178 lines
14 KiB
XML
178 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.721.21529" ID-GBIF-Dataset="8e4ab8e9-98d9-417c-9188-ea0b1161c40b" ID-PMC="PMC5740412" ID-Pensoft-Pub="1313-2970-721-45" ID-PubMed="29308024" ID-ZBK="1300B5278A5041E38D5E1C745846BBDF" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-721-45" ModsDocOrigin="ZooKeys 721" ModsDocTitle="The geographical patterns of distribution of the genus Teuthraustes Simon, 1878 in Ecuador and description of three new species (Scorpiones, Chactidae)" checkinTime="1513120619192" checkinUser="pensoft" docAuthor="Ythier, Eric & Lourenco, Wilson R." docDate="2017" docId="611E2CDE9AAB35AB06C3EF7B9A4D207D" docLanguage="en" docName="ZooKeys 721: 45-63" docOrigin="ZooKeys 721" docSource="http://dx.doi.org/10.3897/zookeys.721.21529" docTitle="Teuthraustes khodayarii Ythier & Lourenco, 2017, sp. n." docType="treatment" docUuid="0F55C3BE-7744-4872-B55A-6B39F7960325" docUuidSource="ZooBank" docVersion="5" lastPageNumber="52" masterDocId="2E37FFB92571FFC5FFA7FFC34F5FFFB3" masterDocTitle="The geographical patterns of distribution of the genus Teuthraustes Simon, 1878 in Ecuador and description of three new species (Scorpiones, Chactidae)" masterLastPageNumber="63" masterPageNumber="45" pageNumber="49" updateTime="1668165195725" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The geographical patterns of distribution of the genus Teuthraustes Simon, 1878 in Ecuador and description of three new species (Scorpiones, Chactidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ythier, Eric</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lourenco, Wilson R.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>721</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>45</mods:start>
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<mods:end>63</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.721.21529</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.721.21529</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-721-45</mods:identifier>
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<mods:identifier type="ZBK">1300B5278A5041E38D5E1C745846BBDF</mods:identifier>
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<mods:identifier type="ZooBank">1300B5278A5041E38D5E1C745846BBDF</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="137679865" LSID="urn:lsid:zoobank.org:act:0F55C3BE-7744-4872-B55A-6B39F7960325" httpUri="http://treatment.plazi.org/id/611E2CDE9AAB35AB06C3EF7B9A4D207D" lastPageId="7" lastPageNumber="52" pageId="4" pageNumber="49">
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<subSubSection pageId="4" pageNumber="49" type="nomenclature">
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<paragraph pageId="4" pageNumber="49">
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<taxonomicName LSID="http://zoobank.org/0F55C3BE-7744-4872-B55A-6B39F7960325" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuthraustes khodayarii" order="Scorpiones" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="khodayarii">Teuthraustes khodayarii</taxonomicName>
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<taxonomicNameLabel pageId="4" pageNumber="49">sp. n.</taxonomicNameLabel>
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Fig. 3, 4, Table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="type material">
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<paragraph pageId="4" pageNumber="49">Type material.</paragraph>
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<paragraph pageId="4" pageNumber="49">
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Ecuador, Orellana Province (formerly Napo), Santa Maria de Huiririma
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-0.71666664">0°43'0"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-75.61667">75°37'0"W</geoCoordinate>
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) VI/1976 (British expedition and local people), rainforest, under log. Pre-adult female holotype. Type material deposited in the
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<normalizedToken originalValue="Muséum">Museum</normalizedToken>
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national
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<normalizedToken originalValue="d’Histoire">d'Histoire</normalizedToken>
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naturelle, Paris, France.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="etymology">
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<paragraph pageId="4" pageNumber="49">Etymology.</paragraph>
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<paragraph pageId="4" pageNumber="49">The specific name honours Dr. Khosro Khodayari, founder, President and CEO of SynTech Research, Davis, California, USA, in recognition of his support for the study of scorpions.</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="diagnosis">
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<paragraph pageId="4" pageNumber="49">Diagnosis.</paragraph>
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<paragraph pageId="4" pageNumber="49">
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Moderate to small scorpion with 35.1 mm in total length. Colouration reddish brown to dark brown. Body and appendages very intensely granulated with a rather thin granulation. Pectines with 6-6 teeth in female. Fixed and movable fingers of chela with 5-5 rows of granules; extremities of fingers with 3 granules. Ventral carinae present on metasomal segments I to IV, but weak on I. Trichobothrial pattern of type C neobothriotaxic
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<normalizedToken originalValue="‘majorante’">'majorante'</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="description">
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<paragraph pageId="4" pageNumber="49">Description.</paragraph>
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<paragraph pageId="4" pageNumber="49">Based on female holotype.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="50" pageId="4" pageNumber="49" type="colouration">
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<paragraph pageId="4" pageNumber="49">Colouration.</paragraph>
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<paragraph lastPageId="5" lastPageNumber="50" pageId="4" pageNumber="49">
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General colouration basically reddish brown to dark brown. Prosoma: carapace reddish brown with a few paler spots; eyes blackish. Mesosoma: tergites reddish brown, with VII slightly paler with some yellow spots; venter and sternites reddish yellow; pectines and genital operculum with the same colour as sternites. Metasoma: segments reddish brown, with darker zones over carinae; vesicle reddish yellow with the tip of the aculeus dark red. Chelicerae: reddish yellow, with some diffuse variegated brown spots;
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<pageBreakToken pageId="5" pageNumber="50" start="start">fingers</pageBreakToken>
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dark red with teeth reddish yellow. Pedipalps: dark brown with blackish carinae; cutting edges of fingers reddish. Legs: reddish yellow to reddish brown without spots.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="52" pageId="5" pageNumber="50" type="morphology">
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<paragraph pageId="5" pageNumber="50">Morphology.</paragraph>
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<paragraph lastPageId="7" lastPageNumber="52" pageId="5" pageNumber="50">
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Prosoma: carapace intensely granulated, with a dense and thin granulation; furrows shallow; sternum pentagonal, wider than long. Mesosoma: tergites almost
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<pageBreakToken pageId="6" pageNumber="51" start="start">acarinate</pageBreakToken>
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, with a thin and dense granulation; sternites smooth and shiny, only VII with a few granulations; spiracles oval to rounded-shaped and conspicuous; pectinal tooth count 6-6, fulcra vestigial. Metasoma: segments I to III wider than long; metasomal tegument on segments I to IV with conspicuous granulations; segment V with spinoid granulations ventrally; carinae on segments
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<normalizedToken originalValue="I–V">I-V</normalizedToken>
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strongly developed; ventral present on all segments, less marked on I; telson with strongly marked granulations, only dorsal face smooth; aculeus shorter than vesicle. Chelicerae: dentition typical of the family
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<taxonomicName family="Chacti dae" lastPageId="7" lastPageNumber="52" lsidName="" pageId="6" pageNumber="51" rank="family">
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Chacti
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<pageBreakToken pageId="7" pageNumber="52" start="start">dae</pageBreakToken>
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</taxonomicName>
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(
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<bibRefCitation author="Vachon, M" journalOrPublisher="Bulletin du Museum d'histoire naturelle" pageId="17" pageNumber="62" pagination="161 - 166" title="De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions." volume="35" year="1963">Vachon 1963</bibRefCitation>
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), and with dense setation ventrally and internally. Pedipalps: femur with dorsal internal, dorsal external and ventral internal carinae strongly marked; ventral external carina absent; dorsal, ventral and internal aspects with granulations; patella granulated with moderate carinae; chela very strongly granulated; internal aspect with conspicuous granules; dentate margins on movable and fixed fingers with 5-5 rows of granules; distal edge of fingers with three granules; trichobothriotaxy of type C; neobothriotaxic
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<normalizedToken originalValue="‘majorante’">'majorante'</normalizedToken>
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(
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<bibRefCitation author="Vachon, M" journalOrPublisher="Bulletin du Museum d'histoire naturelle" pageId="17" pageNumber="62" pagination="857 - 958" title="Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions." volume="140" year="1974">Vachon 1974</bibRefCitation>
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). Legs: tarsi with short setae disposed in a single line.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="52" type="morphometric values">
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<paragraph pageId="7" pageNumber="52">Morphometric values.</paragraph>
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<paragraph pageId="7" pageNumber="52">
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Female holotype of
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<taxonomicName lsidName="T. khodayarii" pageId="7" pageNumber="52" rank="species" species="khodayarii">T. khodayarii</taxonomicName>
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sp. n. Total length (in mm) including telson, 35.1. Carapace: length, 5.3; anterior width, 3.1; posterior width, 5.4. Mesosoma length 11.1. Metasomal segments. I: length, 1.9; width, 3.0; II: length, 2.0; width, 2.7; III: length, 2.2; width, 2.5; IV: length, 2.7; width, 2.3; V: length, 4.8; width, 2.2; depth, 2.0. Telson length, 5.1; vesicle: width, 2.0; depth, 1.6. Pedipalp: femur length, 3.8, width, 1.3; patella length, 3.7, width, 1.8; chela length, 8.1, width, 2.9, depth, 2.8; movable finger length, 4.6.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="52" type="relationships">
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<paragraph pageId="7" pageNumber="52">Relationships.</paragraph>
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<paragraph pageId="7" pageNumber="52">
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The species can be distinguished from the others congeners in particular from
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<taxonomicName class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuthraustes atramentarius" order="Scorpiones" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="atramentarius">Teuthraustes atramentarius</taxonomicName>
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Simon, which is also distributed in Ecuador, but exclusively in the high central Andes, by the following features: (i) a thin but very intense granulation over body and appendages, particularly marked on legs (ii) ocular tubercle much less pronounced, (iii) pectines are wider, (iv) pedipalpal chela are not bulk and elongated, (v) metasomal segments I to III wider than long.
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</paragraph>
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<caption pageId="7" pageNumber="52">
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<paragraph pageId="7" pageNumber="52">
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Figure 3.
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<taxonomicName class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuthraustes khodayarii" order="Scorpiones" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="khodayarii">Teuthraustes khodayarii</taxonomicName>
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sp. n. female holotype.. Habitus, dorsal A and ventral B aspects.
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</paragraph>
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</caption>
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<caption pageId="7" pageNumber="52">
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<paragraph pageId="7" pageNumber="52">
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Figure 4.
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<taxonomicName class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuthraustes khodayarii" order="Scorpiones" pageId="7" pageNumber="52" phylum="Arthropoda" rank="species" species="khodayarii">Teuthraustes khodayarii</taxonomicName>
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sp. n. Female holotype. A Chelicera, dorsal aspect B Disposition of the granulation over the dentate margins of pedipalp-chela movable finger
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<normalizedToken originalValue="C–H">C-H</normalizedToken>
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Trichobothrial pattern
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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Chela, dorso-external and ventral aspects E Femur, dorsal aspect
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<normalizedToken originalValue="F–H">F-H</normalizedToken>
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Patella, dorsal F external G and ventral H aspects. Scale bars 1 mm.v
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</paragraph>
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</caption>
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<caption ID-Table-UUID="1DEC234BA81BA92804405FA9166FC74D" httpUri="http://table.plazi.org/id/1DEC234BA81BA92804405FA9166FC74D" pageId="7" pageNumber="52">
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<paragraph pageId="7" pageNumber="52">
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Table 1. Morphometric values (in mm) of female holotypes of
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<taxonomicName lsidName="T. khodayarii" pageId="7" pageNumber="52" rank="species" species="khodayarii">T. khodayarii</taxonomicName>
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sp. n.,
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<taxonomicName lsidName="T. giupponii" pageId="7" pageNumber="52" rank="species" species="giupponii">T. giupponii</taxonomicName>
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sp. n.,
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<taxonomicName lsidName="T. kuryi" pageId="7" pageNumber="52" rank="species" species="kuryi">T. kuryi</taxonomicName>
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sp. n., and
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<taxonomicName lsidName="T. atramentarius" pageId="7" pageNumber="52" rank="species" species="atramentarius">T. atramentarius</taxonomicName>
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Simon, 1878.
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</paragraph>
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</caption>
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<paragraph pageId="7" pageNumber="52">
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<table pageId="7" pageNumber="52">
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<tr pageId="7" pageNumber="52">
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<th colspan="1" pageId="7" pageNumber="52" rowspan="1">
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<taxonomicName lsidName="T. khodayarii" pageId="7" pageNumber="52" rank="species" species="khodayarii">T. khodayarii</taxonomicName>
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</th>
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<th colspan="1" pageId="7" pageNumber="52" rowspan="1">
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<taxonomicName lsidName="T. giupponii" pageId="7" pageNumber="52" rank="species" species="giupponii">T. giupponii</taxonomicName>
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</th>
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<th colspan="1" pageId="7" pageNumber="52" rowspan="1">
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<taxonomicName lsidName="T. kuryi" pageId="7" pageNumber="52" rank="species" species="kuryi">T. kuryi</taxonomicName>
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</th>
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<th colspan="1" pageId="7" pageNumber="52" rowspan="1">
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<taxonomicName lsidName="T. atramentarius" pageId="7" pageNumber="52" rank="species" species="atramentarius">T. atramentarius</taxonomicName>
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</th>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |