Herbarium Catalogue" box="[1065,1451,1778,1805]" pageId="4" pageNumber="297" refId="ref5719" refString="Kew Herbarium Catalogue (2021) Sansevieria ballyi L. E. Newton. Specimen: K 00099514. Available at: http: // specimens. kew. org / herbarium / K 000099514 (accessed 10 May 2021)" type="url" year="2021">Kew Herbarium Catalogue (2021)</bibRefCitation>
and is wanting at
<collectionCode id="8F89EDDEFF89FFBDFEDDE91FFE3BF8EA" box="[334,416,1814,1840]" pageId="4" pageNumber="297">JSTOR</collectionCode>
Global Plants (https://plants.jstor.org/)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>